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<document id="89DBBDC884D743733EC0B44C5A1CB8EC" ID-CLB-Dataset="53812" ID-DOI="10.11646/zootaxa.5250.1.1" ID-GBIF-Dataset="da8acfed-df61-403e-9a49-7f856ce56196" ID-ISSN="1175­5334" ID-Zenodo-Dep="7705657" ID-ZooBank="4729EAF3-8B78-4AAD-B373-9179D008F239" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678178160677" checkinUser="plazi" docAuthor="WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI &amp; HUANG, JIA" docDate="2023" docId="0393914C2A758721C282F9776C72FAB6" docLanguage="en" docName="zootaxa.5250.1.1.pdf" docOrigin="Zootaxa 5250 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5250.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Stegana (Steganina) nayun Li &amp; Chen, sp. nov." docType="treatment" docVersion="11" lastPageNumber="90" masterDocId="FFAAE9342A228778C215FF896958FB43" masterDocTitle="Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="88" updateTime="1699476226253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7B7D30708D209DD1D7622E24B447C1B2">Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
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<treatment id="0393914C2A758721C282F9776C72FAB6" ID-DOI="http://doi.org/10.5281/zenodo.7706345" ID-GBIF-Taxon="209579547" ID-Zenodo-Dep="7706345" LSID="urn:lsid:plazi:treatment:0393914C2A758721C282F9776C72FAB6" httpUri="http://treatment.plazi.org/id/0393914C2A758721C282F9776C72FAB6" lastPageId="89" lastPageNumber="90" pageId="87" pageNumber="88">
<subSubSection id="C32073D12A75872FC282F9776B83FC5B" box="[151,731,1790,1816]" pageId="87" pageNumber="88" type="nomenclature">
<paragraph id="8B85205A2A75872FC282F9776B83FC5B" blockId="87.[151,731,1790,1853]" box="[151,731,1790,1816]" pageId="87" pageNumber="88">
<heading id="D0CD97362A75872FC282F9776B83FC5B" bold="true" box="[151,731,1790,1816]" fontSize="11" level="1" pageId="87" pageNumber="88" reason="1">
<emphasis id="B94EFC482A75872FC282F9776B83FC5B" bold="true" box="[151,731,1790,1816]" pageId="87" pageNumber="88">
<taxonomicName id="4C3A5BD92A75872FC282F9776B28FC5B" ID-CoL="BDMR7" authority="Li &amp; Chen" authorityName="Li &amp; Chen" box="[151,624,1790,1816]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="nayun" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A75872FC282F97769A1FC5B" bold="true" box="[151,249,1790,1816]" italics="true" pageId="87" pageNumber="88">Stegana</emphasis>
(
<emphasis id="B94EFC482A75872FC31FF97768DCFC5B" bold="true" box="[266,388,1790,1816]" italics="true" pageId="87" pageNumber="88">Steganina</emphasis>
)
<emphasis id="B94EFC482A75872FC380F97768BAFC5B" bold="true" box="[405,482,1790,1816]" italics="true" pageId="87" pageNumber="88">nayun</emphasis>
Li &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel id="A27D41332A75872FC06BF9776B83FC5B" box="[638,731,1790,1816]" pageId="87" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C32073D12A75872FC282F8AB6827FC7E" box="[151,383,1826,1853]" pageId="87" pageNumber="88" type="description">
<paragraph id="8B85205A2A75872FC282F8AB6827FC7E" blockId="87.[151,731,1790,1853]" box="[151,383,1826,1853]" pageId="87" pageNumber="88">
(
<figureCitation id="13013CDF2A75872FC28AF8AB6816FC7F" box="[159,334,1826,1852]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="87" pageNumber="88">Figs. 12C, 12D</figureCitation>
,
<figureCitation id="13013CDF2A75872FC34FF8AB682FFC7E" box="[346,375,1826,1853]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="87" pageNumber="88">54</figureCitation>
)
</paragraph>
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<subSubSection id="C32073D12A75872FC282F8E3684DFCB2" pageId="87" pageNumber="88" type="diagnosis">
<paragraph id="8B85205A2A75872FC282F8E3684DFCB2" blockId="87.[151,1437,1898,2033]" pageId="87" pageNumber="88">
<emphasis id="B94EFC482A75872FC282F8E3684AFCC7" bold="true" box="[151,274,1898,1924]" pageId="87" pageNumber="88">Diagnosis.</emphasis>
This species is similar to
<emphasis id="B94EFC482A75872FC029F8E36B12FCC7" box="[572,586,1898,1924]" italics="true" pageId="87" pageNumber="88">S</emphasis>
.
<taxonomicName id="4C3A5BD92A75872FC04FF8E36D6AFCC7" authority="Wang, Gao, &amp; Chen, 2013" authorityName="Wang, Gao, &amp; Chen" authorityYear="2013" box="[602,1074,1898,1925]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="cylindrica" subGenus="Steganina">
(
<emphasis id="B94EFC482A75872FC076F8E36B29FCC7" box="[611,625,1898,1924]" italics="true" pageId="87" pageNumber="88">S</emphasis>
.)
<emphasis id="B94EFC482A75872FC09FF8E36BA4FCC7" box="[650,764,1898,1924]" italics="true" pageId="87" pageNumber="88">cylindrica</emphasis>
Wang, Gao, &amp; Chen,
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in the shape of male terminalia (as fig.
<quantity id="4CC28DBF2A75872FC2F9F807684FFCEB" box="[236,279,1934,1960]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="87" pageNumber="88" unit="in" value="2.0">2 in</quantity>
<bibRefCitation id="EFAB5DAB2A75872FC308F8066887FCEB" author="Wang, L. &amp; Gao, J. J. &amp; Chen, H. W." box="[285,479,1934,1961]" pageId="87" pageNumber="88" pagination="1993 - 2013" refId="ref74574" refString="Wang, L., Gao, J. J. &amp; Chen, H. W. (2013) Ten new species of the subgenus Steganina (Diptera: Drosophilidae: Stegana) from southern China. Journal of Natural History, 47, 1993 - 2013. https: // doi. org / 10.1080 / 00222933.2012.763055" type="journal article" year="2013">
Wang
<emphasis id="B94EFC482A75872FC370F80668C5FCEB" box="[357,413,1934,1960]" italics="true" pageId="87" pageNumber="88">et al.</emphasis>
2013
</bibRefCitation>
,
<figureCitation id="13013CDF2A75872FC3FFF8076B64FCEA" box="[490,572,1934,1961]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="87" pageNumber="88">Fig. 54</figureCitation>
), but can be distinguished from the latter by having the tubular aedeagus distoventrally with 3 rugae per side, lacking pubescence (
<figureCitation id="13013CDF2A75872FC111F83B6AD7FC8F" box="[772,911,1970,1997]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="87" pageNumber="88">Fig. 54C, D</figureCitation>
); the subepandrial sclerite lacking pubescence (
<figureCitation id="13013CDF2A75872FC28AF85F6850FCB2" box="[159,264,2006,2033]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="87" pageNumber="88">Fig. 54G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7A8721C2D2FF1F6830FA5E" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="description">
<paragraph id="8B85205A2A7A8720C2D2FF1F6ACDF97E" blockId="88.[151,1437,150,573]" pageId="88" pageNumber="89">
<emphasis id="B94EFC482A7A8720C2D2FF1F6801FBF3" bold="true" box="[199,345,150,176]" pageId="88" pageNumber="89">Description.</emphasis>
Male and female: First flagellomere brownish. Clypeus yellow. Mesonotum brown medially, brownish yellow laterally (
<figureCitation id="13013CDF2A7A8720C3D8FF326B6FFB96" box="[461,567,187,213]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="88" pageNumber="89">Fig. 12C</figureCitation>
). Katepisternum yellow (
<figureCitation id="13013CDF2A7A8720C14FFF326A9DFB96" box="[858,965,187,213]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="88" pageNumber="89">Fig. 12D</figureCitation>
). Scutellum brown medially, dark brown laterally, yellow at tip (
<figureCitation id="13013CDF2A7A8720C3B5FF566B50FBBA" box="[416,520,223,249]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="88" pageNumber="89">Fig. 12C</figureCitation>
). Halter mostly grey (
<figureCitation id="13013CDF2A7A8720C117FF566A34FBBA" box="[770,876,223,249]" captionStart="FIGURE 12" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[186,1415,183,1919]" captionTargetId="figure-19@35.[173,1415,181,1919]" captionTargetPageId="35" captionText="FIGURE 12. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) multiprocera Li &amp; Chen, sp. nov.; C, D. S. (S.) nayun Li &amp; Chen, sp. nov.; E, F. S. (S.) nigridentata Li &amp; Chen, sp. nov.; G, H. S. (S.) otphylla Cui &amp; Chen, sp. nov.; I, J. S. (S.) radiciflava Zhang &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705691" httpUri="https://zenodo.org/record/7705691/files/figure.png" pageId="88" pageNumber="89">Fig. 12D</figureCitation>
). Fore femur with a row of 4 long setae on distal part of posteroventral surface. Abdominal tergites brown to dark brown; sternites brown, about 2 times as wide as long. Male terminalia (
<figureCitation id="13013CDF2A7A8720C380FEAE6B78FA02" box="[405,544,294,321]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54AG</figureCitation>
): Epandrium with
<emphasis id="B94EFC482A7A8720C0FEFEAE6A5DFA03" box="[747,773,295,320]" italics="true" pageId="88" pageNumber="89">ca</emphasis>
. 11 setae on dorsal to posterior portions per side (
<figureCitation id="13013CDF2A7A8720C73CFEAE6CC8FA02" box="[1321,1424,294,321]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54A</figureCitation>
). Surstylus with 1 peg-like tooth and some setulae near distal margin, lacking pubescence (
<figureCitation id="13013CDF2A7A8720C663FEC26DA4FA26" box="[1142,1276,330,357]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54A, B</figureCitation>
). Cercus lacking pubescence (
<figureCitation id="13013CDF2A7A8720C341FEE668E6FACA" box="[340,446,366,393]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54A</figureCitation>
). Hypandrium strongly expanded anteromedially (
<figureCitation id="13013CDF2A7A8720C1E3FEE66DD9FACA" box="[1014,1153,366,393]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54C, D</figureCitation>
). Pregonite small, with 2 sensilla (
<figureCitation id="13013CDF2A7A8720C2EFFE1A68D9FAEE" box="[250,385,402,429]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54C, D</figureCitation>
). Aedeagus lacking serrated process (
<figureCitation id="13013CDF2A7A8720C134FE1A6AF1FAEE" box="[801,937,402,429]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54C, D</figureCitation>
). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (
<figureCitation id="13013CDF2A7A8720C070FE3E6BB3FA92" box="[613,747,438,465]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54E, F</figureCitation>
). Subepandrial sclerite slightly rectangle distally (
<figureCitation id="13013CDF2A7A8720C733FE3E6CC8FA92" box="[1318,1424,438,465]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54G</figureCitation>
). Female terminalia (
<figureCitation id="13013CDF2A7A8720C360FE5268A1FAB6" box="[373,505,474,501]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54HJ</figureCitation>
): Epiproct with several setae on distal 1/2 (
<figureCitation id="13013CDF2A7A8720C1FCFE526D0CFAB6" box="[1001,1108,474,501]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54H</figureCitation>
). Cercus and hypoproct with numerous setae and much pubescence (
<figureCitation id="13013CDF2A7A8720C05EFE766B90F95A" box="[587,712,510,537]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54H, J</figureCitation>
). Eight sternite about 1.5 times as long as wide, glabrous on basal 4/5, with several setae and some pubescence on distal 1/5 (
<figureCitation id="13013CDF2A7A8720C13FFDAA6AD0F97E" box="[810,904,546,573]" captionStart="FIGURE 54" captionStartId="88.[151,250,1801,1825]" captionTargetBox="[160,1429,607,1772]" captionTargetId="figure-231@88.[151,1436,597,1777]" captionTargetPageId="88" captionText="FIGURE 54. Stegana (Steganina) nayun Li &amp; Chen, sp. nov., male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89">Fig. 54I</figureCitation>
).
</paragraph>
<caption id="DF4570D22A7A8720C282F8806D7AFCCE" ID-DOI="http://doi.org/10.5281/zenodo.7705802" ID-Zenodo-Dep="7705802" httpUri="https://zenodo.org/record/7705802/files/figure.png" pageId="88" pageNumber="89" startId="88.[151,250,1801,1825]" targetBox="[160,1429,607,1772]" targetPageId="88">
<paragraph id="8B85205A2A7A8720C282F8806D7AFCCE" blockId="88.[151,1436,1801,1933]" pageId="88" pageNumber="89">
<emphasis id="B94EFC482A7A8720C282F8806879FC62" bold="true" box="[151,289,1801,1825]" pageId="88" pageNumber="89">FIGURE 54.</emphasis>
<taxonomicName id="4C3A5BD92A7A8720C33CF8806BE0FC62" authority="Li &amp; Chen" authorityName="Li &amp; Chen" box="[297,696,1801,1825]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="nayun" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A7A8720C33CF8806823FC63" box="[297,379,1801,1824]" italics="true" pageId="88" pageNumber="89">Stegana</emphasis>
(
<emphasis id="B94EFC482A7A8720C39EF88068A8FC63" box="[395,496,1801,1824]" italics="true" pageId="88" pageNumber="89">Steganina</emphasis>
)
<emphasis id="B94EFC482A7A8720C014F8836B67FC62" box="[513,575,1802,1825]" italics="true" pageId="88" pageNumber="89">nayun</emphasis>
Li &amp; Chen
</taxonomicName>
,
<emphasis id="B94EFC482A7A8720C0D1F8806A4EFC62" bold="true" box="[708,790,1801,1825]" pageId="88" pageNumber="89">
<taxonomicNameLabel id="A27D41332A7A8720C0D1F8806A4EFC62" box="[708,790,1801,1825]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (AG) and female terminalia (HJ). A. Epandrium, surstylus, and cercus in lateral view; B. Surstylus in ventral view; C, D. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; E, F.Aedeagal sheath in ventral and lateral views; G. Subepandrial sclerite in ventral view; H. Epiproct and cercus in dorsal view; I. Eighth sternite in ventral view; J. Hypoproct in ventral view.
</paragraph>
</caption>
<paragraph id="8B85205A2A7B8721C282FF1E6830FA5E" blockId="89.[151,1436,150,501]" pageId="89" pageNumber="90">HW = 0.38 (0.42), ch/o = 0.29 (0.38), prorb = 1.05 (1.50), rcorb = 0.79 (0.88), vb = 0.64 (0.93), dcl = 0.43 (0.35), presctl = 0.70 (0.70), sctl = 3.15 (2.39), sterno = 1.19 (0.93), orbito = 1.89 (1.89), dcp = 0.21 (0.21), sctlp = 1.78 (2.22), C = 1.98 (1.95), 4c = 1.05 (1.39), 4v = 1.20 (2.21), 5x = 1.54 (1.91), ac = 10.25 (13.00), M = 0.55 (0.76), C3F = 0.61 (0.70).</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7B8721C2D2FEAE6AA8FAEE" pageId="89" pageNumber="90" type="materials_examined">
<paragraph id="8B85205A2A7B8721C2D2FEAE6AA8FAEE" blockId="89.[151,1436,150,501]" pageId="89" pageNumber="90">
<emphasis id="B94EFC482A7B8721C2D2FEAE68DFFA03" bold="true" box="[199,391,294,321]" pageId="89" pageNumber="90">Type materials.</emphasis>
<materialsCitation id="3B522A072A7B8721C380FEAE6A99FA26" ID-GBIF-Occurrence="4097216461" collectingDate="2017-10-25" collectionCode="SCAU" collectorName="H. W. Chen" country="China" elevation="940" latitude="22.324722" location="Menglian" longLatPrecision="21" longitude="99.583336" municipality="Jinshan" pageId="89" pageNumber="90" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54819EF82A7B8721C380FEAE68A5FA02" box="[405,509,295,321]" pageId="89" pageNumber="90" type="holotype">Holotype</typeStatus>
♂ (
<collectionCode id="ED2BB89F2A7B8721C022FEAE6BDEFA02" box="[567,646,295,321]" pageId="89" pageNumber="90">SCAU</collectionCode>
, No. 125730, DNA #2674),
<collectingCountry id="F32D60CA2A7B8721C1FDFEAE6D1DFA02" box="[1000,1093,295,321]" name="China" pageId="89" pageNumber="90">CHINA</collectingCountry>
:
<collectingMunicipality id="6BE1BA202A7B8721C642FEAE6DF6FA02" box="[1111,1198,295,321]" pageId="89" pageNumber="90">Jinshan</collectingMunicipality>
,
<location id="8EE576812A7B8721C6D5FEAE6C76FA02" LSID="urn:lsid:plazi:treatment:0393914C2A758721C282F9776C72FAB6:8EE576812A7B8721C6D5FEAE6C76FA02" box="[1216,1326,295,321]" country="China" latitude="22.324722" longLatPrecision="21" longitude="99.583336" municipality="Jinshan" name="Menglian" pageId="89" pageNumber="90" stateProvince="Yunnan">Menglian</location>
,
<collectingRegion id="49FEEEB82A7B8721C755FEAE6CC1FA02" box="[1344,1433,295,321]" country="China" name="Yunnan" pageId="89" pageNumber="90">Yunnan</collectingRegion>
,
<geoCoordinate id="EE0E469D2A7B8721C282FEC26844FA26" box="[151,284,331,357]" degrees="22" direction="north" minutes="19" orientation="latitude" pageId="89" pageNumber="90" precision="15" seconds="29" value="22.324722">22°19'29''N</geoCoordinate>
,
<geoCoordinate id="EE0E469D2A7B8721C332FEC368F1FA26" box="[295,425,330,357]" degrees="99" direction="east" minutes="35" orientation="longitude" pageId="89" pageNumber="90" precision="15" seconds="00" value="99.583336">99°35'00''E</geoCoordinate>
, alt.
<quantity id="4CC28DBF2A7B8721C3C8FEC26B78FA26" box="[477,544,331,357]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="89" pageNumber="90" unit="m" value="940.0">
<elevation id="0017C7692A7B8721C3C8FEC26B78FA26" box="[477,544,331,357]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="89" pageNumber="90" unit="m" value="940.0">940m</elevation>
</quantity>
,
<date id="FF84069A2A7B8721C03FFEC36BC5FA26" box="[554,669,330,357]" pageId="89" pageNumber="90" value="2017-10-25">
<collectingDate id="EFC0FF722A7B8721C03FFEC36BC5FA26" box="[554,669,330,357]" pageId="89" pageNumber="90" value="2017-10-25">25.x.2017</collectingDate>
</date>
, ex tree trunk,
<collectorName id="26CF458C2A7B8721C154FEC26A99FA26" box="[833,961,331,357]" pageId="89" pageNumber="90">H.W. Chen</collectorName>
</materialsCitation>
.
<materialsCitation id="3B522A072A7B8721C1DFFEC26B19FACA" ID-GBIF-Occurrence="4097216358" collectingDate="2017-10-25" collectionCode="SCAU" collectorName="H. W. Chen" country="China" elevation="940" latitude="22.324722" location="Menglian" longLatPrecision="21" longitude="99.583336" municipality="Jinshan" pageId="89" pageNumber="90" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54819EF82A7B8721C1DFFEC26D63FA26" box="[970,1083,331,357]" pageId="89" pageNumber="90" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3CEBD32A7B8721C650FEC26D30FA26" box="[1093,1128,331,357]" pageId="89" pageNumber="90" type="female">1♀</specimenCount>
(
<collectionCode id="ED2BB89F2A7B8721C663FEC26D9DFA26" box="[1142,1221,331,357]" pageId="89" pageNumber="90">SCAU</collectionCode>
, No. 125731, DNA #2693), other data same as holotype
</materialsCitation>
;
<materialsCitation id="3B522A072A7B8721C05AFEE66AA8FAEE" ID-GBIF-Occurrence="4097216359" collectingDate="2017-07-18" collectionCode="SCAU" collectorName="Y. L. Wang" country="China" elevation="1500" latitude="30.545" location="Baoxing" longLatPrecision="20" longitude="102.90667" municipality="Fengtongzhai" pageId="89" pageNumber="90" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D3CEBD32A7B8721C05AFEE66B2AFACA" box="[591,626,367,393]" pageId="89" pageNumber="90" type="female">1♀</specimenCount>
(
<collectionCode id="ED2BB89F2A7B8721C091FEE66B8BFACA" box="[644,723,367,393]" pageId="89" pageNumber="90">SCAU</collectionCode>
, No. 125787, DNA #2608),
<collectingMunicipality id="6BE1BA202A7B8721C634FEE66DE4FACA" box="[1057,1212,367,393]" pageId="89" pageNumber="90">Fengtongzhai</collectingMunicipality>
,
<location id="8EE576812A7B8721C6DEFEE66C75FACA" LSID="urn:lsid:plazi:treatment:0393914C2A758721C282F9776C72FAB6:8EE576812A7B8721C6DEFEE66C75FACA" box="[1227,1325,367,393]" country="China" latitude="30.545" longLatPrecision="20" longitude="102.90667" municipality="Fengtongzhai" name="Baoxing" pageId="89" pageNumber="90" stateProvince="Sichuan">Baoxing</location>
,
<collectingRegion id="49FEEEB82A7B8721C72EFEE66CC0FACA" box="[1339,1432,367,393]" country="China" name="Sichuan" pageId="89" pageNumber="90">Sichuan</collectingRegion>
,
<geoCoordinate id="EE0E469D2A7B8721C282FE1A6844FAEE" box="[151,284,403,429]" degrees="30" direction="north" minutes="32" orientation="latitude" pageId="89" pageNumber="90" precision="15" seconds="42" value="30.545">30°32'42''N</geoCoordinate>
,
<geoCoordinate id="EE0E469D2A7B8721C33DFE1B68E0FAEE" box="[296,440,402,429]" degrees="102" direction="east" minutes="54" orientation="longitude" pageId="89" pageNumber="90" precision="15" seconds="24" value="102.90667">102°54'24''E</geoCoordinate>
, alt.
<quantity id="4CC28DBF2A7B8721C3F8FE1B6B65FAEE" box="[493,573,402,429]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="89" pageNumber="90" unit="m" value="1500.0">
<elevation id="0017C7692A7B8721C3F8FE1B6B65FAEE" box="[493,573,402,429]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="89" pageNumber="90" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="FF84069A2A7B8721C05CFE1A6B93FAEE" box="[585,715,403,429]" pageId="89" pageNumber="90" value="2017-07-18">
<collectingDate id="EFC0FF722A7B8721C05CFE1A6B93FAEE" box="[585,715,403,429]" pageId="89" pageNumber="90" value="2017-07-18">18.vii.2017</collectingDate>
</date>
, ex tree trunk,
<collectorName id="26CF458C2A7B8721C164FE1A6AB3FAEE" box="[881,1003,403,429]" pageId="89" pageNumber="90">Y.L. Wang</collectorName>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7B8721C2D2FE3F6BD0FA92" box="[199,648,438,465]" pageId="89" pageNumber="90" type="distribution">
<paragraph id="8B85205A2A7B8721C2D2FE3F6BD0FA92" blockId="89.[151,1436,150,501]" box="[199,648,438,465]" pageId="89" pageNumber="90">
<emphasis id="B94EFC482A7B8721C2D2FE3F6839FA93" bold="true" box="[199,353,438,464]" pageId="89" pageNumber="90">Distribution.</emphasis>
<collectingCountry id="F32D60CA2A7B8721C372FE3E68F2FA92" box="[359,426,439,465]" name="China" pageId="89" pageNumber="90">China</collectingCountry>
(
<collectingRegion id="49FEEEB82A7B8721C3ACFE3E6B40FA92" box="[441,536,439,465]" country="China" name="Sichuan" pageId="89" pageNumber="90">Sichuan</collectingRegion>
,
<collectingRegion id="49FEEEB82A7B8721C037FE3E6B25FA92" box="[546,637,439,465]" country="China" name="Yunnan" pageId="89" pageNumber="90">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A7B8721C2D2FE526C72FAB6" box="[199,1322,475,501]" pageId="89" pageNumber="90" type="etymology">
<paragraph id="8B85205A2A7B8721C2D2FE526C72FAB6" blockId="89.[151,1436,150,501]" box="[199,1322,475,501]" pageId="89" pageNumber="90">
<emphasis id="B94EFC482A7B8721C2D2FE526816FAB6" bold="true" box="[199,334,475,501]" pageId="89" pageNumber="90">Etymology.</emphasis>
From the name of the
<typeStatus id="54819EF82A7B8721C058FE526B25FAB6" box="[589,637,475,501]" pageId="89" pageNumber="90">type</typeStatus>
locality &quot;Jinshan&quot; belonging to &quot;Nayun, Menglian,
<collectingRegion id="49FEEEB82A7B8721C6D4FE526C46FAB6" box="[1217,1310,475,501]" country="China" name="Yunnan" pageId="89" pageNumber="90">Yunnan</collectingRegion>
&quot;.
</paragraph>
</subSubSection>
</treatment>
</document>