238 lines
30 KiB
XML
238 lines
30 KiB
XML
<document id="F12D8F36319EEF545CB69D6BEF09AD09" ID-CLB-Dataset="198290" ID-DOI="10.1080/00222933.2022.2136017" ID-GBIF-Dataset="c9288626-198f-4b45-b923-f134cc0df48a" ID-ISSN="1464-5262" ID-Zenodo-Dep="7380681" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669805002707" checkinUser="plazi" docAuthor="Hyman, Isabel T. & Köhler, Frank" docDate="2022" docId="CE7A781DFFD9FFC0DC91FB42FF7CCF94" docLanguage="en" docName="JNATHIST.56.41-44.1727-1799.pdf" docOrigin="Journal of Natural History 56 (41 - 44)" docSource="http://dx.doi.org/10.1080/00222933.2022.2136017" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Pravonitor monteithi" docType="treatment" docVersion="9" lastPageNumber="1786" masterDocId="32430065FFE0FFFBDD24FFD1FF84CC7C" masterDocTitle="Homoplasy in shells discombobulated the taxonomy: revision of the larger helicarionid land snails of northern Queensland, Australia (Stylommatophora: Helicarionidae)" masterLastPageNumber="1799" masterPageNumber="1727" pageNumber="1784" updateTime="1699412228230" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9DAEC165100273A43652DACC368A37AE">Homoplasy in shells discombobulated the taxonomy: revision of the larger helicarionid land snails of northern Queensland, Australia (Stylommatophora: Helicarionidae)</mods:title>
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<mods:namePart id="ABC62836F17DC5DCCED928D977383D1C">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="926E504DD8A3F8E0856828F2C8C8B79E">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="ECF6A63C04B8E73E668DDD266CB8B900">text</mods:typeOfResource>
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<mods:title id="A90D63C330A5D08A87CC92A8AE5F94C8">Journal of Natural History</mods:title>
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<mods:part id="F33D1454771AC66752DCC15EBF2B4C1F">
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<mods:date id="A308C0CFB4BD8F132B94F59708AD58B3">2022</mods:date>
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<mods:number id="B08C28AF52B68D82272FF0F85A263F4F">2022-11-23</mods:number>
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<mods:number id="7DB05D936BCC2DB39750FF2FCB1562D2">56</mods:number>
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<mods:number id="F71B86C2C499BD6B35FAA78402FD07D1">41 - 44</mods:number>
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<mods:identifier id="18E3C63D24F4BE2B4D6CB8F0335601A9" type="CLB-Dataset">198290</mods:identifier>
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<treatment id="CE7A781DFFD9FFC0DC91FB42FF7CCF94" ID-DOI="http://doi.org/10.5281/zenodo.7383448" ID-GBIF-Taxon="205276705" ID-Zenodo-Dep="7383448" LSID="urn:lsid:plazi:treatment:CE7A781DFFD9FFC0DC91FB42FF7CCF94" httpUri="http://treatment.plazi.org/id/CE7A781DFFD9FFC0DC91FB42FF7CCF94" lastPageId="59" lastPageNumber="1786" pageId="57" pageNumber="1784">
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<subSubSection id="0EC99A80FFD9FFC2DC91FB42FCEAC8D1" box="[437,878,1171,1197]" pageId="57" pageNumber="1784" type="nomenclature">
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<paragraph id="466CC90BFFD9FFC2DC91FB42FCEAC8D1" blockId="57.[437,878,1171,1197]" box="[437,878,1171,1197]" pageId="57" pageNumber="1784">
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<heading id="1D247E67FFD9FFC2DC91FB42FCEAC8D1" box="[437,878,1171,1197]" centered="true" fontSize="11" level="2" pageId="57" pageNumber="1784" reason="5">
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<taxonomicName id="81D3B288FFD9FFC2DC91FB42FCEAC8D1" ID-CoL="32cae32f-7ff7-41c7-a7ec-24c8e79fcfd6" authority="(Stanisic, 2010)" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[437,878,1171,1197]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="monteithi">
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<emphasis id="74A71519FFD9FFC2DC91FB42FD33C8D1" bold="true" box="[437,695,1171,1197]" italics="true" pageId="57" pageNumber="1784">Pravonitor monteithi</emphasis>
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(
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<bibRefCitation id="2242B4FAFFD9FFC2DFE0FB42FCE2C8D1" author="Stanisic J & Shea M & Potter D & Griffiths O" box="[708,870,1171,1197]" pageId="57" pageNumber="1784" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic, 2010</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0EC99A80FFD9FFC2DF3EFB11FC8EC8A6" box="[538,778,1216,1242]" pageId="57" pageNumber="1784" type="description">
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<paragraph id="466CC90BFFD9FFC2DF3EFB11FC8EC8A6" blockId="57.[538,778,1216,1242]" box="[538,778,1216,1242]" pageId="57" pageNumber="1784">
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(
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<figureCitation id="DEE8D58EFFD9FFC2DF06FB11FD14C8A6" box="[546,656,1216,1242]" captionStart="Figure 4" captionStartId="27.[160,225,1613,1636]" captionTargetBox="[255,1060,142,1581]" captionTargetId="figure-3@27.[254,1061,142,1582]" captionTargetPageId="27" captionText="Figure 4. Distribution map. Grey symbols: Geminitor. Blue symbols: Pravonitor. Red symbols: Elatonitor. Biogeographic barriers are marked in black; A = Normanby Basin, B = Black Mountain Corridor, and C = Atherton Tableland." figureDoi="http://doi.org/10.5281/zenodo.7380691" httpUri="https://zenodo.org/record/7380691/files/figure.png" pageId="57" pageNumber="1784">Figures 4</figureCitation>
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,
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<figureCitation id="DEE8D58EFFD9FFC2DFBFFB11FD38C8A6" box="[667,700,1216,1242]" captionStart="Figure 14" captionStartId="47.[160,225,1043,1066]" captionTargetBox="[164,1151,143,1010]" captionTargetId="figure-168@47.[163,1152,142,1011]" captionTargetPageId="47" captionText="Figure 14. Shells of Pravonitor. a) Pravonitor insularum, AM C.170795 (paratype). b) Pravonitor annulus AM C.425415,Gabba Is. c) Pravonitor aquilonia, QM MO48285 (holotype). d) Pravonitor ferrugineus, QM MO12752 (holotype). e). Pravonitor kreffti, AM C.1352, Thursday Is. f) Pravonitor kreffti, AM C.425474, Simpson’s Bay, Utingu. g) Pravonitor monteithi, QM MO48587 (holotype). h) Pravonitor septentrionalis, AM C.425422. i) Pravonitor stuarti, QM MO80213 (paratype). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380720" httpUri="https://zenodo.org/record/7380720/files/figure.png" pageId="57" pageNumber="1784">14</figureCitation>
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(g), 19)
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</paragraph>
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</subSubSection>
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<subSubSection id="0EC99A80FFD9FFC2DD84FB24FD1BC972" box="[160,671,1269,1294]" pageId="57" pageNumber="1784" type="reference_group">
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<paragraph id="466CC90BFFD9FFC2DD84FB24FD1BC972" blockId="57.[160,671,1269,1294]" box="[160,671,1269,1294]" pageId="57" pageNumber="1784">
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<treatmentCitationGroup id="66C3EE25FFD9FFC2DD84FB24FD1BC972" box="[160,671,1269,1294]" pageId="57" pageNumber="1784">
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<treatmentCitation id="C772EF1AFFD9FFC2DD84FB24FD1EC972" author="Stanisic J & Shea M & Potter D & Griffiths O" box="[160,666,1269,1294]" page="297" pageId="57" pageNumber="1784" year="2010">
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<taxonomicName id="81D3B288FFD9FFC2DD84FB24FD1EC972" ID-CoL="7S25L" authority="Stanisic, 2010: 296 - 297" authorityName="Stanisic" authorityPageNumber="296 - 297" authorityYear="2010" box="[160,666,1269,1294]" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="monteithi">
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<emphasis id="74A71519FFD9FFC2DD84FB24FE08C972" box="[160,396,1269,1294]" italics="true" pageId="57" pageNumber="1784">Malandena monteithi</emphasis>
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<bibRefCitation id="2242B4FAFFD9FFC2DCB7FB24FD1EC972" author="Stanisic J & Shea M & Potter D & Griffiths O" box="[403,666,1269,1294]" pageId="57" pageNumber="1784" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic, 2010: 296–297</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="0EC99A80FFD9FFC2DD84FAEFFE69CA4D" pageId="57" pageNumber="1784" type="materials_examined">
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||
<paragraph id="466CC90BFFD9FFC2DD84FAEFFBAAC907" blockId="57.[160,1155,1342,1403]" pageId="57" pageNumber="1784">
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<materialsCitation id="F6BBC356FFD9FFC2DD84FAEFFBAAC907" ID-GBIF-Occurrence="3978770308" collectingDate="1993-11-30" collectorName="G. Monteith & H. Janetzki & D. Cook." country="Australia" elevation="1050" location="Isley Hills" municipality="Qld" pageId="57" pageNumber="1784" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="74A71519FFD9FFC2DD84FAEFFE96C924" bold="true" box="[160,274,1342,1368]" italics="true" pageId="57" pageNumber="1784">
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||
<typeStatus id="996877A9FFD9FFC2DD84FAEFFE89C924" box="[160,269,1342,1368]" pageId="57" pageNumber="1784" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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QMMO48587,
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<location id="430C9FD0FFD9FFC2DCE7FAEEFDA8C924" LSID="urn:lsid:plazi:treatment:CE7A781DFFD9FFC0DC91FB42FF7CCF94:430C9FD0FFD9FFC2DCE7FAEEFDA8C924" box="[451,556,1343,1368]" country="Australia" municipality="Qld" name="Isley Hills" pageId="57" pageNumber="1784">Isley Hills</location>
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, north-eastern
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<collectingMunicipality id="A6085371FFD9FFC2DFF3FAEEFC85C924" box="[727,769,1343,1368]" pageId="57" pageNumber="1784">Qld</collectingMunicipality>
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||
,
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<collectingCountry id="3EC4899BFFD9FFC2DE28FAEEFCE9C924" box="[780,877,1343,1368]" name="Australia" pageId="57" pageNumber="1784">Australia</collectingCountry>
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(17°03ʹS, 145°42ʹE), rainforest,
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<quantity id="812B64EEFFD9FFC2DDC9FAB3FEC7C907" box="[237,323,1378,1403]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="57" pageNumber="1784" unit="m" value="1050.0">
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<elevation id="CDFE2E38FFD9FFC2DDC9FAB3FEC7C907" box="[237,323,1378,1403]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="57" pageNumber="1784" unit="m" value="1050.0">1050 m</elevation>
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</quantity>
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, under logs,
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<date id="326DEFCBFFD9FFC2DCF0FAB3FD29C907" box="[468,685,1378,1403]" pageId="57" pageNumber="1784" value="1993-11-30">
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||
<collectingDate id="22291623FFD9FFC2DCF0FAB3FD29C907" box="[468,685,1378,1403]" pageId="57" pageNumber="1784" value="1993-11-30">30 November 1993</collectingDate>
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</date>
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,
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<collectorName id="EB26ACDDFFD9FFC2DF9CFAB3FCBBC907" box="[696,831,1378,1403]" pageId="57" pageNumber="1784">G. Monteith</collectorName>
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,
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<collectorName id="EB26ACDDFFD9FFC2DE6EFAB3FC41C907" box="[842,965,1378,1403]" pageId="57" pageNumber="1784">H. Janetzki</collectorName>
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,
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<collectorName id="EB26ACDDFFD9FFC2DEEBFAB3FBAAC907" box="[975,1070,1378,1403]" pageId="57" pageNumber="1784">D. Cook.</collectorName>
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</materialsCitation>
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||
</paragraph>
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<paragraph id="466CC90BFFD9FFC2DD84FA7AFE8DC99B" blockId="57.[160,1155,1451,1511]" pageId="57" pageNumber="1784">
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<materialsCitation id="F6BBC356FFD9FFC2DD84FA7AFF34C99B" ID-GBIF-Occurrence="3978770316" collectionCode="AM" pageId="57" pageNumber="1784" specimenCount="1" typeStatus="paratype">
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<emphasis id="74A71519FFD9FFC2DD84FA7AFE9BC9B9" bold="true" box="[160,287,1451,1477]" italics="true" pageId="57" pageNumber="1784">
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<typeStatus id="996877A9FFD9FFC2DD84FA7AFE9EC9B9" box="[160,282,1451,1477]" pageId="57" pageNumber="1784" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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QMMO24798, QMMO48164, QMMO48203, QMMO48693, QMMO78927,
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<collectionCode id="20C251CEFFD9FFC2D979FA7DFB07C9B9" box="[1117,1155,1452,1477]" country="Australia" name="Australian Museum" pageId="57" pageNumber="1784" type="Museum">AM</collectionCode>
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C
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</materialsCitation>
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.425580.
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</paragraph>
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<paragraph id="466CC90BFFD9FFC2DD84F9C6FE69CA4D" blockId="57.[160,493,1559,1585]" box="[160,493,1559,1585]" pageId="57" pageNumber="1784">
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<emphasis id="74A71519FFD9FFC2DD84F9C6FED1CA4D" bold="true" box="[160,341,1559,1585]" italics="true" pageId="57" pageNumber="1784">Other material.</emphasis>
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See
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<tableCitation id="0B51FCB0FFD9FFC2DCB1F9C9FE63CA4D" box="[405,487,1560,1585]" captionStart="Table 1" captionStartId="5.[133,189,450,473]" captionTargetPageId="5" captionText="Table 1. Material studied. Abbreviations: H, holotype; L, lectotype; N, neotype; P, paratype; PL, paralectotype, PN, paraneotype; S, syntype." httpUri="http://table.plazi.org/id/12AC9983FFE5FFFEDDA1FE13FA67CDA5" pageId="57" pageNumber="1784" tableUuid="12AC9983FFE5FFFEDDA1FE13FA67CDA5">Table 1</tableCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0EC99A80FFD9FFC1DD84F9B0FEB3CEE0" lastPageId="58" lastPageNumber="1785" pageId="57" pageNumber="1784" type="diagnosis">
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||
<paragraph id="466CC90BFFD9FFC2DD84F9B0FE90CA07" blockId="57.[159,1156,1633,1728]" box="[160,276,1633,1659]" pageId="57" pageNumber="1784">
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||
<emphasis id="74A71519FFD9FFC2DD84F9B0FE90CA07" bold="true" box="[160,276,1633,1659]" italics="true" pageId="57" pageNumber="1784">Diagnosis</emphasis>
|
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</paragraph>
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<paragraph id="466CC90BFFD9FFC1DD84F955FBF9CC91" blockId="57.[159,1156,1633,1728]" lastBlockId="58.[159,1155,142,237]" lastPageId="58" lastPageNumber="1785" pageId="57" pageNumber="1784">
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||
<emphasis id="74A71519FFD9FFC2DD84F955FE1BCAE2" bold="true" box="[160,415,1668,1694]" italics="true" pageId="57" pageNumber="1784">External morphology.</emphasis>
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Shell (
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<figureCitation id="DEE8D58EFFD9FFC2DCCBF954FDD1CAE2" box="[495,597,1669,1694]" captionStart="Figure 14" captionStartId="47.[160,225,1043,1066]" captionTargetBox="[164,1151,143,1010]" captionTargetId="figure-168@47.[163,1152,142,1011]" captionTargetPageId="47" captionText="Figure 14. Shells of Pravonitor. a) Pravonitor insularum, AM C.170795 (paratype). b) Pravonitor annulus AM C.425415,Gabba Is. c) Pravonitor aquilonia, QM MO48285 (holotype). d) Pravonitor ferrugineus, QM MO12752 (holotype). e). Pravonitor kreffti, AM C.1352, Thursday Is. f) Pravonitor kreffti, AM C.425474, Simpson’s Bay, Utingu. g) Pravonitor monteithi, QM MO48587 (holotype). h) Pravonitor septentrionalis, AM C.425422. i) Pravonitor stuarti, QM MO80213 (paratype). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380720" httpUri="https://zenodo.org/record/7380720/files/figure.png" pageId="57" pageNumber="1784">Figure 14</figureCitation>
|
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(g)) medium-sized (SW
|
||
<quantity id="812B64EEFFD9FFC2DE6FF954FC61CAE2" box="[843,997,1669,1694]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" metricValueMax="1.35" metricValueMin="1.19" pageId="57" pageNumber="1784" unit="mm" value="12.7" valueMax="13.5" valueMin="11.9">11.9–13.5 mm</quantity>
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), amber with a darker underside, 5.1–5.4 whorls, subdiscoidal with a low spire, whorl profile rounded; microsculpture of very fine to obsolete spirals on protoconch and teleoconch; umbilicus very narrow, partially covered by reflected lip. Body white with black eyestalks (in alcohol). Mantle lobes moderately small; shell lappets moderately large, narrow, tapering. Caudal horn large.
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</paragraph>
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<paragraph id="466CC90BFFDAFFC1DD84FEF9FEB3CEE0" blockId="58.[160,1156,296,668]" pageId="58" pageNumber="1785">
|
||
<emphasis id="74A71519FFDAFFC1DD84FEF9FEEBCD3E" bold="true" box="[160,367,296,322]" italics="true" pageId="58" pageNumber="1785">Genital anatomy.</emphasis>
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Genitalia (
|
||
<figureCitation id="DEE8D58EFFDAFFC1DCD1FEF8FDE6CD3E" box="[501,610,297,322]" captionStart="Figure 19" captionStartId="58.[160,225,1669,1692]" captionTargetBox="[162,1152,754,1637]" captionTargetId="figure-162@58.[189,1154,755,1638]" captionTargetPageId="58" captionText="Figure 19. Genitalia of Pravonitor monteithi. a) QM MO48669, Isley Hills. Reproductive system. b-c) QM MO48203, Lamb’s Head. b) Spermatophore. c) Penis interior. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7380734" httpUri="https://zenodo.org/record/7380734/files/figure.png" pageId="58" pageNumber="1785">Figure 19</figureCitation>
|
||
) with short vagina, internally with longitudinal pilasters; bursa copulatrix moderately short, duct broad at base, then narrowing, internally with distinct longitudinal pilasters; bursa elongate. Penis moderately small, bulging at base, tubular medially, swollen proximally with large blind tip; internally with 1–2 pustulose longitudinal pilasters, internal wall sculptured with fine pustules; basal bulge with thickened, folded region internally; basal bulge not enclosed in penial tunica. Epiphallus enters penis through a simple pore; epiphallus longer than penis; epiphallus 2 longer than epiphallus 1; moderately long epiphallic caecum present, with medial attachment of the penial retractor muscle; flagellum long, slender, without obvious internal cryptae. Spermatophore with soft capsule and smooth, elongate tail-pipe with no sculpture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="12AC9983FFDAFFC1DD84F954FC29CAC6" ID-DOI="http://doi.org/10.5281/zenodo.7380734" ID-Zenodo-Dep="7380734" httpUri="https://zenodo.org/record/7380734/files/figure.png" pageId="58" pageNumber="1785" startId="58.[160,225,1669,1692]" targetBox="[162,1152,754,1637]" targetPageId="58">
|
||
<paragraph id="466CC90BFFDAFFC1DD84F954FC29CAC6" blockId="58.[160,1155,1669,1722]" pageId="58" pageNumber="1785">
|
||
<emphasis id="74A71519FFDAFFC1DD84F954FE82CAE0" bold="true" box="[160,262,1669,1692]" pageId="58" pageNumber="1785">Figure 19.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="81D3B288FFDAFFC1DCA5F954FDC2CAE0" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[385,582,1669,1692]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="1785" phylum="Mollusca" rank="species" species="monteithi">
|
||
<emphasis id="74A71519FFDAFFC1DCA5F954FDC2CAE0" box="[385,582,1669,1692]" italics="true" pageId="58" pageNumber="1785">Pravonitor monteithi</emphasis>
|
||
</taxonomicName>
|
||
. a) QM MO48669, Isley Hills. Reproductive system. b-c) QM MO48203, Lamb’s Head. b) Spermatophore. c) Penis interior. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0EC99A80FFDBFFC0DD84FF5FFF7CCF94" pageId="59" pageNumber="1786" type="discussion">
|
||
<paragraph id="466CC90BFFDBFFC0DD84FF5FFE82CCD4" blockId="59.[159,1156,142,1000]" box="[160,262,142,168]" pageId="59" pageNumber="1786">
|
||
<emphasis id="74A71519FFDBFFC0DD84FF5FFE82CCD4" bold="true" box="[160,262,142,168]" italics="true" pageId="59" pageNumber="1786">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="466CC90BFFDBFFC0DD84FF63FEFECEAF" blockId="59.[159,1156,142,1000]" pageId="59" pageNumber="1786">
|
||
<taxonomicName id="81D3B288FFDBFFC0DD84FF63FEF1CCB7" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[160,373,178,203]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="monteithi">
|
||
<emphasis id="74A71519FFDBFFC0DD84FF63FEF1CCB7" box="[160,373,178,203]" italics="true" pageId="59" pageNumber="1786">Pravonitor monteithi</emphasis>
|
||
</taxonomicName>
|
||
was previously included in
|
||
<taxonomicName id="81D3B288FFDBFFC0DFB2FF63FC89CCB7" box="[662,781,178,203]" class="Gastropoda" family="Helicarionidae" genus="Malandena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="genus">
|
||
<emphasis id="74A71519FFDBFFC0DFB2FF63FC89CCB7" box="[662,781,178,203]" italics="true" pageId="59" pageNumber="1786">Malandena</emphasis>
|
||
</taxonomicName>
|
||
(now
|
||
<taxonomicName id="81D3B288FFDBFFC0DE69FF63FC36CCB7" authorityName="Stanisic" authorityYear="2010" box="[845,946,178,203]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="genus">
|
||
<emphasis id="74A71519FFDBFFC0DE69FF63FC36CCB7" box="[845,946,178,203]" italics="true" pageId="59" pageNumber="1786">Elatonitor</emphasis>
|
||
</taxonomicName>
|
||
) and bears a strong external similarity to
|
||
<taxonomicName id="81D3B288FFDBFFC0DC5BFF05FDC5CC91" box="[383,577,212,237]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="suturalis">
|
||
<emphasis id="74A71519FFDBFFC0DC5BFF05FDC5CC91" box="[383,577,212,237]" italics="true" pageId="59" pageNumber="1786">Elatonitor suturalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="81D3B288FFDBFFC0DF52FF05FD75CC91" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[630,753,212,237]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="aquilonia">
|
||
<emphasis id="74A71519FFDBFFC0DF52FF05FD75CC91" box="[630,753,212,237]" italics="true" pageId="59" pageNumber="1786">P. aquilonia</emphasis>
|
||
</taxonomicName>
|
||
. It can be distinguished by its smaller size, finer teleoconch sculpture and rounded whorl periphery. The presence of an epiphallic caecum with a medial attachment of the penial retractor muscle and an internally pustulose penis interior definitively places this species in
|
||
<taxonomicName id="81D3B288FFDBFFC0DF86FEEDFC8ACD29" authorityName="Iredale" authorityYear="1937" box="[674,782,316,341]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="genus">
|
||
<emphasis id="74A71519FFDBFFC0DF86FEEDFC8ACD29" box="[674,782,316,341]" italics="true" pageId="59" pageNumber="1786">Pravonitor</emphasis>
|
||
</taxonomicName>
|
||
. Only a single specimen could be sequenced,and this sequence diverged from within the clade
|
||
<taxonomicName id="81D3B288FFDBFFC0DE11FE8EFC2BCD04" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[821,943,351,376]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="aquilonia">
|
||
<emphasis id="74A71519FFDBFFC0DE11FE8EFC2BCD04" box="[821,943,351,376]" italics="true" pageId="59" pageNumber="1786">P. aquilonia</emphasis>
|
||
</taxonomicName>
|
||
. It is possible that
|
||
<taxonomicName id="81D3B288FFDBFFC0D954FE8EFE80CDE7" baseAuthorityName="Stanisic" baseAuthorityYear="2010" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="monteithi">
|
||
<emphasis id="74A71519FFDBFFC0D954FE8EFE80CDE7" italics="true" pageId="59" pageNumber="1786">P. monteithi</emphasis>
|
||
</taxonomicName>
|
||
should be recognised as a synonym of
|
||
<taxonomicName id="81D3B288FFDBFFC0DFEFFE53FCCECDE7" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[715,842,386,411]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="aquilonia">
|
||
<emphasis id="74A71519FFDBFFC0DFEFFE53FCCECDE7" box="[715,842,386,411]" italics="true" pageId="59" pageNumber="1786">P. aquilonia</emphasis>
|
||
</taxonomicName>
|
||
; however, there are distinct morphological differences between the two and we believe there is insufficient information to synonymise the two species. Specimens identified in the current study as
|
||
<taxonomicName id="81D3B288FFDBFFC0DE89FE16FB07CD9C" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[941,1155,455,480]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="monteithi">
|
||
<emphasis id="74A71519FFDBFFC0DE89FE16FB07CD9C" box="[941,1155,455,480]" italics="true" pageId="59" pageNumber="1786">Pravonitor monteithi</emphasis>
|
||
</taxonomicName>
|
||
had a penis with a large, swollen blind tip, and a basal bulge containing a thickened, folded region, as well as 1–2 penial pilasters. In contrast,
|
||
<taxonomicName id="81D3B288FFDBFFC0DF90FDDDFCAACE59" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[692,814,524,549]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="aquilonia">
|
||
<emphasis id="74A71519FFDBFFC0DF90FDDDFCAACE59" box="[692,814,524,549]" italics="true" pageId="59" pageNumber="1786">P. aquilonia</emphasis>
|
||
</taxonomicName>
|
||
is significantly larger, and has a penis that is narrowed proximally with no blind tip, no penial pilasters, and no thickened, folded region basally.Hence,the two species are retained as separate based on morphological differences despite the close relationship shown in the mitochondrial tree. It is possible that these species have diverged very recently and that the morphological differences are not yet reflected in the tree.
|
||
</paragraph>
|
||
<paragraph id="466CC90BFFDBFFC0DDE4FD0DFF7CCF94" blockId="59.[159,1156,142,1000]" pageId="59" pageNumber="1786">
|
||
This species is recorded from rainforest in the Atherton Tablelands and adjacent coastal ranges (
|
||
<bibRefCitation id="2242B4FAFFDBFFC0DC75FD2EFDA3CF64" author="Stanisic J & Shea M & Potter D & Griffiths O" box="[337,551,767,792]" pageId="59" pageNumber="1786" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic et al. 2010</bibRefCitation>
|
||
); however, in the current study only specimens from Lamb Range and from the adjacent northern parts of the Atherton Tableland (e.g. Tinaroo, Lake Eacham) could be confirmed as belonging to
|
||
<taxonomicName id="81D3B288FFDBFFC0DE51FC95FB84CF21" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[885,1024,836,861]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="monteithi">
|
||
<emphasis id="74A71519FFDBFFC0DE51FC95FB84CF21" box="[885,1024,836,861]" italics="true" pageId="59" pageNumber="1786">P. monteithi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DEE8D58EFFDBFFC0D937FC95FBF2CF21" box="[1043,1142,836,861]" captionStart="Figure 4" captionStartId="27.[160,225,1613,1636]" captionTargetBox="[255,1060,142,1581]" captionTargetId="figure-3@27.[254,1061,142,1582]" captionTargetPageId="27" captionText="Figure 4. Distribution map. Grey symbols: Geminitor. Blue symbols: Pravonitor. Red symbols: Elatonitor. Biogeographic barriers are marked in black; A = Normanby Basin, B = Black Mountain Corridor, and C = Atherton Tableland." figureDoi="http://doi.org/10.5281/zenodo.7380691" httpUri="https://zenodo.org/record/7380691/files/figure.png" pageId="59" pageNumber="1786">Figure 4</figureCitation>
|
||
). Other specimens from the Atherton Tableland preliminarily identified as
|
||
<taxonomicName id="81D3B288FFDBFFC0DEDFFCB6FB07CFFC" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[1019,1155,871,896]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="monteithi">
|
||
<emphasis id="74A71519FFDBFFC0DEDFFCB6FB07CFFC" box="[1019,1155,871,896]" italics="true" pageId="59" pageNumber="1786">P. monteithi</emphasis>
|
||
</taxonomicName>
|
||
were found to be either
|
||
<taxonomicName id="81D3B288FFDBFFC0DC9AFC5BFDBACFDF" box="[446,574,906,931]" class="Gastropoda" family="Helicarionidae" genus="Elatonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="suturalis">
|
||
<emphasis id="74A71519FFDBFFC0DC9AFC5BFDBACFDF" box="[446,574,906,931]" italics="true" pageId="59" pageNumber="1786">M. suturalis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="81D3B288FFDBFFC0DF43FC5BFD6FCFDF" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[615,747,906,931]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="1786" phylum="Mollusca" rank="species" species="aquilonia">
|
||
<emphasis id="74A71519FFDBFFC0DF43FC5BFD6FCFDF" box="[615,747,906,931]" italics="true" pageId="59" pageNumber="1786">P. aquilonia</emphasis>
|
||
</taxonomicName>
|
||
. Further collecting throughout this area to confirm the distribution of these three very similar species would be of great benefit.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |