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<document id="D559E8E334D83A49215C72FEEF9AECF7" ID-CLB-Dataset="198290" ID-DOI="10.1080/00222933.2022.2136017" ID-GBIF-Dataset="c9288626-198f-4b45-b923-f134cc0df48a" ID-ISSN="1464-5262" ID-Zenodo-Dep="7380681" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669805002707" checkinUser="plazi" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2022" docId="CE7A781DFFDFFFBBDF25FB24FC8EC9DB" docLanguage="en" docName="JNATHIST.56.41-44.1727-1799.pdf" docOrigin="Journal of Natural History 56 (41 - 44)" docSource="http://dx.doi.org/10.1080/00222933.2022.2136017" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Elatonitor Stanisic 2010" docType="treatment" docVersion="8" lastPageNumber="1791" masterDocId="32430065FFE0FFFBDD24FFD1FF84CC7C" masterDocTitle="Homoplasy in shells discombobulated the taxonomy: revision of the larger helicarionid land snails of northern Queensland, Australia (Stylommatophora: Helicarionidae)" masterLastPageNumber="1799" masterPageNumber="1727" pageNumber="1790" updateTime="1699412228230" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="393F6324703C0F48681424B6155B6B41">Homoplasy in shells discombobulated the taxonomy: revision of the larger helicarionid land snails of northern Queensland, Australia (Stylommatophora: Helicarionidae)</mods:title>
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<mods:namePart id="E2AE900AAD16144005EEA659F2CC8272">Hyman, Isabel T.</mods:namePart>
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<mods:title id="3A22AFEAD78D0C351AA79B9B275E94AE">Journal of Natural History</mods:title>
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<mods:date id="30B2935333EB342EEC2C6BD66F86E448">2022</mods:date>
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<mods:number id="DDB444416D8E432160EE64E56170C3CB">2022-11-23</mods:number>
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<mods:number id="FC28327E9608BECD06639B915911226F">41 - 44</mods:number>
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<heading id="1D247E67FFDFFFC4DF25FB24FCA6C973" box="[513,802,1269,1295]" centered="true" fontSize="11" level="2" pageId="63" pageNumber="1790" reason="5">
<taxonomicName id="81D3B288FFDFFFC4DF25FB24FCA6C973" ID-CoL="7P3BY" authority="Stanisic, 2010" authorityName="Stanisic" authorityYear="2010" box="[513,802,1269,1295]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="1790" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFDFFFC4DF25FB24FDFFC973" bold="true" box="[513,635,1269,1295]" italics="true" pageId="63" pageNumber="1790">Elatonitor</emphasis>
<bibRefCitation id="2242B4FAFFDFFFC4DFA6FB24FCA6C973" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[642,802,1269,1295]" pageId="63" pageNumber="1790" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic, 2010</bibRefCitation>
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<treatmentCitationGroup id="66C3EE25FFDFFFC4DD84FAFBFDE2C93F" box="[160,614,1322,1347]" pageId="63" pageNumber="1790">
<treatmentCitation id="C772EF1AFFDFFFC4DD84FAFBFE5EC93F" author="Iredale T" box="[160,474,1322,1347]" page="56" pageId="63" pageNumber="1790" year="1933">
<taxonomicName id="81D3B288FFDFFFC4DD84FAFBFE5EC93F" authority="Iredale, 1933: 56" authorityName="Iredale" authorityPageNumber="56" authorityYear="1933" box="[160,474,1322,1347]" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="1790" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFDFFFC4DD84FAFBFE9FC93F" box="[160,283,1322,1347]" italics="true" pageId="63" pageNumber="1790">Malandena</emphasis>
<bibRefCitation id="2242B4FAFFDFFFC4DC07FAFBFE5EC93F" author="Iredale T" box="[291,474,1322,1347]" pageId="63" pageNumber="1790" pagination="37 - 59" refId="ref30578" refString="Iredale T. 1933. Systematic notes on Australian land shells. Rec Aust Mus. 19 (1): 37 - 59. doi: 10.3853 / j. 0067 - 1975.19.1933.690." type="journal article" year="1933">Iredale, 1933: 56</bibRefCitation>
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[
<emphasis id="74A71519FFDFFFC4DCCEFAFBFDD7C93F" box="[490,595,1322,1347]" italics="true" pageId="63" pageNumber="1790">nom. nud</emphasis>
.].
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</paragraph>
<paragraph id="466CC90BFFDFFFC4DD84FA9DFDDCC919" blockId="63.[160,1155,1322,1520]" box="[160,600,1356,1381]" pageId="63" pageNumber="1790">
<treatmentCitationGroup id="66C3EE25FFDFFFC4DD84FA9DFDDCC919" box="[160,600,1356,1381]" pageId="63" pageNumber="1790">
<treatmentCitation id="C772EF1AFFDFFFC4DD84FA9DFE49C919" author="Iredale T" box="[160,461,1356,1381]" page="7" pageId="63" pageNumber="1790" year="1937">
<taxonomicName id="81D3B288FFDFFFC4DD84FA9DFE49C919" authority="Iredale, 1937: 7" authorityName="Iredale" authorityPageNumber="7" authorityYear="1937" box="[160,461,1356,1381]" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="1790" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFDFFFC4DD84FA9DFE9FC919" box="[160,283,1356,1381]" italics="true" pageId="63" pageNumber="1790">Malandena</emphasis>
<bibRefCitation id="2242B4FAFFDFFFC4DC07FA9DFE49C919" author="Iredale T" box="[291,461,1356,1381]" pageId="63" pageNumber="1790" pagination="1 - 39" refId="ref30613" refString="Iredale T. 1937. A basic list of the land Mollusca of Australia. Part II. Aust Zool. 9: 1 - 39." type="journal article" year="1937">Iredale, 1937: 7</bibRefCitation>
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[
<emphasis id="74A71519FFDFFFC4DCF8FA9DFDC1C919" box="[476,581,1356,1381]" italics="true" pageId="63" pageNumber="1790">nom. nud</emphasis>
.].
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</paragraph>
<paragraph id="466CC90BFFDFFFC4DD84FABEFD70C9F4" blockId="63.[160,1155,1322,1520]" box="[160,756,1391,1416]" pageId="63" pageNumber="1790">
<treatmentCitationGroup id="66C3EE25FFDFFFC4DD84FABEFD70C9F4" box="[160,756,1391,1416]" pageId="63" pageNumber="1790">
<treatmentCitation id="C772EF1AFFDFFFC4DD84FABEFDEDC9F4" author="Hyman IT &amp; Ponder WF" box="[160,617,1391,1416]" page="67" pageId="63" pageNumber="1790" year="2010">
<taxonomicName id="81D3B288FFDFFFC4DD84FABEFDEDC9F4" authority="Hyman and Ponder, 2010: 67" authorityName="Hyman and Ponder" authorityPageNumber="67" authorityYear="2010" box="[160,617,1391,1416]" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="1790" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFDFFFC4DD84FABEFE9FC9F4" box="[160,283,1391,1416]" italics="true" pageId="63" pageNumber="1790">Malandena</emphasis>
<bibRefCitation id="2242B4FAFFDFFFC4DC07FABEFDEDC9F4" author="Hyman IT &amp; Ponder WF" box="[291,617,1391,1416]" pageId="63" pageNumber="1790" pagination="1 - 148" refId="ref30470" refString="Hyman IT, Ponder WF. 2010. A morphological phylogenetic analysis and generic revision of Australian Helicarionidae (Gastropoda: Pulmonata: Stylommatophora), and an assessment of the relationships of the family. Zootaxa. 2462 (1): 1 - 148. doi: 10.11646 / zootaxa. 2462.1.1." type="journal article" year="2010">Hyman and Ponder, 2010: 67</bibRefCitation>
</taxonomicName>
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[
<emphasis id="74A71519FFDFFFC4DF5CFABEFD65C9F4" box="[632,737,1391,1416]" italics="true" pageId="63" pageNumber="1790">nom. nud</emphasis>
.].
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</paragraph>
<paragraph id="466CC90BFFDFFFC4DD84FA43FDF8C9D7" blockId="63.[160,1155,1322,1520]" box="[160,636,1426,1451]" pageId="63" pageNumber="1790">
<treatmentCitationGroup id="66C3EE25FFDFFFC4DD84FA43FDF8C9D7" box="[160,636,1426,1451]" pageId="63" pageNumber="1790">
<treatmentCitation id="C772EF1AFFDFFFC4DD84FA43FE75C9D7" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[160,497,1426,1451]" page="296" pageId="63" pageNumber="1790" year="2010">
<taxonomicName id="81D3B288FFDFFFC4DD84FA43FE75C9D7" authority="Stanisic, 2010: 296" authorityName="Stanisic" authorityPageNumber="296" authorityYear="2010" box="[160,497,1426,1451]" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="1790" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFDFFFC4DD84FA43FE9FC9D7" box="[160,283,1426,1451]" italics="true" pageId="63" pageNumber="1790">Malandena</emphasis>
<bibRefCitation id="2242B4FAFFDFFFC4DC07FA43FE75C9D7" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[291,497,1426,1451]" pageId="63" pageNumber="1790" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic, 2010: 296</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<emphasis id="74A71519FFDFFFC4DF24FA43FDEDC9D7" box="[512,617,1426,1451]" italics="true" pageId="63" pageNumber="1790">nom. nud</emphasis>
.].
</treatmentCitationGroup>
</paragraph>
<paragraph id="466CC90BFFDFFFC4DD84FA65FD7DC9B1" blockId="63.[160,1155,1322,1520]" box="[160,761,1460,1485]" pageId="63" pageNumber="1790">
<treatmentCitationGroup id="66C3EE25FFDFFFC4DD84FA65FD7DC9B1" box="[160,761,1460,1485]" pageId="63" pageNumber="1790">
<treatmentCitation id="C772EF1AFFDFFFC4DD84FA65FE6EC9B1" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[160,490,1460,1485]" page="298" pageId="63" pageNumber="1790" year="2010">
<taxonomicName id="81D3B288FFDFFFC4DD84FA65FE6EC9B1" ID-CoL="7P3BY" authority="Stanisic, 2010: 298" authorityName="Stanisic" authorityPageNumber="298" authorityYear="2010" box="[160,490,1460,1485]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="1790" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFDFFFC4DD84FA65FE8EC9B1" box="[160,266,1460,1485]" italics="true" pageId="63" pageNumber="1790">Elatonitor</emphasis>
<bibRefCitation id="2242B4FAFFDFFFC4DC31FA65FE6EC9B1" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[277,490,1460,1485]" pageId="63" pageNumber="1790" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic, 2010: 298</bibRefCitation>
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(of masculine gender).
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</paragraph>
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<subSubSection id="0EC99A80FFDFFFC4DE27FA65FEBAC98C" pageId="63" pageNumber="1790" type="type_taxon">
<paragraph id="466CC90BFFDFFFC4DE27FA65FEBAC98C" blockId="63.[160,1155,1322,1520]" pageId="63" pageNumber="1790">
<typeStatus id="996877A9FFDFFFC4DE27FA65FCBDC9B1" box="[771,825,1460,1485]" pageId="63" pageNumber="1790">Type</typeStatus>
species
<taxonomicName id="81D3B288FFDFFFC4DE84FA65FEBDC98C" ID-CoL="395GM" authority="Stanisic, 2010" authorityName="Stanisic" authorityYear="2010" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="63" pageNumber="1790" phylum="Mollusca" rank="species" species="montanus">
<emphasis id="74A71519FFDFFFC4DE84FA65FB07C9B1" box="[928,1155,1460,1485]" italics="true" pageId="63" pageNumber="1790">Elatonitor montanus</emphasis>
<bibRefCitation id="2242B4FAFFDFFFC4DD84FA06FEBDC98C" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[160,313,1495,1520]" pageId="63" pageNumber="1790" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic, 2010</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0EC99A80FFDFFFBBDD84F9CDFE5BCF5B" lastPageId="64" lastPageNumber="1791" pageId="63" pageNumber="1790" type="diagnosis">
<paragraph id="466CC90BFFDFFFC4DD84F9CDFE19CA4A" blockId="63.[160,1155,1564,1728]" box="[160,413,1564,1590]" pageId="63" pageNumber="1790">
<emphasis id="74A71519FFDFFFC4DD84F9CDFE19CA4A" bold="true" box="[160,413,1564,1590]" italics="true" pageId="63" pageNumber="1790">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="466CC90BFFDFFFBBDD84F9EFFF5BCC91" blockId="63.[160,1155,1564,1728]" lastBlockId="64.[160,1155,142,237]" lastPageId="64" lastPageNumber="1791" pageId="63" pageNumber="1790">
<emphasis id="74A71519FFDFFFC4DD84F9EFFE1BCA24" bold="true" box="[160,415,1598,1624]" italics="true" pageId="63" pageNumber="1790">External appearance.</emphasis>
Shell medium-sized to large (SW
<quantity id="812B64EEFFDFFFC4DE12F9EEFC4CCA24" box="[822,968,1599,1624]" metricMagnitude="-2" metricUnit="m" metricValue="1.6949999999999998" metricValueMax="2.0" metricValueMin="1.39" pageId="63" pageNumber="1790" unit="mm" value="16.95" valueMax="20.0" valueMin="13.9">13.9–20 mm</quantity>
), pale amber to amber, often deep amber below periphery, glossy, 5.3–5.8 whorls, depressedly globose to trochoidal with a moderate spire, whorl profile mildly to strongly angulate. Protoconch with very fine spiral grooves, sometimes becoming obsolete; teleoconch with very fine and dense to coarse, distinct spiral grooves. Body light to dark brown. Mantle lobes small to moderately small; shell lappets long, narrow, tapering. Caudal horn moderately large to large.
</paragraph>
<paragraph id="466CC90BFFA0FFBBDD84FEF9FE5BCF5B" blockId="64.[160,1156,296,807]" pageId="64" pageNumber="1791">
<emphasis id="74A71519FFA0FFBBDD84FEF9FEFCCD3E" bold="true" box="[160,376,296,322]" italics="true" pageId="64" pageNumber="1791">Genital anatomy.</emphasis>
Ovotestis of 8–9 lobes, embedded in digestive gland. Carrefour visible; talon usually embedded in albumen gland. Spermoviduct not folded. Free oviduct long, with elongate capsular gland; free oviduct internally sculptured with longitudinal pilasters; capsular gland with no internal sculpture. Vagina short to long, sculptured internally with longitudinal pilasters or a network of wavy ridges. Bursa copulatrix inserted on the vagina, moderately short to moderately long, duct narrow, sometimes slightly broader at base, internally with primarily longitudinal sculpture; bursa elongate oval. Penis moderately small to long, tubular; internally with 2–4 longitudinal pilasters which sometimes merge or divide, internal wall sculptured with longitudinal wrinkles or ridges; penis and sometimes part of epiphallus enclosed in penial tunica. Epiphallus enters penis through a simple pore or a short verge; epiphallus longer than penis; epiphallus 2 equal to or longer than epiphallus 1; epiphallic caecum absent; flagellum long, slender, without obvious internal cryptae. Spermatophore a soft-walled capsule with hard tail-pipe; capsule very elongate, with elongate tip; tail-pipe short, sculptured with short hooks, becoming more dense towards tip.
</paragraph>
</subSubSection>
<subSubSection id="0EC99A80FFA0FFBBDD84FCB2FC8EC9DB" pageId="64" pageNumber="1791" type="discussion">
<paragraph id="466CC90BFFA0FFBBDD84FCB2FE82CF01" blockId="64.[160,1156,867,1447]" box="[160,262,867,893]" pageId="64" pageNumber="1791">
<emphasis id="74A71519FFA0FFBBDD84FCB2FE82CF01" bold="true" box="[160,262,867,893]" italics="true" pageId="64" pageNumber="1791">Remarks</emphasis>
</paragraph>
<paragraph id="466CC90BFFA0FFBBDD84FC57FCA7C813" blockId="64.[160,1156,867,1447]" pageId="64" pageNumber="1791">
<taxonomicName id="81D3B288FFA0FFBBDD84FC57FE8ECFE3" authorityName="Stanisic" authorityYear="2010" box="[160,266,902,927]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFA0FFBBDD84FC57FE8ECFE3" box="[160,266,902,927]" italics="true" pageId="64" pageNumber="1791">Elatonitor</emphasis>
</taxonomicName>
was introduced for a single species,
<taxonomicName id="81D3B288FFA0FFBBDF84FC57FCA2CFE3" authorityName="Stanisic" authorityYear="2010" box="[672,806,902,927]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="species" species="montanus">
<emphasis id="74A71519FFA0FFBBDF84FC57FCA2CFE3" box="[672,806,902,927]" italics="true" pageId="64" pageNumber="1791">E. montanus</emphasis>
</taxonomicName>
, based on the sharply angulate whorls and very fine, very dense spiral sculpture of its shell. In the current study we demonstrate that there is a close relationship between
<taxonomicName id="81D3B288FFA0FFBBDE1CFC1DFC41CF99" authorityName="Stanisic" authorityYear="2010" box="[824,965,972,997]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="species" species="montanus">
<emphasis id="74A71519FFA0FFBBDE1CFC1DFC41CF99" box="[824,965,972,997]" italics="true" pageId="64" pageNumber="1791">E. montanus</emphasis>
</taxonomicName>
and
<taxonomicName id="81D3B288FFA0FFBBD923FC1DFF7FC87B" authorityName="Hyman &amp; Köhler" authorityYear="2022" baseAuthorityName="Hyman &amp; Köhler" baseAuthorityYear="2022" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="species" species="suturalis">
<emphasis id="74A71519FFA0FFBBD923FC1DFF7FC87B" italics="true" pageId="64" pageNumber="1791">Malandena suturalis</emphasis>
</taxonomicName>
, sole remaining member of genus
<taxonomicName id="81D3B288FFA0FFBBDFBFFC3FFC92C87B" authorityName="Stanisic" authorityYear="2010" box="[667,790,1006,1031]" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFA0FFBBDFBFFC3FFC92C87B" box="[667,790,1006,1031]" italics="true" pageId="64" pageNumber="1791">Malandena</emphasis>
</taxonomicName>
. However,
<taxonomicName id="81D3B288FFA0FFBBDEBFFC3FFB92C87B" authorityName="Stanisic" authorityYear="2010" box="[923,1046,1006,1031]" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFA0FFBBDEBFFC3FFB92C87B" box="[923,1046,1006,1031]" italics="true" pageId="64" pageNumber="1791">Malandena</emphasis>
</taxonomicName>
, while in current usage, has never been validly described and therefore remains unavailable (
<bibRefCitation id="2242B4FAFFA0FFBBD91DFBC0FEF1C831" author="Kohler F &amp; Bouchet P" pageId="64" pageNumber="1791" pagination="150 - 159" refId="ref30807" refString="Kohler F, Bouchet P. 2020. On unavailable genus-group names introduced by Tom Iredale for Australian non-marine gastropods: nomenclatural clarifications and descriptions of new genera. Molluscan Res. 40 (2): 150 - 159. doi: 10.1080 / 13235818.2020.1724603." type="journal article" year="2020">Köhler and Bouchet 2020</bibRefCitation>
). Consequently,
<taxonomicName id="81D3B288FFA0FFBBDF1CFBE5FD26C831" authorityName="Stanisic" authorityYear="2010" box="[568,674,1076,1101]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFA0FFBBDF1CFBE5FD26C831" box="[568,674,1076,1101]" italics="true" pageId="64" pageNumber="1791">Elatonitor</emphasis>
</taxonomicName>
takes its place. Like all names ending in ‘nitor’, the genus name
<taxonomicName id="81D3B288FFA0FFBBDC88FB87FD92C813" authorityName="Stanisic" authorityYear="2010" box="[428,534,1110,1135]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFA0FFBBDC88FB87FD92C813" box="[428,534,1110,1135]" italics="true" pageId="64" pageNumber="1791">Elatonitor</emphasis>
</taxonomicName>
is of masculine gender.
</paragraph>
<paragraph id="466CC90BFFA0FFBBDDE4FBA8FC8EC9DB" blockId="64.[160,1156,867,1447]" pageId="64" pageNumber="1791">
<taxonomicName id="81D3B288FFA0FFBBDDE4FBA8FEAEC8EE" authorityName="Stanisic" authorityYear="2010" box="[192,298,1145,1170]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFA0FFBBDDE4FBA8FEAEC8EE" box="[192,298,1145,1170]" italics="true" pageId="64" pageNumber="1791">Elatonitor</emphasis>
</taxonomicName>
is characterised by its medium-sized to large, glossy shell (shell
<quantity id="812B64EEFFA0FFBBDED5FBA8FBE3C8EE" box="[1009,1127,1145,1170]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="64" pageNumber="1791" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
in width), depressedly globose to trochoidal shape with an angulate periphery and usually a supraperipheral band, and by anatomical features including the presence of at least a short vagina, a long, slender penis with longitudinal pilasters and longitudinal ridges or wrinkles, and an epiphallus with no caecum and with a slender flagellum. In particular, it can be distinguished from the similar genera
<taxonomicName id="81D3B288FFA0FFBBDF8FFAF7FC98C943" authorityName="Iredale" authorityYear="1937" box="[683,796,1318,1343]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFA0FFBBDF8FFAF7FC98C943" box="[683,796,1318,1343]" italics="true" pageId="64" pageNumber="1791">Pravonitor</emphasis>
</taxonomicName>
and
<taxonomicName id="81D3B288FFA0FFBBDE7CFAF7FC47C943" authorityName="Hyman &amp; Köhler" authorityYear="2022" box="[856,963,1318,1343]" class="Gastropoda" family="Helicarionidae" genus="Geminitor" kingdom="Animalia" order="Stylommatophora" pageId="64" pageNumber="1791" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFA0FFBBDE7CFAF7FC47C943" box="[856,963,1318,1343]" italics="true" pageId="64" pageNumber="1791">Geminitor</emphasis>
</taxonomicName>
by its large size, angulate shell periphery, the lack of an epiphallic caecum and the single-chambered penis with primarily longitudinal sculpture. This genus is found in rainforest and vine thicket from Mt Bartle Frere to Cooktown (
<bibRefCitation id="2242B4FAFFA0FFBBDF0EFA5FFD79C9DB" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[554,765,1422,1447]" pageId="64" pageNumber="1791" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic et al. 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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