treatments-xml/data/03/85/F3/0385F373FFA5B44DFF6C7AE2BACC036E.xml

<document id="70EC71F7FE8176132F84ACFA2DE00225" ID-DOI="10.11646/zootaxa.3860.2.1" ID-GBIF-Dataset="75d1fc6b-8d95-48e6-a52a-1a9ab0ce9fb0" ID-ISSN="1175-5326" ID-Zenodo-Dep="229557" ID-ZooBank="DD34DE75-74F3-42B1-9224-DC3BF9F3CCC7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460903743442" checkinUser="plazi" docAuthor="Aballay, Fernando H., Chani-Posse, Mariana R., Ayón, María Rosana, Maldonado, María Belén &amp; Centeno, Néstor D." docDate="2014" docId="0385F373FFA5B44DFF6C7AE2BACC036E" docLanguage="en" docName="zt03860p124.pdf" docOrigin="Zootaxa 3860 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Staphylinidae" docType="key" docVersion="8" lastPageNumber="104" masterDocId="FFBC8B0BFFA7B44EFFFB7968BF50042D" masterDocTitle="An illustrated key to and diagnoses of the species of Staphylinidae (Coleoptera) associated with decaying carcasses in Argentina" masterLastPageNumber="124" masterPageNumber="101" pageNumber="103" updateTime="1698360573237" updateUser="plazi">
<mods:mods id="D97E91FBF47F4E5F62192B6AD9A9D18D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B6E66E2D5788F674E51302023E4B14A8">
<mods:title id="1AC08B8BD02EB9ECA964D49A39E57FC6">An illustrated key to and diagnoses of the species of Staphylinidae (Coleoptera) associated with decaying carcasses in Argentina</mods:title>
</mods:titleInfo>
<mods:name id="969C66797D8257EAD1A638367D744189" type="personal">
<mods:role id="C2B5E474A8A0204EA6E6DF2D23307D15">
<mods:roleTerm id="661390BCE8C23FBD37484824BF6D81D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="434E1FB34EB1EA7C5B6E34AAE5792ED1">Aballay, Fernando H.</mods:namePart>
</mods:name>
<mods:name id="FA76B8D77D90546EE0B034841D0FD261" type="personal">
<mods:role id="45D3D3978E572A9E7F7E63037054C02A">
<mods:roleTerm id="147B076D3A819132169A0BED0D8ACB08">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AAF50BEA11B57EB65CE54DCDF2D8C982">Chani-Posse, Mariana R.</mods:namePart>
</mods:name>
<mods:name id="FF6549B46D82AC0CDCF5F7C73DE04E6A" type="personal">
<mods:role id="FBFAC99B6D5F367696BDD12F83FFC930">
<mods:roleTerm id="2F99033F84997F9214E431868B6353D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9FB08F40AD7CD2F1CB14FA3D8748EA5">Ayón, María Rosana</mods:namePart>
</mods:name>
<mods:name id="EDC3D2D8CF5BA9546091F723C6170BFD" type="personal">
<mods:role id="3834F74FE63808EBC6A1636F4AD9F8F3">
<mods:roleTerm id="5B96F352C611CDE2E5D5D7E12011E18B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE6D414B541E4FAB558D2CECBB79A9BF">Maldonado, María Belén</mods:namePart>
</mods:name>
<mods:name id="5F17E166AB96337F4CB8F24C6ECC16AA" type="personal">
<mods:role id="706FA29044B403C3D6E3F68C0C788D8A">
<mods:roleTerm id="41856F959EE9A46F96C2FC87A36A0060">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7798200CB907F860A71927A13F725A6F">Centeno, Néstor D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0ECFD108716EC9EFAC1B2FD61EED20DC">text</mods:typeOfResource>
<mods:relatedItem id="AB7595BA5F7F4F339095BEEF798A6635" type="host">
<mods:titleInfo id="029F647BD6E970D0BB88706EC76AA19C">
<mods:title id="168280EC8C925037C3FACDFE72F774D3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8FF94824C42868B2CA374A8A0324EE7E">
<mods:date id="73E8301704AB0CA6A6B7AE3116FB6047">2014</mods:date>
<mods:detail id="D075FFAA1167EEE910619DC7FEA572B4" type="volume">
<mods:number id="C365C1284C45DE0CB4F4E89E5955BAE7">3860</mods:number>
</mods:detail>
<mods:detail id="67FEB50FB55FD602FFBA0FB9E2C4B899" type="issue">
<mods:number id="D03897567337127EF3D209AE8621CCFF">2</mods:number>
</mods:detail>
<mods:extent id="2A33DDB02E12DAEC10BF9ED3DFA66D6E" unit="page">
<mods:start id="9829317B903B56F4B0FED98291368EE8">101</mods:start>
<mods:end id="0E594C3DEC49D8C9B3D3EE8931B009F8">124</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BFA389F7619FB0D18C96B21629E88071">journal article</mods:classification>
<mods:identifier id="3ED7F8B5D6BFCC5D032A25B585652103" type="DOI">10.11646/zootaxa.3860.2.1</mods:identifier>
<mods:identifier id="7FDF690BC830177C0A64EC2D984F7EB8" type="GBIF-Dataset">75d1fc6b-8d95-48e6-a52a-1a9ab0ce9fb0</mods:identifier>
<mods:identifier id="515607733A08D8241293784B5F4794C9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BC2B52C3115DFF32276F9BFFA8F8F962" type="Zenodo-Dep">229557</mods:identifier>
<mods:identifier id="F467D82ABA2CBA10FA12B5426CA4343F" type="ZooBank">DD34DE75-74F3-42B1-9224-DC3BF9F3CCC7</mods:identifier>
</mods:mods>
<treatment id="0385F373FFA5B44DFF6C7AE2BACC036E" ID-DOI="http://doi.org/10.5281/zenodo.5688943" ID-GBIF-Taxon="119587841" ID-Zenodo-Dep="5688943" LSID="urn:lsid:plazi:treatment:0385F373FFA5B44DFF6C7AE2BACC036E" httpUri="http://treatment.plazi.org/id/0385F373FFA5B44DFF6C7AE2BACC036E" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103">
<subSubSection id="C33611EEFFA5B44CFF6C7AE2BCAE0789" box="[151,1022,906,932]" pageId="2" pageNumber="103" type="nomenclature">
<paragraph id="8B934265FFA5B44CFF6C7AE2BCAE0789" blockId="2.[151,1022,906,932]" box="[151,1022,906,932]" pageId="2" pageNumber="103">
<heading id="D0DBF509FFA5B44CFF6C7AE2BCAE0789" bold="true" box="[151,1022,906,932]" fontSize="11" level="1" pageId="2" pageNumber="103" reason="1">
<emphasis id="B9589E77FFA5B44CFF6C7AE2BCAE0789" bold="true" box="[151,1022,906,932]" pageId="2" pageNumber="103">
Key to species of 
<taxonomicName id="4C2C39E6FFA5B44CFE957AE2BD490789" ID-CoL="8VR4S" box="[366,537,906,932]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
associated with carcasses in 
<collectingCountry id="F33B02F5FFA5B44CFC847AE2BCAE0789" box="[895,1022,906,932]" name="Argentina" pageId="2" pageNumber="103">Argentina</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33611EEFFA5B44DFF6C7AB9BACC036E" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103" type="key">
<keyStep id="30D85AC0FFA5B44CFF6C7AB9BACB0029" pageId="2" pageNumber="103">
<paragraph id="8B934265FFA5B44CFF6C7AB9BACC07CB" blockId="2.[151,1436,977,1979]" box="[151,1436,977,998]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7AB9BACC07CB" box="[151,1436,977,998]" pageId="2" pageNumber="103">
1 Antennae inserted posterior to a line drawn between anterior margin of eyes (
<figureCitation id="13175EE0FFA5B44CFC3D7AB9BB5C07CB" box="[966,1036,977,998]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">Fig. 25</figureCitation>
)................... 2. 
<taxonomicName id="4C2C39E6FFA5B44CFAE77AB9BACC07CB" ID-CoL="8VR4S" box="[1308,1436,977,998]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="family">Aleocharinae</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA5B44CFF6C7A87BACB0029" blockId="2.[151,1436,977,1979]" box="[151,1435,1007,1028]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7A87BACB0029" box="[151,1435,1007,1028]" pageId="2" pageNumber="103">
- Antennae inserted anterior to a line drawn between anterior margin of eyes (
<figureCitation id="13175EE0FFA5B44CFC407A87BB510029" box="[955,1025,1007,1028]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">Fig. 26</figureCitation>
)................................ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA5B44CFF6C7D66BE6C0052" pageId="2" pageNumber="103">
<paragraph id="8B934265FFA5B44CFF6C7D66BE92006F" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7D66BE92006F" pageId="2" pageNumber="103">
2 Maxillary and labial palpi with apical pseudosegment, 5-articled and 4-articled respectively (
<figureCitation id="13175EE0FFA5B44CFBA07D66BBF3000E" box="[1115,1187,1038,1059]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">Fig. 27</figureCitation>
); tarsal formula 5-5-5.. 3 
<taxonomicName id="4C2C39E6FFA5B44CFF1A7D45BE92006F" ID-CoL="8VRG3" authority="Gravenhorst" authorityName="Gravenhorst" box="[225,450,1069,1090]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA5B44CFF1A7D45BE15006F" box="[225,325,1069,1090]" italics="true" pageId="2" pageNumber="103">Aleochara</emphasis>
Gravenhorst
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA5B44CFF6C7D24BE6C0052" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7D24BE6C0052" pageId="2" pageNumber="103">
- Maxillary and labial palpi without apical pseudosegment, 4-articled and 3-articled respectively; tarsal formula 4-5-5... 
<taxonomicName id="4C2C39E6FFA5B44CFAA27D24BE6C0052" ID-CoL="8VRBL" authority="Thomson" authorityName="Thomson" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA5B44CFAA27D24BACC004C" box="[1369,1436,1100,1121]" italics="true" pageId="2" pageNumber="103">Atheta</emphasis>
Thomson
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA5B44CFF6C7DE1BACB0134" pageId="2" pageNumber="103">
<paragraph id="8B934265FFA5B44CFF6C7DE1BACC00D7" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7DE1BACC00D7" pageId="2" pageNumber="103">
3 Pronotum with two longitudinal, subparallel rows of setiferous punctures, arranged in more or less defined lines (
<figureCitation id="13175EE0FFA5B44CFAE47DE1BA3800B3" box="[1311,1384,1161,1182]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">Fig. 30</figureCitation>
)......................................................................... subgenus 
<taxonomicName id="4C2C39E6FFA5B44CFB867DCFBACB0091" authority="Mulsant and Rey" authorityName="Mulsant and Rey" box="[1149,1435,1191,1212]" class="Insecta" family="Staphylinidae" genus="Coprochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA5B44CFB867DCFBBA10091" box="[1149,1265,1191,1212]" italics="true" pageId="2" pageNumber="103">Coprochara</emphasis>
Mulsant and Rey
</taxonomicName>
Head and pronotum scarcely pubescent, elytra with two small apico-medial spots not well defined and blurred (
<figureCitation id="13175EE0FFA5B44CFAA97DAEBAC700F6" box="[1362,1431,1222,1243]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="2" pageNumber="103">Figs. 4</figureCitation>
, 
<figureCitation id="13175EE0FFA5B44CFEF07D8DBE7600D7" box="[267,294,1253,1274]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">30</figureCitation>
)…........................................................ 
<taxonomicName id="4C2C39E6FFA5B44CFC1D7D8DBACC00D7" ID-CoL="BMYC" authority="Fairmaire &amp; Germain" authorityName="Fairmaire &amp; Germain" box="[998,1436,1253,1274]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="signaticollis">
<emphasis id="B9589E77FFA5B44CFC1D7D8DBB9600D7" box="[998,1222,1253,1274]" italics="true" pageId="2" pageNumber="103">Aleochara signaticollis</emphasis>
Fairmaire &amp; Germain
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA5B44CFF6C7C6CBACB0134" blockId="2.[151,1436,977,1979]" box="[151,1435,1284,1305]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7C6CBACB0134" box="[151,1435,1284,1305]" pageId="2" pageNumber="103">
- Pronotum evenly pubescent (
<figureCitation id="13175EE0FFA5B44CFE037C6CBD3B0134" box="[504,619,1284,1305]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">Figs. 28, 29</figureCitation>
).................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA5B44CFF6C7C4ABACC01FC" pageId="2" pageNumber="103">
<paragraph id="8B934265FFA5B44CFF6C7C4ABACC0159" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7C4ABACC0159" pageId="2" pageNumber="103">
4 Mesosternum completely carinate along midline (
<figureCitation id="13175EE0FFA5B44CFD467C4ABC55011A" box="[701,773,1314,1335]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">Fig. 31</figureCitation>
)....................... subgenus 
<taxonomicName id="4C2C39E6FFA5B44CFB787C4ABE0B017B" authority="Mulsant and Rey Antennal" authorityName="Mulsant and Rey Antennal" class="Insecta" family="Staphylinidae" genus="Xenochara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA5B44CFB787C4ABBBC011A" box="[1155,1260,1314,1335]" italics="true" pageId="2" pageNumber="103">Xenochara</emphasis>
Mulsant and Rey Antennal
</taxonomicName>
segments 4–6 slightly elongate to quadrate, 5–10 slightly transverse; pronotum and elytra densely pubescent; elytral pubescence directed posterad (
<figureCitation id="13175EE0FFA5B44CFDA97C37BDC60159" box="[594,662,1375,1396]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="2" pageNumber="103">Figs. 3</figureCitation>
, 
<figureCitation id="13175EE0FFA5B44CFD5A7C37BDEA0159" box="[673,698,1375,1396]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">29</figureCitation>
)........................................ 
<taxonomicName id="4C2C39E6FFA5B44CFB5E7C37BACC0159" ID-CoL="BNCJ" authority="Klug" authorityName="Klug" box="[1189,1436,1375,1396]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="puberula">
<emphasis id="B9589E77FFA5B44CFB5E7C37BA350159" box="[1189,1381,1375,1396]" italics="true" pageId="2" pageNumber="103">Aleochara puberula</emphasis>
Klug
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA5B44CFF6C7C16BACC01FC" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7C16BACC01FC" pageId="2" pageNumber="103">
- Mesosternum not carinate along midline (
<figureCitation id="13175EE0FFA5B44CFD8A7C16BDE901BE" box="[625,697,1406,1427]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">Fig. 32</figureCitation>
); body robust and compact......... subgenus 
<taxonomicName id="4C2C39E6FFA5B44CFB737C16BE0D019F" ID-CoL="8VRG3" authority="Mulsant and Rey Antennal" authorityName="Mulsant and Rey Antennal" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA5B44CFB737C16BBBC01BE" box="[1160,1260,1406,1427]" italics="true" pageId="2" pageNumber="103">Aleochara</emphasis>
Mulsant and Rey Antennal
</taxonomicName>
segments 4–6 distinctly transverse; head and pronotum sparsely pubescent; elytra with two latero-medial brown spots (
<figureCitation id="13175EE0FFA5B44CFEB87CD4BED701FC" box="[323,391,1468,1489]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="2" pageNumber="103">Figs. 2</figureCitation>
, 
<figureCitation id="13175EE0FFA5B44CFE6A7CD4BEFA01FC" box="[401,426,1468,1489]" captionStart="FIGURE 25 – 32" captionStartId="10.[151,250,1941,1963]" captionTargetBox="[263,1331,183,1921]" captionTargetId="figure@10.[260,1331,183,1922]" captionTargetPageId="10" captionText="FIGURE 25 – 32. Head (lateral view): 25, Aleochara puberula; 26, Philonthus longicornis. Maxillary palpus: 27, A. puberula. Head and pronotum: 28, A. bonariensis; 29, A. puberula; 30, A. signaticollis. Mesosternum: 31, A. puberula; 32, A. signaticollis. Abbreviations: Mc; mesosternum carina; MxP, maxillary palpus." httpUri="https://zenodo.org/record/229560/files/figure.png" pageId="2" pageNumber="103">28</figureCitation>
)........................................................... 
<taxonomicName id="4C2C39E6FFA5B44CFB847CD4BACC01FC" ID-CoL="BMQM" authority="Lynch" authorityName="Lynch" box="[1151,1436,1468,1489]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B9589E77FFA5B44CFB847CD4BA0901FC" box="[1151,1369,1468,1489]" italics="true" pageId="2" pageNumber="103">Aleochara bonariensis</emphasis>
Lynch
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA5B44CFF6C7CB2BACC0266" pageId="2" pageNumber="103">
<paragraph id="8B934265FFA5B44CFF6C7CB2BACC0201" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7CB2BACC0201" pageId="2" pageNumber="103">
5 Abdomen with complete sternum 2 (seven sterna can be counted) (
<figureCitation id="13175EE0FFA5B44CFC9A7CB2BCF701C2" box="[865,935,1498,1519]" captionStart="FIGURE 42 – 49" captionStartId="13.[151,250,1680,1702]" captionTargetBox="[158,1436,509,1655]" captionTargetId="figure@13.[151,1436,509,1655]" captionTargetPageId="13" captionText="FIGURE 42 – 49. Abdomen: 42, Platydracus scabrosus; 43, Anotylus sp. Front tarsus: 44, Belonuchus rufipennis; 45, Philonthus longicornis; 46, Xenopygus analis; 47, Styngetus viduus. Femur: 48, Xenopygus analis; 49, Styngetus viduus. Abbreviation: Fe, femur." httpUri="https://zenodo.org/record/229563/files/figure.png" pageId="2" pageNumber="103">Fig. 43</figureCitation>
)................................ 
<taxonomicName id="4C2C39E6FFA5B44CFAC97CB2BACB01C2" ID-CoL="8VR5V" box="[1330,1435,1498,1519]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="subFamily" subFamily="Oxytelinae">Oxytelinae</taxonomicName>
Pronotum transverse, with longitudinal impressions (
<figureCitation id="13175EE0FFA5B44CFCF27C91BC000223" box="[777,848,1529,1550]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="2" pageNumber="103">Fig. 35</figureCitation>
); scutellum with tripartite impression, of a fleur-de-lis shape.................................................................................. 
<taxonomicName id="4C2C39E6FFA5B44CFB1C7F7FBACC0201" ID-CoL="8VRGY" authority="Thomson" authorityName="Thomson" box="[1255,1436,1559,1580]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA5B44CFB1C7F7FBA6B0201" box="[1255,1339,1559,1580]" italics="true" pageId="2" pageNumber="103">Anotylus</emphasis>
Thomson
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA5B44CFF6C7F5EBACC0266" blockId="2.[151,1436,977,1979]" box="[151,1436,1590,1611]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7F5EBACC0266" box="[151,1436,1590,1611]" pageId="2" pageNumber="103">
- Abdomen with sternum 2 absent or rudimentary, only six complete sterna are visible (
<figureCitation id="13175EE0FFA5B44CFBEC7F5EBB0F0266" box="[1047,1119,1590,1611]" captionStart="FIGURE 42 – 49" captionStartId="13.[151,250,1680,1702]" captionTargetBox="[158,1436,509,1655]" captionTargetId="figure@13.[151,1436,509,1655]" captionTargetPageId="13" captionText="FIGURE 42 – 49. Abdomen: 42, Platydracus scabrosus; 43, Anotylus sp. Front tarsus: 44, Belonuchus rufipennis; 45, Philonthus longicornis; 46, Xenopygus analis; 47, Styngetus viduus. Femur: 48, Xenopygus analis; 49, Styngetus viduus. Abbreviation: Fe, femur." httpUri="https://zenodo.org/record/229563/files/figure.png" pageId="2" pageNumber="103">Fig. 42</figureCitation>
);............ 6 
<taxonomicName id="4C2C39E6FFA5B44CFAE37F5EBACC0266" ID-CoL="8VR6D" box="[1304,1436,1590,1611]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA5B44CFF6C7F3DBACB032E" pageId="2" pageNumber="103">
<paragraph id="8B934265FFA5B44CFF6C7F3DBACB02C9" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7F3DBACB02C9" pageId="2" pageNumber="103">
6 With sclerotized plate (solid or divided into two sclerites) in front of prosternum (
<figureCitation id="13175EE0FFA5B44CFC147F3DBB650247" box="[1007,1077,1621,1642]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="2" pageNumber="103">Fig. 33</figureCitation>
)................... 
<taxonomicName id="4C2C39E6FFA5B44CFAD87F3DBACC0247" ID-CoL="8VRBW" box="[1315,1436,1621,1642]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="tribe" tribe="Xantholinini">Xantholinini</taxonomicName>
6.1 Pronotal disc impunctate except near margins; head, pronotum and abdomen of metallic color; length&gt; 
<quantity id="4CD4EF80FFA5B44CFAF07F1CBA1E02A4" box="[1291,1358,1652,1673]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="103" unit="mm" value="20.0">20 mm</quantity>
(
<figureCitation id="13175EE0FFA5B44CFAA07F1CBAC402A4" box="[1371,1428,1652,1673]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="2" pageNumber="103">Fig. 6</figureCitation>
)....................................................................... 
<taxonomicName id="4C2C39E6FFA5B44CFBA27FFABACB028A" ID-CoL="3CBN4" authority="Mannerheim" authorityName="Mannerheim" box="[1113,1435,1682,1703]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="chalybaeus">
<emphasis id="B9589E77FFA5B44CFBA27FFABA4B028A" box="[1113,1307,1682,1703]" italics="true" pageId="2" pageNumber="103">Eulissus chalybaeus</emphasis>
Mannerheim
</taxonomicName>
6.2 Pronotal disc with distinct dorsal and lateral rows of punctures; head, pronotum and abdomen brownish, glossy but never of metallic color; length distinctly &lt;
<quantity id="4CD4EF80FFA5B44CFD5A7FA7BDB402C9" box="[673,740,1743,1764]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="103" unit="mm" value="20.0">20 mm</quantity>
(
<figureCitation id="13175EE0FFA5B44CFD097FA7BC7B02C9" box="[754,811,1743,1764]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="2" pageNumber="103">Fig. 7</figureCitation>
)............................ 
<taxonomicName id="4C2C39E6FFA5B44CFB737FA7BACB02C9" ID-CoL="8VRWD" authority="Coiffait and Saiz" authorityName="Coiffait and Saiz" box="[1160,1435,1743,1764]" class="Insecta" family="Staphylinidae" genus="Neohypnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA5B44CFB737FA7BBA202C9" box="[1160,1266,1743,1764]" italics="true" pageId="2" pageNumber="103">Neohypnus</emphasis>
Coiffait and Sáiz
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA5B44CFF6C7F86BACB032E" blockId="2.[151,1436,977,1979]" box="[151,1435,1774,1795]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7F86BACB032E" box="[151,1435,1774,1795]" pageId="2" pageNumber="103">
- Without distinct sclerotized plate in front of prosternum (
<figureCitation id="13175EE0FFA5B44CFCFE7F86BC1D032E" box="[773,845,1774,1795]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="2" pageNumber="103">Fig. 34</figureCitation>
)........................ 7. 
<taxonomicName id="4C2C39E6FFA5B44CFB767F86BACB032E" ID-CoL="8VRBD" authority="(Staphylininae)" baseAuthorityName="Staphylininae" box="[1165,1435,1774,1795]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini (Staphylininae)</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA5B44CFF6C7E65BACB0353" pageId="2" pageNumber="103">
<paragraph id="8B934265FFA5B44CFF6C7E65BACB036C" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7E65BACB036C" pageId="2" pageNumber="103">
7 Superior line of pronotal hypomeron deflexed ventrad to join inferior line before reaching anterior angle of pronotum (
<figureCitation id="13175EE0FFA5B44CFAB47E65BAC4030F" box="[1359,1428,1805,1826]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="2" pageNumber="103">Fig. 36</figureCitation>
)................................................................................................... 8.
</keyLead>
</paragraph>
<paragraph id="8B934265FFA5B44CFF6C7E22BACB0353" blockId="2.[151,1436,977,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7E22BACB0353" pageId="2" pageNumber="103">
- Superior line of pronotal hypomeron not deflexed ventrad, reaching anterior angle of pronotum without joining inferior line (
<figureCitation id="13175EE0FFA5B44CFF137E01BE7E0353" box="[232,302,1897,1918]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="2" pageNumber="103">Fig. 37</figureCitation>
)................................................................................20. Xanthopygina
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA5B44CFF6C7EEFBACC0396" pageId="2" pageNumber="103">
<paragraph id="8B934265FFA5B44CFF6C7EEFBACB03B1" blockId="2.[151,1436,977,1979]" box="[151,1435,1927,1948]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7EEFBACB03B1" box="[151,1435,1927,1948]" pageId="2" pageNumber="103">
8 Ligula distinctly emarginated or bilobed apically (
<figureCitation id="13175EE0FFA5B44CFD3A7EEFBC5903B1" box="[705,777,1927,1948]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="2" pageNumber="103">Fig. 38</figureCitation>
); always large, length 
<quantity id="4CD4EF80FFA5B44CFC1A7EEFBB1703B1" box="[993,1095,1927,1948]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="2" pageNumber="103" unit="mm" value="21.0" valueMax="30.0" valueMin="12.0">12–30 mm</quantity>
................ 9. 
<taxonomicName id="4C2C39E6FFA5B44CFAD87EEFBACB03B1" ID-CoL="8VRDQ" box="[1315,1435,1927,1948]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="subTribe" subTribe="Staphylinina">Staphylinina</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA5B44CFF6C7ECEBACC0396" blockId="2.[151,1436,977,1979]" box="[151,1436,1958,1979]" pageId="2" pageNumber="103">
<keyLead id="30DDE150FFA5B44CFF6C7ECEBACC0396" box="[151,1436,1958,1979]" pageId="2" pageNumber="103">
- Ligula distinctly entire, rounded or slightly sinuate apically (
<figureCitation id="13175EE0FFA5B44CFCDD7ECEBC3E0396" box="[806,878,1958,1979]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="2" pageNumber="103">Fig. 39</figureCitation>
); usually smaller, length 
<quantity id="4CD4EF80FFA5B44CFBA57ECEBBE90396" box="[1118,1209,1958,1979]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="2" pageNumber="103" unit="mm" value="9.5" valueMax="15.0" valueMin="4.0">4–15 mm</quantity>
......12. 
<taxonomicName id="4C2C39E6FFA5B44CFAD67ECEBACC0396" ID-CoL="8VRDC" box="[1325,1436,1958,1979]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C79FFBACC04E6" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C79FFBACC0481" blockId="3.[151,1436,151,1859]" box="[151,1436,151,172]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C79FFBACC0481" box="[151,1436,151,172]" pageId="3" pageNumber="104">
9 Central part of pronotal disc densely punctate (
<figureCitation id="13175EE0FFA4B44DFD5979FFBDAE0481" box="[674,766,151,172]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="3" pageNumber="104">Figs. 8, 9</figureCitation>
, 
<figureCitation id="13175EE0FFA4B44DFCF379FFBC710481" box="[776,801,151,172]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="3" pageNumber="104">40</figureCitation>
)............................... 10. 
<taxonomicName id="4C2C39E6FFA4B44DFB3C79FFBACC0481" ID-CoL="8VRZC" authority="Thomson" authorityName="Thomson" box="[1223,1436,151,172]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA4B44DFB3C79FFBA6B0481" box="[1223,1339,151,172]" italics="true" pageId="3" pageNumber="104">Platydracus</emphasis>
Thomson
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C79DEBACC04E6" blockId="3.[151,1436,151,1859]" box="[151,1436,182,203]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C79DEBACC04E6" box="[151,1436,182,203]" pageId="3" pageNumber="104">
- Central part of pronotal disc glossy and impunctate (
<figureCitation id="13175EE0FFA4B44DFD2579DEBC0104E6" box="[734,849,182,203]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="3" pageNumber="104">Figs. 10, 11</figureCitation>
, 
<figureCitation id="13175EE0FFA4B44DFCA079DEBC2504E6" box="[859,885,182,203]" captionStart="FIGURE 33 – 41" captionStartId="11.[151,250,1195,1217]" captionTargetBox="[151,1436,193,1173]" captionTargetId="figure@11.[151,1436,193,1173]" captionTargetPageId="11" captionText="FIGURE 33 – 41. Prosternum: 33, Eulissus chalybaeus; 34, Creophilus maxillosus; 35, Anotylus sp; 40, Platydracus chrysotrichopterus, 41, Creophilus maxillosus. Pronotal hypomeron: 36, Philonthus longicornis; 37, Oligotergus ogloblini. Ligula: 38, Creophilus maxillosus; 39, Xenopygus analis. Abbreviations: Lg, ligula; IL, inferior line; Pp, plate of pronotum; SL, superior line." httpUri="https://zenodo.org/record/229561/files/figure.png" pageId="3" pageNumber="104">41</figureCitation>
)............................11. 
<taxonomicName id="4C2C39E6FFA4B44DFB0A79DEBACC04E6" ID-CoL="8VRLQ" authority="Leach" authorityName="Leach" box="[1265,1436,182,203]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA4B44DFB0A79DEBA0B04E6" box="[1265,1371,182,203]" italics="true" pageId="3" pageNumber="104">Creophilus</emphasis>
Leach
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C79BDBACC056B" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C79BDBACC0525" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C79BDBACC0525" pageId="3" pageNumber="104">
10 Head, pronotum and elytra of metallic color, abdomen entirely black with distinct white pubescence (usually head and pronotum metallic green to blue and elytra metallic red) (
<figureCitation id="13175EE0FFA4B44DFD35799BBC570525" box="[718,775,243,264]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="3" pageNumber="104">Fig. 8</figureCitation>
)................... 
<taxonomicName id="4C2C39E6FFA4B44DFC0C799BBACC0525" ID-CoL="4JQW5" authority="Scheerpeltz" authorityName="Scheerpeltz" box="[1015,1436,243,264]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="chrysotrichopterus">
<emphasis id="B9589E77FFA4B44DFC0C799BBA750525" box="[1015,1317,243,264]" italics="true" pageId="3" pageNumber="104">Platydracus chrysotrichopterus</emphasis>
Scheerpeltz
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C787ABACC056B" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C787ABACC056B" pageId="3" pageNumber="104">
- Head, pronotum and elytra dull black to piceous black, abdomen black with last visible segment brown to reddish brown (
<figureCitation id="13175EE0FFA4B44DFA8E787ABFBE056B" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="3" pageNumber="104">Fig. 9</figureCitation>
).......................................................................... 
<taxonomicName id="4C2C39E6FFA4B44DFB897859BACC056B" ID-CoL="6VPQP" authority="Curtis" authorityName="Curtis" box="[1138,1436,305,326]" class="Insecta" family="Staphylinidae" genus="Platydracus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="B9589E77FFA4B44DFB897859BA1B056B" box="[1138,1355,305,326]" italics="true" pageId="3" pageNumber="104">Platydracus scabrosus</emphasis>
(Curtis)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7827BACB05ED" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7827BACC05AE" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7827BACC05AE" pageId="3" pageNumber="104">
11 With distinctive white pubescence especially on anterior angles of pronotum, across middle of elytra and on abdominal terga 4 and 5 (
<figureCitation id="13175EE0FFA4B44DFED87806BE3805AE" box="[291,360,366,387]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="3" pageNumber="104">Fig. 10</figureCitation>
).............................................................. 
<taxonomicName id="4C2C39E6FFA4B44DFBAC7806BACC05AE" ID-CoL="ZD3Z" authority="Linnaeus" authorityName="Linnaeus" box="[1111,1436,366,387]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="maxillosus">
<emphasis id="B9589E77FFA4B44DFBAC7806BA7D05AE" box="[1111,1325,366,387]" italics="true" pageId="3" pageNumber="104">Creophilus maxillosus</emphasis>
(Linnaeus)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C78E5BACB05ED" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C78E5BACB05ED" pageId="3" pageNumber="104">
- With extensive pattern of white and golden brown vestiture on anterior angles of pronotum, elytra, and abdomen (
<figureCitation id="13175EE0FFA4B44DFADF78E5BA38058F" box="[1316,1384,397,418]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="3" pageNumber="104">Fig. 11</figureCitation>
)........................................................................... 
<taxonomicName id="4C2C39E6FFA4B44DFBBD78C3BACB05ED" ID-CoL="ZD48" authority="Mannerheim" authorityName="Mannerheim" box="[1094,1435,427,448]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="B9589E77FFA4B44DFBBD78C3BA4B05ED" box="[1094,1307,427,448]" italics="true" pageId="3" pageNumber="104">Creophilus variegatus</emphasis>
Mannerheim
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C78A2BACB069B" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C78A2BACB0677" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C78A2BACB0677" pageId="3" pageNumber="104">
12 Front tarsi with first four segments more or less cylindrical with only regular, unmodified marginal setae on ventral surface (
<figureCitation id="13175EE0FFA4B44DFF137881BE6105D3" box="[232,305,489,510]" captionStart="FIGURE 42 – 49" captionStartId="13.[151,250,1680,1702]" captionTargetBox="[158,1436,509,1655]" captionTargetId="figure@13.[151,1436,509,1655]" captionTargetPageId="13" captionText="FIGURE 42 – 49. Abdomen: 42, Platydracus scabrosus; 43, Anotylus sp. Front tarsus: 44, Belonuchus rufipennis; 45, Philonthus longicornis; 46, Xenopygus analis; 47, Styngetus viduus. Femur: 48, Xenopygus analis; 49, Styngetus viduus. Abbreviation: Fe, femur." httpUri="https://zenodo.org/record/229563/files/figure.png" pageId="3" pageNumber="104">Fig. 44</figureCitation>
): lateral puncture of pronotum bearing long seta situated away from superior line at a distance at least three times as large as diameter of puncture (
<figureCitation id="13175EE0FFA4B44DFDFE7B6FBD1B0631" box="[517,587,519,540]" captionStart="FIGURE 50 – 51" captionStartId="14.[151,250,613,635]" captionTargetBox="[151,1436,157,586]" captionTargetId="figure@14.[151,1436,157,586]" captionTargetPageId="14" captionText="FIGURE 50 – 51. Pronotum (lateral view): 50, Belonuchus rufipennis; 51, Philonthus bonariensis. Abbreviations: Lp, lateral puncture of pronotum." httpUri="https://zenodo.org/record/229564/files/figure.png" pageId="3" pageNumber="104">Fig. 50</figureCitation>
)................................................... 
<taxonomicName id="4C2C39E6FFA4B44DFB457B6FBACC0631" ID-CoL="8VRHV" authority="Nordmann" authorityName="Nordmann" box="[1214,1436,519,540]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA4B44DFB457B6FBA7E0631" box="[1214,1326,519,540]" italics="true" pageId="3" pageNumber="104">Belonuchus</emphasis>
Nordmann
</taxonomicName>
Head, pronotum and two last abdominal segments black, elytra and first four abdominal segments reddish-yellow to rufous (
<figureCitation id="13175EE0FFA4B44DFF137B2DBE770677" box="[232,295,581,602]" captionStart="FIGURE 1 – 12" captionStartId="7.[151,250,1916,1938]" captionTargetBox="[272,1322,208,1883]" captionTargetId="figure@7.[266,1322,195,1892]" captionTargetPageId="7" captionText="FIGURE 1 – 12. Habitus: 1, Atheta sp.; 2, Aleochara bonariensis Lynch; 3, A. puberula Klug; 4, A. signaticollis Fairmaire; 5, Anotylus Thomson; 6, Eulissus chalybaeus Mannerheim; 7, Neohypnus sp.; 8, Platydracus chrysotrichopterus Scheerpeltz; 9, Platydracus scabrosus (Curtis); 10; Creophilus maxillosus (Linnaeus); 11. Creophilus variegatus Mannerheim; 12, Belonuchus rufipennis (Fabricius)." httpUri="https://zenodo.org/record/229558/files/figure.png" pageId="3" pageNumber="104">Fig.12</figureCitation>
)................................................................... 
<taxonomicName id="4C2C39E6FFA4B44DFBAE7B2DBACB0677" ID-CoL="LCB5" authority="Fabricius" authorityName="Fabricius" box="[1109,1435,581,602]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="rufipennis">
<emphasis id="B9589E77FFA4B44DFBAE7B2DBA7C0677" box="[1109,1324,581,602]" italics="true" pageId="3" pageNumber="104">Belonuchus rufipennis</emphasis>
(Fabricius)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7B0CBACB069B" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7B0CBACB069B" pageId="3" pageNumber="104">
- Front tarsi with first four segments more or less flattened dorsoventrally and widened distally, with modified pale (adhesive) setae on ventral surface (
<figureCitation id="13175EE0FFA4B44DFE357BEABD4E06BA" box="[462,542,642,663]" captionStart="FIGURE 42 – 49" captionStartId="13.[151,250,1680,1702]" captionTargetBox="[158,1436,509,1655]" captionTargetId="figure@13.[151,1436,509,1655]" captionTargetPageId="13" captionText="FIGURE 42 – 49. Abdomen: 42, Platydracus scabrosus; 43, Anotylus sp. Front tarsus: 44, Belonuchus rufipennis; 45, Philonthus longicornis; 46, Xenopygus analis; 47, Styngetus viduus. Femur: 48, Xenopygus analis; 49, Styngetus viduus. Abbreviation: Fe, femur." httpUri="https://zenodo.org/record/229563/files/figure.png" pageId="3" pageNumber="104">Figs. 45</figureCitation>
); lateral puncture of pronotum bearing long seta situated close to superior line or at a distance no more than three times the diameter of puncture (
<figureCitation id="13175EE0FFA4B44DFD367BC9BC43069B" box="[717,787,673,694]" captionStart="FIGURE 50 – 51" captionStartId="14.[151,250,613,635]" captionTargetBox="[151,1436,157,586]" captionTargetId="figure@14.[151,1436,157,586]" captionTargetPageId="14" captionText="FIGURE 50 – 51. Pronotum (lateral view): 50, Belonuchus rufipennis; 51, Philonthus bonariensis. Abbreviations: Lp, lateral puncture of pronotum." httpUri="https://zenodo.org/record/229564/files/figure.png" pageId="3" pageNumber="104">Fig. 51</figureCitation>
)................................. 13. 
<taxonomicName id="4C2C39E6FFA4B44DFB237BC9BACB069B" ID-CoL="8VRYR" authority="Stephens" authorityName="Stephens" box="[1240,1435,673,694]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA4B44DFB237BC9BA10069B" box="[1240,1344,673,694]" italics="true" pageId="3" pageNumber="104">Philonthus</emphasis>
Stephens
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7BD7BACB06DE" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7BD7BACB06F9" blockId="3.[151,1436,151,1859]" box="[151,1435,703,724]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7BD7BACB06F9" box="[151,1435,703,724]" pageId="3" pageNumber="104">
13 Pronotum moderately to distinctly narrowed anteriad (
<figureCitation id="13175EE0FFA4B44DFD127BD7BC0E06F9" box="[745,862,703,724]" captionStart="FIGURE 52 – 59" captionStartId="15.[151,250,1416,1438]" captionTargetBox="[151,1436,155,1401]" captionTargetId="figure@15.[151,1436,155,1401]" captionTargetPageId="15" captionText="FIGURE 52 – 59. Head: 52, Philonthus quadraticeps; 53, P. discoideus; 54, P. bicoloristylus. Pronotum: 55, P. bicoloristylus; 56, P. bonariensis; 57, Philonthus quadraticeps. 58, 59. Pronotal hypomeron: 58, Xenopygus analis; 59, Oligotergus ogloblini. Abbreviations: 1 – 5, punctures on dorsal rows of pronotum; Pcp, postcoxal process." httpUri="https://zenodo.org/record/229565/files/figure.png" pageId="3" pageNumber="104">Figs. 55, 56</figureCitation>
)............................................. 15
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7BB6BACB06DE" blockId="3.[151,1436,151,1859]" box="[151,1435,734,755]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7BB6BACB06DE" box="[151,1435,734,755]" pageId="3" pageNumber="104">
- Pronotum parallel-sided (
<figureCitation id="13175EE0FFA4B44DFE2E7BB6BD4A06DE" box="[469,538,734,755]" captionStart="FIGURE 52 – 59" captionStartId="15.[151,250,1416,1438]" captionTargetBox="[151,1436,155,1401]" captionTargetId="figure@15.[151,1436,155,1401]" captionTargetPageId="15" captionText="FIGURE 52 – 59. Head: 52, Philonthus quadraticeps; 53, P. discoideus; 54, P. bicoloristylus. Pronotum: 55, P. bicoloristylus; 56, P. bonariensis; 57, Philonthus quadraticeps. 58, 59. Pronotal hypomeron: 58, Xenopygus analis; 59, Oligotergus ogloblini. Abbreviations: 1 – 5, punctures on dorsal rows of pronotum; Pcp, postcoxal process." httpUri="https://zenodo.org/record/229565/files/figure.png" pageId="3" pageNumber="104">Fig. 57</figureCitation>
)........................................................................ 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7B95BACB0743" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7B95BAC6071C" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7B95BAC6071C" pageId="3" pageNumber="104">
14 Eyes distinctly shorter than temples seen from above (
<figureCitation id="13175EE0FFA4B44DFCFA7B95BC1C073F" box="[769,844,765,786]" captionStart="FIGURE 52 – 59" captionStartId="15.[151,250,1416,1438]" captionTargetBox="[151,1436,155,1401]" captionTargetId="figure@15.[151,1436,155,1401]" captionTargetPageId="15" captionText="FIGURE 52 – 59. Head: 52, Philonthus quadraticeps; 53, P. discoideus; 54, P. bicoloristylus. Pronotum: 55, P. bicoloristylus; 56, P. bonariensis; 57, Philonthus quadraticeps. 58, 59. Pronotal hypomeron: 58, Xenopygus analis; 59, Oligotergus ogloblini. Abbreviations: 1 – 5, punctures on dorsal rows of pronotum; Pcp, postcoxal process." httpUri="https://zenodo.org/record/229565/files/figure.png" pageId="3" pageNumber="104">Fig. 52</figureCitation>
); transverse distance between punctures on elytra distinctly larger than diameters of punctures............................................. 
<taxonomicName id="4C2C39E6FFA4B44DFBBF7A74BAC6071C" ID-CoL="4GBN3" authority="Boheman" authorityName="Boheman" box="[1092,1430,796,817]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis id="B9589E77FFA4B44DFBBF7A74BA63071C" box="[1092,1331,796,817]" italics="true" pageId="3" pageNumber="104">Philonthus quadraticeps</emphasis>
Boheman
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7A52BACB0743" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7A52BACB0743" pageId="3" pageNumber="104">
- Eyes distinctly longer than temples seen from above (
<figureCitation id="13175EE0FFA4B44DFD1F7A52BC650762" box="[740,821,826,847]" captionStart="FIGURE 52 – 59" captionStartId="15.[151,250,1416,1438]" captionTargetBox="[151,1436,155,1401]" captionTargetId="figure@15.[151,1436,155,1401]" captionTargetPageId="15" captionText="FIGURE 52 – 59. Head: 52, Philonthus quadraticeps; 53, P. discoideus; 54, P. bicoloristylus. Pronotum: 55, P. bicoloristylus; 56, P. bonariensis; 57, Philonthus quadraticeps. 58, 59. Pronotal hypomeron: 58, Xenopygus analis; 59, Oligotergus ogloblini. Abbreviations: 1 – 5, punctures on dorsal rows of pronotum; Pcp, postcoxal process." httpUri="https://zenodo.org/record/229565/files/figure.png" pageId="3" pageNumber="104">Figs. 53</figureCitation>
); transverse distance between punctures on elytra about as large as to distinctly smaller than diameters of punctures............................... 
<taxonomicName id="4C2C39E6FFA4B44DFBC37A31BACB0743" ID-CoL="779TK" authority="Gravenhorst" authorityName="Gravenhorst" box="[1080,1435,857,878]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="discoideus">
<emphasis id="B9589E77FFA4B44DFBC37A31BA5E0743" box="[1080,1294,857,878]" italics="true" pageId="3" pageNumber="104">Philonthus discoideus</emphasis>
(Gravenhorst)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7A1FBACC0786" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7A1FBACB07A1" blockId="3.[151,1436,151,1859]" box="[151,1435,887,908]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7A1FBACB07A1" box="[151,1435,887,908]" pageId="3" pageNumber="104">15 Eyes about as long as or shorter than temples seen from above................................................. 16</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7AFEBACC0786" blockId="3.[151,1436,151,1859]" box="[151,1436,918,939]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7AFEBACC0786" box="[151,1436,918,939]" pageId="3" pageNumber="104">
- Eyes distinctly longer than temples seen from above (
<figureCitation id="13175EE0FFA4B44DFD187AFEBC7A0786" box="[739,810,918,939]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 15</figureCitation>
, 
<figureCitation id="13175EE0FFA4B44DFCCF7AFEBC1D0786" box="[820,845,918,939]" captionStart="FIGURE 52 – 59" captionStartId="15.[151,250,1416,1438]" captionTargetBox="[151,1436,155,1401]" captionTargetId="figure@15.[151,1436,155,1401]" captionTargetPageId="15" captionText="FIGURE 52 – 59. Head: 52, Philonthus quadraticeps; 53, P. discoideus; 54, P. bicoloristylus. Pronotum: 55, P. bicoloristylus; 56, P. bonariensis; 57, Philonthus quadraticeps. 58, 59. Pronotal hypomeron: 58, Xenopygus analis; 59, Oligotergus ogloblini. Abbreviations: 1 – 5, punctures on dorsal rows of pronotum; Pcp, postcoxal process." httpUri="https://zenodo.org/record/229565/files/figure.png" pageId="3" pageNumber="104">54</figureCitation>
)................. 
<taxonomicName id="4C2C39E6FFA4B44DFBD67AFEBACC0786" ID-CoL="76XHG" authority="Chani-Posse" authorityName="Chani-Posse" box="[1069,1436,918,939]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="bicoloristylus">
<emphasis id="B9589E77FFA4B44DFBD67AFEBA4E0786" box="[1069,1310,918,939]" italics="true" pageId="3" pageNumber="104">Philonthus bicoloristylus</emphasis>
Chani-Posse
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7ADDBACC07C4" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7ADDBACB07E7" blockId="3.[151,1436,151,1859]" box="[151,1435,949,970]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7ADDBACB07E7" box="[151,1435,949,970]" pageId="3" pageNumber="104">16 Head about as wide as long............................................................................. 17</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7ABCBACC07C4" blockId="3.[151,1436,151,1859]" box="[151,1436,980,1001]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7ABCBACC07C4" box="[151,1436,980,1001]" pageId="3" pageNumber="104">
- Head moderately to distinctly wider than long (
<figureCitation id="13175EE0FFA4B44DFD5D7ABCBDBC07C4" box="[678,748,980,1001]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 16</figureCitation>
)........................... 
<taxonomicName id="4C2C39E6FFA4B44DFBBF7ABCBACC07C4" ID-CoL="779K5" authority="Chani-Posse" authorityName="Chani-Posse" box="[1092,1436,980,1001]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="bruchianus">
<emphasis id="B9589E77FFA4B44DFBBF7ABCBA4D07C4" box="[1092,1309,980,1001]" italics="true" pageId="3" pageNumber="104">Philonthus bruchianus</emphasis>
Chani-Posse
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7A9ABACB000B" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7A9ABACB002A" blockId="3.[151,1436,151,1859]" box="[151,1435,1010,1031]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7A9ABACB002A" box="[151,1435,1010,1031]" pageId="3" pageNumber="104">17 Antennal segments 8–10 distinctly elongate................................................................ 19</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7D79BACB000B" blockId="3.[151,1436,151,1859]" box="[151,1435,1041,1062]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7D79BACB000B" box="[151,1435,1041,1062]" pageId="3" pageNumber="104">- Antennal segments 8–10 not elongate..................................................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7D47BACC008C" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7D47BACB004E" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7D47BACB004E" pageId="3" pageNumber="104">
18 Pronotum with dorsal rows of punctures each with five punctures, rarely with four (if so, only at one row); first segment of hind tarsus distinctly longer than last segment (
<figureCitation id="13175EE0FFA4B44DFD5E7D26BDBB004E" box="[677,747,1102,1123]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 17</figureCitation>
)............................. 
<taxonomicName id="4C2C39E6FFA4B44DFBB47D26BACB004E" ID-CoL="779NH" authority="Erichson" authorityName="Erichson" box="[1103,1435,1102,1123]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="flavolimbatus">
<emphasis id="B9589E77FFA4B44DFBB47D26BA10004E" box="[1103,1344,1102,1123]" italics="true" pageId="3" pageNumber="104">Philonthus flavolimbatus</emphasis>
Erichson
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7D05BACC008C" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7D05BACC008C" pageId="3" pageNumber="104">
- Pronotum with dorsal rows of punctures each with four punctures; first segment of hind tarsus as long as or shorter than last segment (
<figureCitation id="13175EE0FFA4B44DFEC57DE4BED3008C" box="[318,387,1164,1185]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 18</figureCitation>
)............................................................ 
<taxonomicName id="4C2C39E6FFA4B44DFBAB7DE4BACC008C" ID-CoL="76XJQ" authority="Bernhauer" authorityName="Bernhauer" box="[1104,1436,1164,1185]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="bonariensis">
<emphasis id="B9589E77FFA4B44DFBAB7DE4BA62008C" box="[1104,1330,1164,1185]" italics="true" pageId="3" pageNumber="104">Philonthus bonariensis</emphasis>
Bernhauer
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7DC2BACB0136" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7DC2BACB00F3" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7DC2BACB00F3" pageId="3" pageNumber="104">
19 Pronotum with dorsal rows of punctures each with five punctures; apex of male tergum 10 not emarginate medioapically (
<figureCitation id="13175EE0FFA4B44DFA8E7DC2BFAA00F3" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 19</figureCitation>
)........................................................................ 
<taxonomicName id="4C2C39E6FFA4B44DFB9F7DA1BACB00F3" ID-CoL="4G9X6" authority="Stephens" authorityName="Stephens" box="[1124,1435,1225,1246]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B9589E77FFA4B44DFB9F7DA1BA6F00F3" box="[1124,1343,1225,1246]" italics="true" pageId="3" pageNumber="104">Philonthus longicornis</emphasis>
Stephens
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7D8FBACB0136" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7D8FBACB0136" pageId="3" pageNumber="104">
- Pronotum with dorsal rows of punctures each with four punctures; apex of male tergum 10 emarginate medio-apically (
<figureCitation id="13175EE0FFA4B44DFAB47D8FBAC400D1" box="[1359,1428,1255,1276]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 20</figureCitation>
)................................................................................ 
<taxonomicName id="4C2C39E6FFA4B44DFB5F7C6EBACB0136" ID-CoL="779HV" authority="Herman" authorityName="Herman" box="[1188,1435,1286,1307]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="B9589E77FFA4B44DFB5F7C6EBA180136" box="[1188,1352,1286,1307]" italics="true" pageId="3" pageNumber="104">Philonthus argus</emphasis>
Herman
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7C4DBACB01FE" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7C4DBACB01BB" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7C4DBACB01BB" pageId="3" pageNumber="104">
20 First four segments of front tarsus slightly dilated, much narrower than apex of protibia and with few translucent tenent setae on either side of ventral apex (
<figureCitation id="13175EE0FFA4B44DFDFE7C2CBD1C0174" box="[517,588,1348,1369]" captionStart="FIGURE 42 – 49" captionStartId="13.[151,250,1680,1702]" captionTargetBox="[158,1436,509,1655]" captionTargetId="figure@13.[151,1436,509,1655]" captionTargetPageId="13" captionText="FIGURE 42 – 49. Abdomen: 42, Platydracus scabrosus; 43, Anotylus sp. Front tarsus: 44, Belonuchus rufipennis; 45, Philonthus longicornis; 46, Xenopygus analis; 47, Styngetus viduus. Femur: 48, Xenopygus analis; 49, Styngetus viduus. Abbreviation: Fe, femur." httpUri="https://zenodo.org/record/229563/files/figure.png" pageId="3" pageNumber="104">Fig. 47</figureCitation>
); hind femur of male with posterior margin crenulate (
<figureCitation id="13175EE0FFA4B44DFBAD7C2CBBCC0174" box="[1110,1180,1348,1369]" captionStart="FIGURE 42 – 49" captionStartId="13.[151,250,1680,1702]" captionTargetBox="[158,1436,509,1655]" captionTargetId="figure@13.[151,1436,509,1655]" captionTargetPageId="13" captionText="FIGURE 42 – 49. Abdomen: 42, Platydracus scabrosus; 43, Anotylus sp. Front tarsus: 44, Belonuchus rufipennis; 45, Philonthus longicornis; 46, Xenopygus analis; 47, Styngetus viduus. Femur: 48, Xenopygus analis; 49, Styngetus viduus. Abbreviation: Fe, femur." httpUri="https://zenodo.org/record/229563/files/figure.png" pageId="3" pageNumber="104">Fig. 49</figureCitation>
)....... 
<taxonomicName id="4C2C39E6FFA4B44DFAF97C2CBACC0174" ID-CoL="8X7WD" authority="Sharp" authorityName="Sharp" box="[1282,1436,1348,1369]" class="Insecta" family="Staphylinidae" genus="Styngetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA4B44DFAF97C2CBA0D0174" box="[1282,1373,1348,1369]" italics="true" pageId="3" pageNumber="104">Styngetus</emphasis>
Sharp
</taxonomicName>
Head and pronotum glossy black and densely punctate; elytra metallic blue, abdomen black with last visible segment brown to reddish brown (
<figureCitation id="13175EE0FFA4B44DFE077CE9BD1001BB" box="[508,576,1409,1430]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 21</figureCitation>
)................................................. 
<taxonomicName id="4C2C39E6FFA4B44DFB777CE9BACB01BB" ID-CoL="53BDK" authority="Erichson" authorityName="Erichson" box="[1164,1435,1409,1430]" class="Insecta" family="Staphylinidae" genus="Styngetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="viduus">
<emphasis id="B9589E77FFA4B44DFB777CE9BA7F01BB" box="[1164,1327,1409,1430]" italics="true" pageId="3" pageNumber="104">Styngetus viduus</emphasis>
(Erichson)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7CF7BACB01FE" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7CF7BACB01FE" pageId="3" pageNumber="104">
- First four segments of front tarsus distinctly dilated, about as wide as or wider than apex of protibia and with dense pad of translucent tenent setae below (
<figureCitation id="13175EE0FFA4B44DFDCD7CD6BD2801FE" box="[566,632,1470,1491]" captionStart="FIGURE 42 – 49" captionStartId="13.[151,250,1680,1702]" captionTargetBox="[158,1436,509,1655]" captionTargetId="figure@13.[151,1436,509,1655]" captionTargetPageId="13" captionText="FIGURE 42 – 49. Abdomen: 42, Platydracus scabrosus; 43, Anotylus sp. Front tarsus: 44, Belonuchus rufipennis; 45, Philonthus longicornis; 46, Xenopygus analis; 47, Styngetus viduus. Femur: 48, Xenopygus analis; 49, Styngetus viduus. Abbreviation: Fe, femur." httpUri="https://zenodo.org/record/229563/files/figure.png" pageId="3" pageNumber="104">Fig.46</figureCitation>
); hind femur of male not crenulate (
<figureCitation id="13175EE0FFA4B44DFC2A7CD6BB4701FE" box="[977,1047,1470,1491]" captionStart="FIGURE 42 – 49" captionStartId="13.[151,250,1680,1702]" captionTargetBox="[158,1436,509,1655]" captionTargetId="figure@13.[151,1436,509,1655]" captionTargetPageId="13" captionText="FIGURE 42 – 49. Abdomen: 42, Platydracus scabrosus; 43, Anotylus sp. Front tarsus: 44, Belonuchus rufipennis; 45, Philonthus longicornis; 46, Xenopygus analis; 47, Styngetus viduus. Femur: 48, Xenopygus analis; 49, Styngetus viduus. Abbreviation: Fe, femur." httpUri="https://zenodo.org/record/229563/files/figure.png" pageId="3" pageNumber="104">Fig. 48</figureCitation>
)............................. 21
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7CB5BACB0263" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7CB5BACB0202" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7CB5BACB0202" pageId="3" pageNumber="104">
21 Eyes about as long as or shorter than temples seen from above; pronotal hypomeron with a small translucent postcoxal process very close behind coxal articulation (
<figureCitation id="13175EE0FFA4B44DFDBF7C94BDDA023C" box="[580,650,1532,1553]" captionStart="FIGURE 52 – 59" captionStartId="15.[151,250,1416,1438]" captionTargetBox="[151,1436,155,1401]" captionTargetId="figure@15.[151,1436,155,1401]" captionTargetPageId="15" captionText="FIGURE 52 – 59. Head: 52, Philonthus quadraticeps; 53, P. discoideus; 54, P. bicoloristylus. Pronotum: 55, P. bicoloristylus; 56, P. bonariensis; 57, Philonthus quadraticeps. 58, 59. Pronotal hypomeron: 58, Xenopygus analis; 59, Oligotergus ogloblini. Abbreviations: 1 – 5, punctures on dorsal rows of pronotum; Pcp, postcoxal process." httpUri="https://zenodo.org/record/229565/files/figure.png" pageId="3" pageNumber="104">Fig. 58</figureCitation>
)................................................ 
<taxonomicName id="4C2C39E6FFA4B44DFB297C94BACC023C" ID-CoL="8X7Y3" authority="Bernhuer" authorityName="Bernhuer" box="[1234,1436,1532,1553]" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA4B44DFB297C94BA6B023C" box="[1234,1339,1532,1553]" italics="true" pageId="3" pageNumber="104">Xenopygus</emphasis>
Bernhuer
</taxonomicName>
Head, pronotum and elytra of metallic color (
<figureCitation id="13175EE0FFA4B44DFD687F72BD890202" box="[659,729,1562,1583]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 22</figureCitation>
)................................... 
<taxonomicName id="4C2C39E6FFA4B44DFB7D7F72BACB0202" ID-CoL="5CBCW" authority="Erichson" authorityName="Erichson" box="[1158,1435,1562,1583]" class="Insecta" family="Staphylinidae" genus="Xenopygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B9589E77FFA4B44DFB7D7F72BA600202" box="[1158,1328,1562,1583]" italics="true" pageId="3" pageNumber="104">Xenopygus analis</emphasis>
(Erichson)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7F51BACB0263" blockId="3.[151,1436,151,1859]" box="[151,1435,1593,1614]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7F51BACB0263" box="[151,1435,1593,1614]" pageId="3" pageNumber="104">
- Eyes distinctly longer than temples seen from above; pronotal hypomeron without postcoxal process (
<figureCitation id="13175EE0FFA4B44DFB4A7F51BBA70263" box="[1201,1271,1593,1614]" captionStart="FIGURE 52 – 59" captionStartId="15.[151,250,1416,1438]" captionTargetBox="[151,1436,155,1401]" captionTargetId="figure@15.[151,1436,155,1401]" captionTargetPageId="15" captionText="FIGURE 52 – 59. Head: 52, Philonthus quadraticeps; 53, P. discoideus; 54, P. bicoloristylus. Pronotum: 55, P. bicoloristylus; 56, P. bonariensis; 57, Philonthus quadraticeps. 58, 59. Pronotal hypomeron: 58, Xenopygus analis; 59, Oligotergus ogloblini. Abbreviations: 1 – 5, punctures on dorsal rows of pronotum; Pcp, postcoxal process." httpUri="https://zenodo.org/record/229565/files/figure.png" pageId="3" pageNumber="104">Fig. 59</figureCitation>
)........... 22
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D85AC0FFA4B44DFF6C7F3FBACC036E" pageId="3" pageNumber="104">
<paragraph id="8B934265FFA4B44DFF6C7F3FBACB02E4" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7F3FBACB02E4" pageId="3" pageNumber="104">
22 Antenna with segments 7–10 transverse; head, especially the frons and vertex, with dense contiguous punctures............................................................................................... 
<taxonomicName id="4C2C39E6FFA4B44DFB2D7F1EBACC02A6" ID-CoL="8X7M6" authority="Blackwelder" authorityName="Blackwelder" box="[1238,1436,1654,1675]" class="Insecta" family="Staphylinidae" genus="Nordus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA4B44DFB2D7F1EBA4C02A6" box="[1238,1308,1654,1675]" italics="true" pageId="3" pageNumber="104">Nordus</emphasis>
Blackwelder
</taxonomicName>
Head, pronotum, elytra and scutellum golden orange, antennal segments yellow, abdomen black except apical margin of segment 7 and entire segment 8 golden orange (
<figureCitation id="13175EE0FFA4B44DFD707FDCBD8102E4" box="[651,721,1716,1737]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 23</figureCitation>
).................................... 
<taxonomicName id="4C2C39E6FFA4B44DFB7E7FDCBACB02E4" ID-CoL="47Q8X" authority="Chatzimanolis" authorityName="Chatzimanolis" box="[1157,1435,1716,1737]" class="Insecta" family="Staphylinidae" genus="Nordus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="elytisi">
<emphasis id="B9589E77FFA4B44DFB7E7FDCBA5A02E4" box="[1157,1290,1716,1737]" italics="true" pageId="3" pageNumber="104">Nordus elytisi</emphasis>
Chatzimanolis
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B934265FFA4B44DFF6C7FBABACC036E" blockId="3.[151,1436,151,1859]" pageId="3" pageNumber="104">
<keyLead id="30DDE150FFA4B44DFF6C7FBABACC036E" pageId="3" pageNumber="104">
- Antenna with segments 7–10 elongate or conical; head with frons and vertex only sparsely punctate, but often with dense microsculpture.......................................................................... 
<taxonomicName id="4C2C39E6FFA4B44DFB137F99BACC032B" ID-CoL="8X7MG" authority="Bierig" authorityName="Bierig" box="[1256,1436,1777,1798]" class="Insecta" family="Staphylinidae" genus="Oligotergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B9589E77FFA4B44DFB137F99BA0A032B" box="[1256,1370,1777,1798]" italics="true" pageId="3" pageNumber="104">Oligotergus</emphasis>
Bierig
</taxonomicName>
Head, pronotum and elytra black to piceous black, abdominal segments black to piceous black with apical margins of segments 3–6 and entire segments 7 and 8 brown to reddish brown (
<figureCitation id="13175EE0FFA4B44DFC887E46BCE9036E" box="[883,953,1838,1859]" captionStart="FIGURE 13 – 24" captionStartId="8.[151,250,1911,1933]" captionTargetBox="[257,1354,196,1875]" captionTargetId="figure@8.[237,1354,193,1895]" captionTargetPageId="8" captionText="FIGURE 13 – 24. Habitus: 13, Philonthus quadraticeps Boheman; 14, P. discoideus (Gravenhorst); 15, P. bicoloristylus Chani- Posse; 16, P. bruchianus Chani-Posse; 17, P. flavolimbatus Erichson; 18, P. bonariensis Bernhauer; 19, P. longicornis Stephens; 20, P. argus Herman; 21, Styngetus viduus (Erichson); 22, Xenopygus analis (Erichson); 23, Nordus elytisi Chatzimanolis; 24, Oligotergus ogloblini Bernhauer." httpUri="https://zenodo.org/record/229559/files/figure.png" pageId="3" pageNumber="104">Fig. 24</figureCitation>
)............. 
<taxonomicName id="4C2C39E6FFA4B44DFB987E46BACC036E" ID-CoL="497PT" authority="Bernhauer" authorityName="Bernhauer" box="[1123,1436,1838,1859]" class="Insecta" family="Staphylinidae" genus="Oligotergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="ogloblini">
<emphasis id="B9589E77FFA4B44DFB987E46BA62036E" box="[1123,1330,1838,1859]" italics="true" pageId="3" pageNumber="104">Oligotergus ogloblini</emphasis>
Bernhauer
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>