157 lines
18 KiB
XML
157 lines
18 KiB
XML
<document id="C9CB1502A67A2EAFA88B3B7D0DD4DDE3" ID-CLB-Dataset="198290" ID-DOI="10.1080/00222933.2022.2136017" ID-GBIF-Dataset="c9288626-198f-4b45-b923-f134cc0df48a" ID-ISSN="1464-5262" ID-Zenodo-Dep="7380681" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669805002707" checkinUser="plazi" docAuthor="Hyman, Isabel T. & Köhler, Frank" docDate="2022" docId="CE7A781DFFFEFFE4DF32FB24FF7CC92F" docLanguage="en" docName="JNATHIST.56.41-44.1727-1799.pdf" docOrigin="Journal of Natural History 56 (41 - 44)" docSource="http://dx.doi.org/10.1080/00222933.2022.2136017" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Geminitor Hyman & Köhler 2022, gen. nov." docType="treatment" docUuid="A2434D25-ADF5-493F-A17C-304B5F3CC182" docUuidSource="ZooBank" docVersion="8" lastPageNumber="1758" masterDocId="32430065FFE0FFFBDD24FFD1FF84CC7C" masterDocTitle="Homoplasy in shells discombobulated the taxonomy: revision of the larger helicarionid land snails of northern Queensland, Australia (Stylommatophora: Helicarionidae)" masterLastPageNumber="1799" masterPageNumber="1727" pageNumber="1757" updateTime="1699412228230" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="2C5482559585D1BEAEF1E6C813C7F178" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="42A8FA6D5414B8144FA794D3A0B778D4">
|
||
<mods:title id="0386A63816BC3EEEE4FE72E27E8BEA68">Homoplasy in shells discombobulated the taxonomy: revision of the larger helicarionid land snails of northern Queensland, Australia (Stylommatophora: Helicarionidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="9DA24396CA800C35595B39E27837F34C" type="personal">
|
||
<mods:role id="35B377DFEA045EAD03F7FF72AFD254C0">
|
||
<mods:roleTerm id="DEF27F237208E1BEB22CEE5B5A4594EE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1AC232C004D5D208F41BB0A51745B8BA">Hyman, Isabel T.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="42855B34EC5DFB10FDE1A2C380806BAD" type="personal">
|
||
<mods:role id="DCE822331328F9DAB0B59C8F373BD23F">
|
||
<mods:roleTerm id="061B43683C5F3B2EA98F3A1AABD2C216">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C0BF4B7DBB835FD4853E137816BA42F0">Köhler, Frank</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3B8ECE34ACA70C1DE41040159F170A59">text</mods:typeOfResource>
|
||
<mods:relatedItem id="D5F6FDBD3642AB711965D73E76F91B65" type="host">
|
||
<mods:titleInfo id="F4367698ADD85BB9DDDDDCE6C9C3D301">
|
||
<mods:title id="81AF64A0912C07E8CCF72ECEB56F70A2">Journal of Natural History</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="567A5F1C25AEB3F7F14993A02B909A5F">
|
||
<mods:date id="937B7AED3846200A24B102A7835F72B9">2022</mods:date>
|
||
<mods:detail id="6E7B1B2EE7D533A694229825BA2895A8" type="pubDate">
|
||
<mods:number id="D5FEF9EB9CE8151B48F16F7FDBE6B9A6">2022-11-23</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CAEBBBC351886C9B33BAC9B5D1DFC0D0" type="volume">
|
||
<mods:number id="4C33228E14201597D7507BF0E195BD5F">56</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="37069C6040E96C565C4A9F047A64CC0A" type="issue">
|
||
<mods:number id="8C587D89454C78F28819DE408C89AE13">41 - 44</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="22ABAD65E1297419CCC390EB8884BC0D" unit="page">
|
||
<mods:start id="E979EF21C203854968ACFF3B520721A4">1727</mods:start>
|
||
<mods:end id="899E38B9046B25705801D0B0E5E351FB">1799</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="5CA7BBE6A03C4408ABBEA71F2EA14B4F">
|
||
<mods:url id="6AEC5DC004722836B45956D787C3900E">http://dx.doi.org/10.1080/00222933.2022.2136017</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="5EC95B342FBC0A7F508301A0BBD7B28B">journal article</mods:classification>
|
||
<mods:identifier id="2172C1AF630B8F059A9A9793DDCCEFEC" type="CLB-Dataset">198290</mods:identifier>
|
||
<mods:identifier id="241A348C23C39B7721B5E5FB3A469E19" type="DOI">10.1080/00222933.2022.2136017</mods:identifier>
|
||
<mods:identifier id="2E8A312CA3447F1ED56469C4C1D51FB1" type="GBIF-Dataset">c9288626-198f-4b45-b923-f134cc0df48a</mods:identifier>
|
||
<mods:identifier id="EA44A4C2CA0406ACCD889024B94FA84F" type="ISSN">1464-5262</mods:identifier>
|
||
<mods:identifier id="47DB30431460E0C321EDED8A31B45AB4" type="Zenodo-Dep">7380681</mods:identifier>
|
||
<mods:identifier id="4A24E7FC27DDAB47177937FFC5503190" type="ZooBank">83362BEC-6E6B-4B65-AC1E-F49762C744B8</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CE7A781DFFFEFFE4DF32FB24FF7CC92F" ID-DOI="http://doi.org/10.5281/zenodo.7383416" ID-GBIF-Taxon="205276711" ID-Zenodo-Dep="7383416" LSID="urn:lsid:zoobank.org:act:A2434D25-ADF5-493F-A17C-304B5F3CC182" httpUri="http://treatment.plazi.org/id/CE7A781DFFFEFFE4DF32FB24FF7CC92F" lastPageId="31" lastPageNumber="1758" pageId="30" pageNumber="1757">
|
||
<subSubSection id="0EC99A80FFFEFFE5DF32FB24FC89C973" box="[534,781,1269,1295]" pageId="30" pageNumber="1757" type="nomenclature">
|
||
<paragraph id="466CC90BFFFEFFE5DF32FB24FC89C973" blockId="30.[534,781,1269,1295]" box="[534,781,1269,1295]" pageId="30" pageNumber="1757">
|
||
<heading id="1D247E67FFFEFFE5DF32FB24FC89C973" bold="true" box="[534,781,1269,1295]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="1757" reason="3">
|
||
<emphasis id="74A71519FFFEFFE5DF32FB24FC89C973" bold="true" box="[534,781,1269,1295]" pageId="30" pageNumber="1757">
|
||
<taxonomicName id="81D3B288FFFEFFE5DF32FB24FD15C973" ID-CoL="c4c018e1-5e32-4389-a605-9446671daa5b" authority="Hyman & Köhler, 2022" authorityName="Hyman & Köhler" authorityYear="2022" box="[534,657,1269,1295]" class="Gastropoda" family="Helicarionidae" genus="Geminitor" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="1757" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||
<emphasis id="74A71519FFFEFFE5DF32FB24FD15C973" bold="true" box="[534,657,1269,1295]" italics="true" pageId="30" pageNumber="1757">Geminitor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6F94A862FFFEFFE5DFBCFB24FC89C973" box="[664,781,1269,1295]" pageId="30" pageNumber="1757" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EC99A80FFFEFFE5DD84FAF8FD09C93E" box="[160,653,1321,1346]" pageId="30" pageNumber="1757" type="type_taxon">
|
||
<paragraph id="466CC90BFFFEFFE5DD84FAF8FD09C93E" blockId="30.[160,653,1321,1346]" box="[160,653,1321,1346]" pageId="30" pageNumber="1757">
|
||
<typeStatus id="996877A9FFFEFFE5DD84FAF8FF52C93E" box="[160,214,1321,1346]" pageId="30" pageNumber="1757">Type</typeStatus>
|
||
species
|
||
<taxonomicName id="81D3B288FFFEFFE5DC13FAF8FD03C93E" ID-CoL="6VXD7" authority="Stanisic, 2010" authorityName="Stanisic" authorityYear="2010" box="[311,647,1321,1346]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="1757" phylum="Mollusca" rank="species" species="laura">
|
||
<emphasis id="74A71519FFFEFFE5DC13FAF8FE63C93E" box="[311,487,1321,1346]" italics="true" pageId="30" pageNumber="1757">Pravonitor laura</emphasis>
|
||
<bibRefCitation id="2242B4FAFFFEFFE5DCCBFAF8FD03C93E" author="Stanisic J & Shea M & Potter D & Griffiths O" box="[495,647,1321,1346]" pageId="30" pageNumber="1757" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EC99A80FFFEFFE5DD84FABFFD7EC9B1" pageId="30" pageNumber="1757" type="etymology">
|
||
<paragraph id="466CC90BFFFEFFE5DD84FABFFE99C9F4" blockId="30.[160,1155,1390,1485]" box="[160,285,1390,1416]" pageId="30" pageNumber="1757">
|
||
<emphasis id="74A71519FFFEFFE5DD84FABFFE99C9F4" bold="true" box="[160,285,1390,1416]" italics="true" pageId="30" pageNumber="1757">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="466CC90BFFFEFFE5DD84FA43FD7EC9B1" blockId="30.[160,1155,1390,1485]" pageId="30" pageNumber="1757">
|
||
From
|
||
<emphasis id="74A71519FFFEFFE5DDC0FA43FEC4C9D7" box="[228,320,1426,1451]" italics="true" pageId="30" pageNumber="1757">geminus</emphasis>
|
||
(Latin = twin-born, twin), referring to the double-chambered penis, and
|
||
<taxonomicName id="81D3B288FFFEFFE5DD84FA65FF52C9B1" baseAuthorityName="Hyman and Kohler" baseAuthorityYear="2018" box="[160,214,1460,1485]" class="Gastropoda" family="Helicarionidae" genus="Nitor" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="1757" phylum="Mollusca" rank="genus">
|
||
<emphasis id="74A71519FFFEFFE5DD84FA65FF52C9B1" box="[160,214,1460,1485]" italics="true" pageId="30" pageNumber="1757">Nitor</emphasis>
|
||
</taxonomicName>
|
||
, a helicarionid snail genus; of masculine gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EC99A80FFFEFFE4DD84FA28FF5BCC91" lastPageId="31" lastPageNumber="1758" pageId="30" pageNumber="1757" type="diagnosis">
|
||
<paragraph id="466CC90BFFFEFFE5DD84FA28FE19CA6F" blockId="30.[160,1156,1529,1728]" box="[160,413,1529,1555]" pageId="30" pageNumber="1757">
|
||
<emphasis id="74A71519FFFEFFE5DD84FA28FE19CA6F" bold="true" box="[160,413,1529,1555]" italics="true" pageId="30" pageNumber="1757">Differential diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="466CC90BFFFEFFE4DD84F9CDFF5BCC91" blockId="30.[160,1156,1529,1728]" lastBlockId="31.[160,1155,142,237]" lastPageId="31" lastPageNumber="1758" pageId="30" pageNumber="1757">
|
||
<emphasis id="74A71519FFFEFFE5DD84F9CDFE27CA4A" bold="true" box="[160,419,1564,1590]" italics="true" pageId="30" pageNumber="1757">External appearance.</emphasis>
|
||
Shell small to medium-sized (SW
|
||
<quantity id="812B64EEFFFEFFE5DE66F9CCFC5FCA4A" box="[834,987,1565,1590]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" metricValueMax="1.46" metricValueMin="0.6599999999999999" pageId="30" pageNumber="1757" unit="mm" value="10.6" valueMax="14.6" valueMin="6.6">6.6–14.6 mm</quantity>
|
||
), pale gold to amber, often paler on the underside, glossy, 4.8–6.3 whorls, discoidal to subdiscoidal with a low to moderately elevated spire; whorls rounded; umbilicus narrow, sealed with a mucous plug; lip slightly reflected over umbilicus. Protoconch with moderately coarse spiral grooves, becoming finer and more crowded on teleoconch. Body colour pale brown to white, often with a grey tail tip and black eyestalks. Mantle lobes small; shell lappets moderately long, triangular, wide at base, tapering to a point. Caudal horn moderately large.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EC99A80FFFFFFE4DD84FEF1FF69CEC4" pageId="31" pageNumber="1758" type="description">
|
||
<paragraph id="466CC90BFFFFFFE4DD84FEF1FF69CEC4" blockId="31.[160,1156,288,696]" pageId="31" pageNumber="1758">
|
||
<emphasis id="74A71519FFFFFFE4DD84FEF1FEFDCD46" bold="true" box="[160,377,288,314]" italics="true" pageId="31" pageNumber="1758">Genital anatomy.</emphasis>
|
||
Ovotestis embedded in digestive gland. Carrefour visible; talon usually embedded in albumen gland. Spermoviduct not folded. Free oviduct long, with elongate capsular gland; free oviduct internally smooth. Vagina very short. Bursa copulatrix moderately short, inserted on vagina, base of duct usually swollen, then narrowing; bursa elongate. Penis with two chambers, distal chamber with longitudinal pilasters and ridges, upper chamber sculptured with rounded, transverse plates. Verge-like structure sometimes present between penial chambers. Penis tunica attached by muscle fibres to middle of epiphallus; epiphallus enters penis through simple pore or small verge; epiphallus 2 longer than or shorter than epiphallus 1; epiphallus 2 broader than epiphallus 1; epiphallic caecum moderately long to long, penial retractor muscle attached to base of caecum. Long, slender epiphallic flagellum with axial filament present; no internal cryptae visible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EC99A80FFFFFFE4DD84FD3AFF7CC92F" pageId="31" pageNumber="1758" type="discussion">
|
||
<paragraph id="466CC90BFFFFFFE4DD84FD3AFE82CF79" blockId="31.[160,1156,747,1363]" box="[160,262,747,773]" pageId="31" pageNumber="1758">
|
||
<emphasis id="74A71519FFFFFFE4DD84FD3AFE82CF79" bold="true" box="[160,262,747,773]" italics="true" pageId="31" pageNumber="1758">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="466CC90BFFFFFFE4DD84FCDEFE42CFEC" blockId="31.[160,1156,747,1363]" pageId="31" pageNumber="1758">
|
||
<taxonomicName id="81D3B288FFFFFFE4DD84FCDEFE8FCF54" authorityName="Hyman & Köhler" authorityYear="2022" box="[160,267,783,808]" class="Gastropoda" family="Helicarionidae" genus="Geminitor" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="1758" phylum="Mollusca" rank="genus">
|
||
<emphasis id="74A71519FFFFFFE4DD84FCDEFE8FCF54" box="[160,267,783,808]" italics="true" pageId="31" pageNumber="1758">Geminitor</emphasis>
|
||
</taxonomicName>
|
||
is introduced for small to medium-sized snails with a relatively pale, unbanded, discoidal to subdiscoidal shell and a two-chambered penis. Two species are removed from
|
||
<taxonomicName id="81D3B288FFFFFFE4DD84FC85FE95CF11" box="[160,273,852,877]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="1758" phylum="Mollusca" rank="genus">
|
||
<emphasis id="74A71519FFFFFFE4DD84FC85FE95CF11" box="[160,273,852,877]" italics="true" pageId="31" pageNumber="1758">Pravonitor</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="74A71519FFFFFFE4DC06FC85FE6ACF11" box="[290,494,852,877]" italics="true" pageId="31" pageNumber="1758">
|
||
<taxonomicName id="81D3B288FFFFFFE4DC06FC85FEF8CF11" authorityName="Stanisic" authorityYear="2010" box="[290,380,852,877]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="1758" phylum="Mollusca" rank="species" species="laura">G. laura</taxonomicName>
|
||
,
|
||
<taxonomicName id="81D3B288FFFFFFE4DCADFC85FE6ACF11" box="[393,494,852,877]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="1758" phylum="Mollusca" rank="species" species="villaris">G. villaris</taxonomicName>
|
||
</emphasis>
|
||
) and three species are described as new (
|
||
<emphasis id="74A71519FFFFFFE4DEF2FC85FE3DCFEC" italics="true" pageId="31" pageNumber="1758">
|
||
<taxonomicName id="81D3B288FFFFFFE4DEF2FC85FBDBCF11" authorityName="Hyman & Köhler" authorityYear="2022" box="[982,1119,852,877]" class="Gastropoda" family="Helicarionidae" genus="Geminitor" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="1758" phylum="Mollusca" rank="species" species="kullaensis">G. kullaensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="81D3B288FFFFFFE4D949FC85FEABCFEC" authorityName="Hyman & Köhler" authorityYear="2022" class="Gastropoda" family="Helicarionidae" genus="Geminitor" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="1758" phylum="Mollusca" rank="species" species="wenlockensis">G. wenlockensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="81D3B288FFFFFFE4DC1EFCA6FE3DCFEC" authorityName="Hyman & Köhler" authorityYear="2022" box="[314,441,887,912]" class="Gastropoda" family="Helicarionidae" genus="Geminitor" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="1758" phylum="Mollusca" rank="species" species="macveae">G. macveae</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="466CC90BFFFFFFE4DDE4FC4BFDE7C897" blockId="31.[160,1156,747,1363]" pageId="31" pageNumber="1758">
|
||
<taxonomicName id="81D3B288FFFFFFE4DDE4FC4BFEAFCFCF" authorityName="Hyman & Köhler" authorityYear="2022" box="[192,299,922,947]" class="Gastropoda" family="Helicarionidae" genus="Geminitor" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="1758" phylum="Mollusca" rank="genus">
|
||
<emphasis id="74A71519FFFFFFE4DDE4FC4BFEAFCFCF" box="[192,299,922,947]" italics="true" pageId="31" pageNumber="1758">Geminitor</emphasis>
|
||
</taxonomicName>
|
||
can be characterised by its small to medium-sized, pale gold to amber, glossy shell, often paler on the underside, with rather coarse microspiral sculpture on the protoconch (becoming finer on the teleoconch). The shell is discoidal to subdiscoidal in shape, with a low to moderately elevated spire and a rounded whorl profile. Anatomically, it is distinguished particularly by its two-chambered penis, often with a beehive-shaped verge leading into the second chamber, and/or a series of overlapping plates around the opening between the two chambers. Other key features include the long, slender reproductive system with visible carrefour, the very short vagina, the broad epiphallus 2 compared to epiphallus 1, the long caecum with basal attachment of the penial retractor muscle, and the long, slender flagellum.
|
||
</paragraph>
|
||
<paragraph id="466CC90BFFFFFFE4DDE4FB25FF7CC92F" blockId="31.[160,1156,747,1363]" pageId="31" pageNumber="1758">
|
||
<materialsCitation id="F6BBC356FFFFFFE4DDE4FB25FE8EC94C" ID-GBIF-Occurrence="3978770311" location="Cairns" pageId="31" pageNumber="1758" specimenCount="1" stateProvince="York">
|
||
This genus is restricted to Cape
|
||
<collectingRegion id="841707E9FFFFFFE4DF39FB25FDCBC971" box="[541,591,1268,1293]" country="United Kingdom" name="York" pageId="31" pageNumber="1758">York</collectingRegion>
|
||
Peninsula, being found from just north of
|
||
<location id="430C9FD0FFFFFFE4D906FB25FBE3C971" LSID="urn:lsid:plazi:treatment:CE7A781DFFFEFFE4DF32FB24FF7CC92F:430C9FD0FFFFFFE4D906FB25FBE3C971" box="[1058,1127,1268,1293]" name="Cairns" pageId="31" pageNumber="1758" stateProvince="York">Cairns</location>
|
||
to Bramwell
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F6BBC356FFFFFFE4DC37FAC6FF70C92F" ID-GBIF-Occurrence="3978770304" location="This area" pageId="31" pageNumber="1758" specimenCount="1" stateProvince="York">
|
||
<location id="430C9FD0FFFFFFE4DC37FAC6FEF2C94C" LSID="urn:lsid:plazi:treatment:CE7A781DFFFEFFE4DF32FB24FF7CC92F:430C9FD0FFFFFFE4DC37FAC6FEF2C94C" box="[275,374,1303,1328]" name="This area" pageId="31" pageNumber="1758" stateProvince="York">This area</location>
|
||
is under-collected and it is likely that there are additional undescribed species
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |