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<document id="6D7E68DC35179E36175133C7B0E56237" ID-CLB-Dataset="198290" ID-DOI="10.1080/00222933.2022.2136017" ID-GBIF-Dataset="c9288626-198f-4b45-b923-f134cc0df48a" ID-ISSN="1464-5262" ID-Zenodo-Dep="7380681" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669805002707" checkinUser="plazi" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2022" docId="CE7A781DFFD7FFC2DCC6FEB5FDCDC833" docLanguage="en" docName="JNATHIST.56.41-44.1727-1799.pdf" docOrigin="Journal of Natural History 56 (41 - 44)" docSource="http://dx.doi.org/10.1080/00222933.2022.2136017" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Pravonitor kreffti" docType="treatment" docVersion="9" lastPageNumber="1784" masterDocId="32430065FFE0FFFBDD24FFD1FF84CC7C" masterDocTitle="Homoplasy in shells discombobulated the taxonomy: revision of the larger helicarionid land snails of northern Queensland, Australia (Stylommatophora: Helicarionidae)" masterLastPageNumber="1799" masterPageNumber="1727" pageNumber="1782" updateTime="1699412228230" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="01D0B6E849F23C2058017F2BE33EF681">Homoplasy in shells discombobulated the taxonomy: revision of the larger helicarionid land snails of northern Queensland, Australia (Stylommatophora: Helicarionidae)</mods:title>
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<mods:namePart id="2E01082499430F0551F4E6143BBE40F3">Hyman, Isabel T.</mods:namePart>
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<mods:title id="BAB9648AB24EF5A07FCDF9CAD06E73DB">Journal of Natural History</mods:title>
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<mods:date id="F31AB066654DB4633DC85AD351EB4FBF">2022</mods:date>
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<taxonomicName id="81D3B288FFD7FFCCDCC6FEB5FCC5CD02" ID-CoL="6VXCV" authority="(Cox, 1864)" baseAuthorityName="Cox" baseAuthorityYear="1864" box="[482,833,356,382]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="1782" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD7FFCCDCC6FEB5FD32CD02" bold="true" box="[482,694,356,382]" italics="true" pageId="55" pageNumber="1782">Pravonitor kreffti</emphasis>
(
<bibRefCitation id="2242B4FAFFD7FFCCDFE0FEB5FCBDCD02" author="Cox JC" box="[708,825,356,382]" pageId="55" pageNumber="1782" refId="ref29536" refString="Cox JC. 1864. Catalogue of specimens of the Australian land shells in the collection of James C. Cox. Sydney: John Alex Engel. doi: 10.5962 / bhl. title. 12929." type="book" year="1864">Cox, 1864</bibRefCitation>
)
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<paragraph id="466CC90BFFD7FFCCDF2BFE72FC90CDC1" blockId="55.[527,788,419,445]" box="[527,788,419,445]" pageId="55" pageNumber="1782">
(
<figureCitation id="DEE8D58EFFD7FFCCDF33FE72FD07CDC1" box="[535,643,419,445]" captionStart="Figure 4" captionStartId="27.[160,225,1613,1636]" captionTargetBox="[255,1060,142,1581]" captionTargetId="figure-3@27.[254,1061,142,1582]" captionTargetPageId="27" captionText="Figure 4. Distribution map. Grey symbols: Geminitor. Blue symbols: Pravonitor. Red symbols: Elatonitor. Biogeographic barriers are marked in black; A = Normanby Basin, B = Black Mountain Corridor, and C = Atherton Tableland." figureDoi="http://doi.org/10.5281/zenodo.7380691" httpUri="https://zenodo.org/record/7380691/files/figure.png" pageId="55" pageNumber="1782">Figures 4</figureCitation>
,
<figureCitation id="DEE8D58EFFD7FFCCDFB5FE72FD34CDC1" box="[657,688,419,445]" captionStart="Figure 14" captionStartId="47.[160,225,1043,1066]" captionTargetBox="[164,1151,143,1010]" captionTargetId="figure-168@47.[163,1152,142,1011]" captionTargetPageId="47" captionText="Figure 14. Shells of Pravonitor. a) Pravonitor insularum, AM C.170795 (paratype). b) Pravonitor annulus AM C.425415,Gabba Is. c) Pravonitor aquilonia, QM MO48285 (holotype). d) Pravonitor ferrugineus, QM MO12752 (holotype). e). Pravonitor kreffti, AM C.1352, Thursday Is. f) Pravonitor kreffti, AM C.425474, Simpsons Bay, Utingu. g) Pravonitor monteithi, QM MO48587 (holotype). h) Pravonitor septentrionalis, AM C.425422. i) Pravonitor stuarti, QM MO80213 (paratype). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380720" httpUri="https://zenodo.org/record/7380720/files/figure.png" pageId="55" pageNumber="1782">14</figureCitation>
(ef), 18)
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<treatmentCitation id="C772EF1AFFD7FFCCDD84FE09FE3ECD8D" author="Cox JC" box="[160,442,472,497]" page="21" pageId="55" pageNumber="1782" year="1864">
<taxonomicName id="81D3B288FFD7FFCCDD84FE09FE3ECD8D" authority="Cox, 1864: 21" authorityName="Cox" authorityPageNumber="21" authorityYear="1864" box="[160,442,472,497]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="1782" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD7FFCCDD84FE09FE9ECD8D" box="[160,282,472,497]" italics="true" pageId="55" pageNumber="1782">Helix kreffti</emphasis>
<bibRefCitation id="2242B4FAFFD7FFCCDC06FE09FE3ECD8D" author="Cox JC" box="[290,442,472,497]" pageId="55" pageNumber="1782" refId="ref29536" refString="Cox JC. 1864. Catalogue of specimens of the Australian land shells in the collection of James C. Cox. Sydney: John Alex Engel. doi: 10.5962 / bhl. title. 12929." type="book" year="1864">Cox, 1864: 21</bibRefCitation>
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.
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<treatmentCitationGroup id="66C3EE25FFD7FFCCDD84FE2BFDEDCE6F" box="[160,617,506,531]" pageId="55" pageNumber="1782">
<treatmentCitation id="C772EF1AFFD7FFCCDD84FE2BFDE2CE6F" author="Brazier J" box="[160,614,506,531]" page="118" pageId="55" pageNumber="1782" year="1876">
<taxonomicName id="81D3B288FFD7FFCCDD84FE2BFDE2CE6F" authority=": Brazier 1876: 118" authorityName="Brazier" authorityPageNumber="118" authorityYear="1876" box="[160,614,506,531]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="1782" phylum="Mollusca" rank="species" species="kreffti" subGenus="Thalassia">
<emphasis id="74A71519FFD7FFCCDD84FE2BFF51CE6F" box="[160,213,506,531]" italics="true" pageId="55" pageNumber="1782">Helix</emphasis>
(
<emphasis id="74A71519FFD7FFCCDDC0FE2BFECDCE6F" box="[228,329,506,531]" italics="true" pageId="55" pageNumber="1782">Thalassia</emphasis>
)
<emphasis id="74A71519FFD7FFCCDC7CFE2BFE12CE6F" box="[344,406,506,531]" italics="true" pageId="55" pageNumber="1782">kreffti</emphasis>
:
<bibRefCitation id="2242B4FAFFD7FFCCDC87FE2BFDE2CE6F" author="Brazier J" box="[419,614,506,531]" pageId="55" pageNumber="1782" pagination="117 - 133" refId="ref29454" refString="Brazier J. 1876. List of land shells collected during the Chevert expedition. Proc Linn Soc NSW. 1: 117 - 133. doi: 10.5962 / bhl. part. 12392" type="journal article" year="1876">Brazier 1876: 118</bibRefCitation>
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.
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<treatmentCitationGroup id="66C3EE25FFD7FFCCDD84FDCCFBC4CE4A" box="[160,1088,541,566]" pageId="55" pageNumber="1782">
<taxonomicName id="81D3B288FFD7FFCCDD84FDCCFCF6CE4A" ID-CoL="af2598cd-4976-485b-9186-ab5b979149de" authority="Sect." authorityName="Sect." authorityPageNumber="219" authorityYear="1847" box="[160,882,541,566]" class="Gastropoda" family="Ariophantidae" genus="Nanina" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="1782" phylum="Mollusca" rank="genus">
<emphasis id="74A71519FFD7FFCCDD84FDCCFF6BCE4A" box="[160,239,541,566]" italics="true" pageId="55" pageNumber="1782">Nanina</emphasis>
(Subg.
<emphasis id="74A71519FFD7FFCCDC61FDCCFEFBCE4A" box="[325,383,541,566]" italics="true" pageId="55" pageNumber="1782">Xesta</emphasis>
Sect.
<emphasis id="74A71519FFD7FFCCDCE7FDCCFDE4CE4A" box="[451,608,541,566]" italics="true" pageId="55" pageNumber="1782">Macrochlamys</emphasis>
)
<treatmentCitation id="C772EF1AFFD7FFCCDF4BFDCCFCF6CE4A" author="Tryon GW" box="[623,882,541,566]" page="219" pageId="55" pageNumber="1782" year="1886">
<emphasis id="74A71519FFD7FFCCDF4BFDCCFD29CE4A" box="[623,685,541,566]" italics="true" pageId="55" pageNumber="1782">kreffti</emphasis>
:
<bibRefCitation id="2242B4FAFFD7FFCCDF9EFDCCFCF6CE4A" author="Tryon GW" box="[698,882,541,566]" pageId="55" pageNumber="1782" refId="ref32172" refString="Tryon GW jnr. 1886. Manual of conchology; structural and systematic. Second series: pulmonata. Volume 2. Zonitidae. Philadelphia: G. W. Tryon, jnr." type="book" year="1886">Tryon 1886: 219</bibRefCitation>
</treatmentCitation>
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, pl. 63, figs 9899.
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<treatmentCitationGroup id="66C3EE25FFD7FFCCDD84FD91FDFBCE25" box="[160,639,576,601]" pageId="55" pageNumber="1782">
<treatmentCitation id="C772EF1AFFD7FFCCDD84FD91FDFFCE25" author="Cox JC" box="[160,635,576,601]" page="6" pageId="55" pageNumber="1782" year="1909">
<taxonomicName id="81D3B288FFD7FFCCDD84FD91FDFFCE25" authority=": Cox 1909: 6" authorityName="Cox" authorityPageNumber="6" authorityYear="1909" box="[160,635,576,601]" class="Gastropoda" family="Ariophantidae" genus="Nanina" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="1782" phylum="Mollusca" rank="species" species="kreffti" subGenus="Macrochlamys">
<emphasis id="74A71519FFD7FFCCDD84FD91FF6BCE25" box="[160,239,576,601]" italics="true" pageId="55" pageNumber="1782">Nanina</emphasis>
(
<emphasis id="74A71519FFD7FFCCDDDAFD91FE1FCE25" box="[254,411,576,601]" italics="true" pageId="55" pageNumber="1782">Macrochlamys</emphasis>
)
<emphasis id="74A71519FFD7FFCCDC8FFD91FE6DCE25" box="[427,489,576,601]" italics="true" pageId="55" pageNumber="1782">kreffti</emphasis>
:
<bibRefCitation id="2242B4FAFFD7FFCCDCD2FD91FDFFCE25" author="Cox JC" box="[502,635,576,601]" pageId="55" pageNumber="1782" refId="ref29601" refString="Cox JC. 1909. A list of the subclass Pulmonata found in Australia, Part I., not including Tasmania, Lord Howe's Island (under the NSW Government), or the New Guinea Mainland and adjacent islands. Sydney: Frederick W. White." type="book" year="1909">Cox 1909: 6</bibRefCitation>
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.
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</paragraph>
<paragraph id="466CC90BFFD7FFCCDD84FDB3FDDFCE07" blockId="55.[160,1088,472,670]" box="[160,603,610,635]" pageId="55" pageNumber="1782">
<treatmentCitationGroup id="66C3EE25FFD7FFCCDD84FDB3FDDFCE07" box="[160,603,610,635]" pageId="55" pageNumber="1782">
<treatmentCitation id="C772EF1AFFD7FFCCDD84FDB3FDD3CE07" author="Iredale T" box="[160,599,610,635]" page="3" pageId="55" pageNumber="1782" year="1937">
<taxonomicName id="81D3B288FFD7FFCCDD84FDB3FDD3CE07" ID-CoL="6RZ4Q" authority=": Iredale 1937: 3" authorityName=": Iredale" authorityPageNumber="3" authorityYear="1937" box="[160,599,610,635]" class="Gastropoda" family="Helicarionidae" genus="Nitor" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="1782" phylum="Mollusca" rank="species" species="kreffti" subGenus="Pravonitor">
<emphasis id="74A71519FFD7FFCCDD84FDB3FF52CE07" box="[160,214,610,635]" italics="true" pageId="55" pageNumber="1782">Nitor</emphasis>
(
<emphasis id="74A71519FFD7FFCCDDC1FDB3FED2CE07" box="[229,342,610,635]" italics="true" pageId="55" pageNumber="1782">Pravonitor</emphasis>
)
<emphasis id="74A71519FFD7FFCCDC41FDB3FE27CE07" box="[357,419,610,635]" italics="true" pageId="55" pageNumber="1782">kreffti</emphasis>
:
<bibRefCitation id="2242B4FAFFD7FFCCDC94FDB3FDD3CE07" author="Iredale T" box="[432,599,610,635]" pageId="55" pageNumber="1782" pagination="1 - 39" refId="ref30613" refString="Iredale T. 1937. A basic list of the land Mollusca of Australia. Part II. Aust Zool. 9: 1 - 39." type="journal article" year="1937">Iredale 1937: 3</bibRefCitation>
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.
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<paragraph id="466CC90BFFD7FFCCDD84FD54FC54CEE2" blockId="55.[160,1088,472,670]" box="[160,976,645,670]" pageId="55" pageNumber="1782">
<treatmentCitationGroup id="66C3EE25FFD7FFCCDD84FD54FC54CEE2" box="[160,976,645,670]" pageId="55" pageNumber="1782">
<treatmentCitation id="C772EF1AFFD7FFCCDD84FD54FDA5CEE2" author="Smith BJ" box="[160,545,645,670]" page="241" pageId="55" pageNumber="1782" year="1992">
<taxonomicName id="81D3B288FFD7FFCCDD84FD54FDA5CEE2" ID-CoL="6VXCV" authority=": Smith, 1992: 241" authorityName="Smith" authorityPageNumber="241" authorityYear="1992" box="[160,545,645,670]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="1782" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD7FFCCDD84FD54FED3CEE2" box="[160,343,645,670]" italics="true" pageId="55" pageNumber="1782">Pravonitor kreffti</emphasis>
:
<bibRefCitation id="2242B4FAFFD7FFCCDC40FD54FDA5CEE2" author="Smith BJ" box="[356,545,645,670]" pageId="55" pageNumber="1782" pagination="1 - 408" refId="ref31733" refString="Smith BJ 1992. Non-marine Mollusca. In: Houston W, editor. Zoological catalogue of Australia (Vol. 8). Canberra: Australian Government Publishing Service; pp. 1 - 408." type="book chapter" year="1992">Smith, 1992: 241</bibRefCitation>
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;
<treatmentCitation id="C772EF1AFFD7FFCCDF08FD54FCEDCEE2" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[556,873,645,670]" page="300" pageId="55" pageNumber="1782" year="2010">
<bibRefCitation id="2242B4FAFFD7FFCCDF08FD54FCEDCEE2" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[556,873,645,670]" pageId="55" pageNumber="1782" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic et al. 2010: 300301</bibRefCitation>
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(in part).
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<emphasis id="74A71519FFD7FFCCDD84F95CFE8DCAD8" bold="true" box="[160,265,1677,1700]" pageId="55" pageNumber="1782">Figure 18.</emphasis>
Genitalia of
<taxonomicName id="81D3B288FFD7FFCCDCA9F95CFD0FCAD8" authority=", QM MO" authorityName="QM MO" box="[397,651,1677,1700]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="1782" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD7FFCCDCA9F95CFDB4CAD8" box="[397,560,1677,1700]" italics="true" pageId="55" pageNumber="1782">Pravonitor kreffti</emphasis>
, QM MO
</taxonomicName>
86003 (neotype), Thursday Island. a) Reproductive system. b) Penis interior. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="0EC99A80FFD8FFC3DD84FF5FFE69CD9C" pageId="56" pageNumber="1783" type="materials_examined">
<paragraph id="466CC90BFFD8FFC3DD84FF5FFEFBCCD4" blockId="56.[160,1156,142,237]" box="[160,383,142,168]" pageId="56" pageNumber="1783">
<emphasis id="74A71519FFD8FFC3DD84FF5FFEFBCCD4" bold="true" box="[160,383,142,168]" italics="true" pageId="56" pageNumber="1783">Material examined</emphasis>
</paragraph>
<paragraph id="466CC90BFFD8FFC3DD84FF60FEB9CC91" blockId="56.[160,1156,142,237]" pageId="56" pageNumber="1783">
<materialsCitation id="F6BBC356FFD8FFC3DD84FF60FEB9CC91" ID-GBIF-Occurrence="3978770318" country="Australia" county="North Qld" location="Status" municipality="Far North Qld" pageId="56" pageNumber="1783" specimenCount="1" typeStatus="holotype">
<emphasis id="74A71519FFD8FFC3DD84FF60FE96CCB7" bold="true" box="[160,274,177,203]" italics="true" pageId="56" pageNumber="1783">
<typeStatus id="996877A9FFD8FFC3DD84FF60FE89CCB7" box="[160,269,177,203]" pageId="56" pageNumber="1783" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="430C9FD0FFD8FFC3DC05FF63FEE2CCB7" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD8FFC3DC05FF63FEE2CCB7" box="[289,358,178,203]" country="Australia" county="North Qld" municipality="Far North Qld" name="Status" pageId="56" pageNumber="1783">Status</location>
unknown, whereabouts unknown presumed lost,
<location id="430C9FD0FFD8FFC3DEBFFF63FB8BCCB7" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD8FFC3DEBFFF63FB8BCCB7" box="[923,1039,178,203]" country="Australia" county="North Qld" municipality="Far North Qld" name="Cape York" pageId="56" pageNumber="1783">Cape York</location>
,
<collectingMunicipality id="A6085371FFD8FFC3D93DFF63FF4ECC91" pageId="56" pageNumber="1783">
Far
<collectingCounty id="AF0DB187FFD8FFC3D967FF63FF4ECC91" pageId="56" pageNumber="1783">North Qld</collectingCounty>
</collectingMunicipality>
,
<collectingCountry id="3EC4899BFFD8FFC3DDF1FF05FEBDCC91" box="[213,313,212,237]" name="Australia" pageId="56" pageNumber="1783">Australia</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="466CC90BFFD8FFC3DD84FEC8FD60CD29" blockId="56.[160,1155,281,341]" pageId="56" pageNumber="1783">
<materialsCitation id="F6BBC356FFD8FFC3DD84FEC8FD60CD29" ID-GBIF-Occurrence="3978770317" collectingDate="1985-04-11" collectionCode="QM" collectorName="J. Stanisic &amp; D. Potter." country="Australia" county="North Qld" location="Green Hill" municipality="Far North Qld" pageId="56" pageNumber="1783" specimenCode="MO86003" specimenCount="1" typeStatus="neotype">
<emphasis id="74A71519FFD8FFC3DD84FEC8FE8DCD4F" bold="true" box="[160,265,281,307]" italics="true" pageId="56" pageNumber="1783">
<typeStatus id="996877A9FFD8FFC3DD84FEC8FE80CD4F" box="[160,260,281,307]" pageId="56" pageNumber="1783" type="neotype">Neotype</typeStatus>
.
</emphasis>
<collectionCode id="20C251CEFFD8FFC3DC3DFECBFEC5CD4F" box="[281,321,282,307]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="56" pageNumber="1783" type="Museum">QM</collectionCode>
<specimenCode id="16756170FFD8FFC3DC62FECBFE31CD4F" box="[326,437,282,307]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="56" pageNumber="1783" type="Museum">MO86003</specimenCode>
.
<location id="430C9FD0FFD8FFC3DC9AFECBFDA9CD4F" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD8FFC3DC9AFECBFDA9CD4F" box="[446,557,282,307]" country="Australia" county="North Qld" municipality="Far North Qld" name="Green Hill" pageId="56" pageNumber="1783">Green Hill</location>
,
<location id="430C9FD0FFD8FFC3DF12FECBFD63CD4F" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD8FFC3DF12FECBFD63CD4F" box="[566,743,282,307]" country="Australia" county="North Qld" municipality="Far North Qld" name="Thursday Island" pageId="56" pageNumber="1783">Thursday Island</location>
,
<location id="430C9FD0FFD8FFC3DFD4FECBFCFCCD4F" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD8FFC3DFD4FECBFCFCCD4F" box="[752,888,282,307]" country="Australia" county="North Qld" municipality="Far North Qld" name="Torres Strait" pageId="56" pageNumber="1783">Torres Strait</location>
,
<collectingMunicipality id="A6085371FFD8FFC3DEA4FECBFB9DCD4F" box="[896,1049,282,307]" pageId="56" pageNumber="1783">
Far
<collectingCounty id="AF0DB187FFD8FFC3DE8DFECBFB9DCD4F" box="[937,1049,282,307]" pageId="56" pageNumber="1783">North Qld</collectingCounty>
</collectingMunicipality>
,
<collectingCountry id="3EC4899BFFD8FFC3D906FECBFB07CD4F" box="[1058,1155,282,307]" name="Australia" pageId="56" pageNumber="1783">Australia</collectingCountry>
(10.58, 142.28),
<date id="326DEFCBFFD8FFC3DC7BFEEDFE73CD29" box="[351,503,316,341]" pageId="56" pageNumber="1783" value="1985-04-11">
<collectingDate id="22291623FFD8FFC3DC7BFEEDFE73CD29" box="[351,503,316,341]" pageId="56" pageNumber="1783" value="1985-04-11">11 April 1985</collectingDate>
</date>
,
<collectorName id="EB26ACDDFFD8FFC3DF26FEEDFDF4CD29" box="[514,624,316,341]" pageId="56" pageNumber="1783">J. Stanisic</collectorName>
,
<collectorName id="EB26ACDDFFD8FFC3DF5EFEEDFD60CD29" box="[634,740,316,341]" pageId="56" pageNumber="1783">D. Potter.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="466CC90BFFD8FFC3DD84FE50FD79CDE7" blockId="56.[160,765,385,411]" box="[160,765,385,411]" pageId="56" pageNumber="1783">
<emphasis id="74A71519FFD8FFC3DD84FE50FEB9CDE7" bold="true" box="[160,317,385,411]" italics="true" pageId="56" pageNumber="1783">Paraneotype.</emphasis>
<materialsCitation id="F6BBC356FFD8FFC3DC69FE53FD7CCDE7" ID-GBIF-Occurrence="3978770336" box="[333,760,386,411]" collectingDate="1985-04-11" collectionCode="QM" collectorName="J. Stanisic &amp; D. Potter." country="Australia" county="North Qld" location="Green Hill" municipality="Far North Qld" pageId="56" pageNumber="1783" specimenCode="MO15572" specimenCount="1">
<collectionCode id="20C251CEFFD8FFC3DC69FE53FEF1CDE7" box="[333,373,386,411]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="56" pageNumber="1783" type="Museum">QM</collectionCode>
<specimenCode id="16756170FFD8FFC3DC58FE53FE6DCDE7" box="[380,489,386,411]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="56" pageNumber="1783" type="Museum">MO15572</specimenCode>
(same data as neotype)
</materialsCitation>
.
</paragraph>
<paragraph id="466CC90BFFD8FFC3DD84FE17FE69CD9C" blockId="56.[160,493,454,480]" box="[160,493,454,480]" pageId="56" pageNumber="1783">
<emphasis id="74A71519FFD8FFC3DD84FE17FED1CD9C" bold="true" box="[160,341,454,480]" italics="true" pageId="56" pageNumber="1783">Other material.</emphasis>
See
<tableCitation id="0B51FCB0FFD8FFC3DCB1FE16FE63CD9C" box="[405,487,455,480]" captionStart="Table 1" captionStartId="5.[133,189,450,473]" captionTargetPageId="5" captionText="Table 1. Material studied. Abbreviations: H, holotype; L, lectotype; N, neotype; P, paratype; PL, paralectotype, PN, paraneotype; S, syntype." httpUri="http://table.plazi.org/id/12AC9983FFE5FFFEDDA1FE13FA67CDA5" pageId="56" pageNumber="1783" tableUuid="12AC9983FFE5FFFEDDA1FE13FA67CDA5">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0EC99A80FFD8FFC3DD84FDDDFD93C8EA" pageId="56" pageNumber="1783" type="diagnosis">
<paragraph id="466CC90BFFD8FFC3DD84FDDDFE90CE5A" blockId="56.[160,1155,524,792]" box="[160,276,524,550]" pageId="56" pageNumber="1783">
<emphasis id="74A71519FFD8FFC3DD84FDDDFE90CE5A" bold="true" box="[160,276,524,550]" italics="true" pageId="56" pageNumber="1783">Diagnosis</emphasis>
</paragraph>
<paragraph id="466CC90BFFD8FFC3DD84FDFFFF5BCF64" blockId="56.[160,1155,524,792]" pageId="56" pageNumber="1783">
<emphasis id="74A71519FFD8FFC3DD84FDFFFE26CE34" bold="true" box="[160,418,558,584]" italics="true" pageId="56" pageNumber="1783">External morphology.</emphasis>
Shell (
<figureCitation id="DEE8D58EFFD8FFC3DCD3FDFEFDE0CE34" box="[503,612,559,584]" captionStart="Figure 14" captionStartId="47.[160,225,1043,1066]" captionTargetBox="[164,1151,143,1010]" captionTargetId="figure-168@47.[163,1152,142,1011]" captionTargetPageId="47" captionText="Figure 14. Shells of Pravonitor. a) Pravonitor insularum, AM C.170795 (paratype). b) Pravonitor annulus AM C.425415,Gabba Is. c) Pravonitor aquilonia, QM MO48285 (holotype). d) Pravonitor ferrugineus, QM MO12752 (holotype). e). Pravonitor kreffti, AM C.1352, Thursday Is. f) Pravonitor kreffti, AM C.425474, Simpsons Bay, Utingu. g) Pravonitor monteithi, QM MO48587 (holotype). h) Pravonitor septentrionalis, AM C.425422. i) Pravonitor stuarti, QM MO80213 (paratype). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380720" httpUri="https://zenodo.org/record/7380720/files/figure.png" pageId="56" pageNumber="1783">Figure 14</figureCitation>
(ef)) medium-sized (SW
<quantity id="812B64EEFFD8FFC3DE5BFDFEFBA5CE34" box="[895,1057,559,584]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" metricValueMax="1.3699999999999999" metricValueMin="1.25" pageId="56" pageNumber="1783" unit="mm" value="13.1" valueMax="13.7" valueMin="12.5">12.513.7 mm</quantity>
), golden brown, 5.05.7 whorls, subdiscoidal to subglobose with a rounded whorl profile; shell microsculpture with very fine to obsolete spiral grooves; umbilicus very narrow, sealed with a mucous plug, partially covered by reflected lip. Body grey with a cream sole (in alcohol). Right and median mantle lobes moderately small, left lobe small; shell lappets moderately large, narrow, tapering, right lappet longer than left. Caudal horn moderately large.
</paragraph>
<paragraph id="466CC90BFFD8FFC3DD84FC95FD93C8EA" blockId="56.[160,1156,836,1174]" pageId="56" pageNumber="1783">
<emphasis id="74A71519FFD8FFC3DD84FC95FEF5CF22" bold="true" box="[160,369,836,862]" italics="true" pageId="56" pageNumber="1783">Genital anatomy.</emphasis>
Genitalia (
<figureCitation id="DEE8D58EFFD8FFC3DCD1FC94FDE5CF22" box="[501,609,837,862]" captionStart="Figure 18" captionStartId="55.[160,225,1677,1700]" captionTargetBox="[166,1148,765,1645]" captionTargetId="figure-127@55.[166,1122,784,1646]" captionTargetPageId="55" captionText="Figure 18. Genitalia of Pravonitor kreffti, QM MO86003 (neotype), Thursday Island. a) Reproductive system. b) Penis interior. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7380730" httpUri="https://zenodo.org/record/7380730/files/figure.png" pageId="56" pageNumber="1783">Figure 18</figureCitation>
) with short vagina, internally with broad longitudinal pilasters; bursa copulatrix moderately long, duct broad at base, then narrowing, internally with broad longitudinal pilasters; bursa elongate. Penis large, narrow at base and becoming broader proximally, with a large blind tip; internally with no pilasters or with one pustulose longitudinal pilaster, internal wall sculptured with fine pustules; approx. 5075% enclosed in penial tunica; penis sometimes folded longitudinally inside tunica. Epiphallus enters penis through a simple pore; epiphallus similar in length to penis; epiphallus 2 longer than epiphallus 1; moderately short epiphallic caecum present, with medial or apical attachment of the penial retractor muscle; flagellum long, slender, without obvious internal cryptae.
</paragraph>
</subSubSection>
<subSubSection id="0EC99A80FFD8FFC2DD84FB10FDCDC833" lastPageId="57" lastPageNumber="1784" pageId="56" pageNumber="1783" type="discussion">
<paragraph id="466CC90BFFD8FFC3DD84FB10FE82C8A7" blockId="56.[159,1156,1217,1728]" box="[160,262,1217,1243]" pageId="56" pageNumber="1783">
<emphasis id="74A71519FFD8FFC3DD84FB10FE82C8A7" bold="true" box="[160,262,1217,1243]" italics="true" pageId="56" pageNumber="1783">Remarks</emphasis>
</paragraph>
<paragraph id="466CC90BFFD8FFC3DD84FB34FD75CA6F" blockId="56.[159,1156,1217,1728]" pageId="56" pageNumber="1783">
The correct identity of this species requires some clarification. It was first described by
<bibRefCitation id="2242B4FAFFD8FFC3D97DFB34FF62C95C" author="Cox JC" pageId="56" pageNumber="1783" refId="ref29536" refString="Cox JC. 1864. Catalogue of specimens of the Australian land shells in the collection of James C. Cox. Sydney: John Alex Engel. doi: 10.5962 / bhl. title. 12929." type="book" year="1864">Cox (1864)</bibRefCitation>
based on a single specimen collected by MacGillivray, with no figure and with only a brief description (including the dimensions diam. maj. 0.57, min. 0.49, alt 0.31 (uncertain), 6 whorls), and a rather general
<typeStatus id="996877A9FFD8FFC3DF10FA9CFDE2C91A" box="[564,614,1357,1382]" pageId="56" pageNumber="1783">type</typeStatus>
locality of simply Cape York. Shortly afterwards,
<bibRefCitation id="2242B4FAFFD8FFC3DD84FABEFE9CC9F4" author="Cox JC" box="[160,280,1391,1416]" pageId="56" pageNumber="1783" refId="ref29574" refString="Cox JC. 1868. A monograph of Australian land shells. Sydney: William Maddock. doi: 10.5962 / bhl. title. 1292." type="book" year="1868">Cox (1868)</bibRefCitation>
synonymised the species with
<taxonomicName id="81D3B288FFD8FFC3DF5DFABEFD5DC9F4" authorityName="Reeve" authorityYear="1854" box="[633,729,1391,1416]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="1783" phylum="Mollusca" rank="species" species="villaris">
<emphasis id="74A71519FFD8FFC3DF5DFABEFD5DC9F4" box="[633,729,1391,1416]" italics="true" pageId="56" pageNumber="1783">P. villaris</emphasis>
</taxonomicName>
, and figured the specimen under this name, with dimensions (width
<quantity id="812B64EEFFD8FFC3DCD3FA43FDECC9D7" box="[503,616,1426,1451]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="56" pageNumber="1783" unit="in" value="0.5">0.5 inches</quantity>
; height
<quantity id="812B64EEFFD8FFC3DFE4FA43FCBBC9D7" box="[704,831,1426,1451]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" pageId="56" pageNumber="1783" unit="in" value="0.25">0.25 inches</quantity>
, shell 5 whorls) matching the figure given at natural size. Here,
<bibRefCitation id="2242B4FAFFD8FFC3DF3EFA64FD16C9B2" author="Cox JC" box="[538,658,1461,1486]" pageId="56" pageNumber="1783" refId="ref29574" refString="Cox JC. 1868. A monograph of Australian land shells. Sydney: William Maddock. doi: 10.5962 / bhl. title. 1292." type="book" year="1868">Cox (1868)</bibRefCitation>
gives the locality as Cape
<collectingRegion id="841707E9FFD8FFC3DEE5FA64FC77C9B2" box="[961,1011,1461,1486]" country="United Kingdom" name="York" pageId="56" pageNumber="1783">York</collectingRegion>
(Cox); Rocky Isles, near Cape Flattery (MacGillivray); however, this second locality is probably referring to
<taxonomicName id="81D3B288FFD8FFC3DD9BFA2BFEA6CA6F" authorityName="Reeve" authorityYear="1854" box="[191,290,1530,1555]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="1783" phylum="Mollusca" rank="species" species="villaris">
<emphasis id="74A71519FFD8FFC3DD9BFA2BFEA6CA6F" box="[191,290,1530,1555]" italics="true" pageId="56" pageNumber="1783">G. villaris</emphasis>
</taxonomicName>
, which was described from Lizard Island.
</paragraph>
<paragraph id="466CC90BFFD8FFC3DDE4F9CCFBBACABC" blockId="56.[159,1156,1217,1728]" pageId="56" pageNumber="1783">
Subsequent authors including
<bibRefCitation id="2242B4FAFFD8FFC3DF27F9CCFD10CA4A" author="Brazier J" box="[515,660,1565,1590]" pageId="56" pageNumber="1783" pagination="117 - 133" refId="ref29454" refString="Brazier J. 1876. List of land shells collected during the Chevert expedition. Proc Linn Soc NSW. 1: 117 - 133. doi: 10.5962 / bhl. part. 12392" type="journal article" year="1876">Brazier (1876)</bibRefCitation>
,
<bibRefCitation id="2242B4FAFFD8FFC3DFB8F9CCFCA6CA4A" author="Tryon GW" box="[668,802,1565,1590]" pageId="56" pageNumber="1783" refId="ref32172" refString="Tryon GW jnr. 1886. Manual of conchology; structural and systematic. Second series: pulmonata. Volume 2. Zonitidae. Philadelphia: G. W. Tryon, jnr." type="book" year="1886">Tryon (1886)</bibRefCitation>
and
<bibRefCitation id="2242B4FAFFD8FFC3DE77F9CCFC40CA4A" author="Cox JC" box="[851,964,1565,1590]" pageId="56" pageNumber="1783" refId="ref29601" refString="Cox JC. 1909. A list of the subclass Pulmonata found in Australia, Part I., not including Tasmania, Lord Howe's Island (under the NSW Government), or the New Guinea Mainland and adjacent islands. Sydney: Frederick W. White." type="book" year="1909">Cox (1909)</bibRefCitation>
treated
<taxonomicName id="81D3B288FFD8FFC3D93DF9CCFBD0CA4A" box="[1049,1108,1565,1590]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="1783" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD8FFC3D93DF9CCFBD0CA4A" box="[1049,1108,1565,1590]" italics="true" pageId="56" pageNumber="1783">kreffti</emphasis>
</taxonomicName>
and
<taxonomicName id="81D3B288FFD8FFC3DDBBF9EEFF64CA24" authorityName="Reeve" authorityYear="1854" box="[159,224,1599,1624]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="1783" phylum="Mollusca" rank="species" species="villaris">
<emphasis id="74A71519FFD8FFC3DDBBF9EEFF64CA24" box="[159,224,1599,1624]" italics="true" pageId="56" pageNumber="1783">villaris</emphasis>
</taxonomicName>
as separate species.
<bibRefCitation id="2242B4FAFFD8FFC3DC9EF9EEFDC8CA24" author="Brazier J" box="[442,588,1599,1624]" pageId="56" pageNumber="1783" pagination="117 - 133" refId="ref29454" refString="Brazier J. 1876. List of land shells collected during the Chevert expedition. Proc Linn Soc NSW. 1: 117 - 133. doi: 10.5962 / bhl. part. 12392" type="journal article" year="1876">Brazier (1876)</bibRefCitation>
stated that he believed Coxs
<typeStatus id="996877A9FFD8FFC3DEA2F9EEFC32CA24" box="[902,950,1599,1624]" pageId="56" pageNumber="1783">type</typeStatus>
shell to come from Darnley Island in the Torres Strait, given the close similarity of the
<taxonomicName id="81D3B288FFD8FFC3DE49F9B3FC2CCA07" box="[877,936,1634,1659]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="1783" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD8FFC3DE49F9B3FC2CCA07" box="[877,936,1634,1659]" italics="true" pageId="56" pageNumber="1783">kreffti</emphasis>
</taxonomicName>
<typeStatus id="996877A9FFD8FFC3DE94F9B3FC64CA07" box="[944,992,1634,1659]" pageId="56" pageNumber="1783">type</typeStatus>
to the Darnley Island species,and recorded
<taxonomicName id="81D3B288FFD8FFC3DCEBF954FD94CAE2" authorityName="Reeve" authorityYear="1854" box="[463,528,1669,1694]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="1783" phylum="Mollusca" rank="species" species="villaris">
<emphasis id="74A71519FFD8FFC3DCEBF954FD94CAE2" box="[463,528,1669,1694]" italics="true" pageId="56" pageNumber="1783">villaris</emphasis>
</taxonomicName>
from Cape
<collectingRegion id="841707E9FFD8FFC3DFADF954FD3ECAE2" box="[649,698,1669,1694]" country="United Kingdom" name="York" pageId="56" pageNumber="1783">York</collectingRegion>
and Albany Island (this latter reference was probably actually
<taxonomicName id="81D3B288FFD8FFC3DC7BF976FD8ACABC" box="[351,526,1703,1728]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="1783" phylum="Mollusca" rank="species" species="septentrionalis">
<emphasis id="74A71519FFD8FFC3DC7BF976FD8ACABC" box="[351,526,1703,1728]" italics="true" pageId="56" pageNumber="1783">P. septentrionalis</emphasis>
</taxonomicName>
, since
<taxonomicName id="81D3B288FFD8FFC3DF72F976FD37CABC" authorityName="Reeve" authorityYear="1854" box="[598,691,1703,1728]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="1783" phylum="Mollusca" rank="species" species="villaris">
<emphasis id="74A71519FFD8FFC3DF72F976FD37CABC" box="[598,691,1703,1728]" italics="true" pageId="56" pageNumber="1783">G. villaris</emphasis>
</taxonomicName>
is found considerably farther south).
</paragraph>
<paragraph id="466CC90BFFD9FFC2DDE4FF5FFD07CED3" blockId="57.[160,1157,142,1103]" pageId="57" pageNumber="1784">
<materialsCitation id="F6BBC356FFD9FFC2DDE4FF5FFB04CCB6" ID-GBIF-Occurrence="3978770333" location="Among" municipality="Cape York" pageId="57" pageNumber="1784" specimenCount="1">
<location id="430C9FD0FFD9FFC2DDE4FF5FFE91CCDB" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DDE4FF5FFE91CCDB" box="[192,277,142,167]" municipality="Cape York" name="Among" pageId="57" pageNumber="1784">Among</location>
material from
<collectingMunicipality id="A6085371FFD9FFC2DC9AFF5FFDB1CCDB" box="[446,565,142,167]" pageId="57" pageNumber="1784">Cape York</collectingMunicipality>
, there are two species represented: one is relatively small (shells of 5.05.5 whorls are
<quantity id="812B64EEFFD9FFC2DF14FF60FD33CCB6" box="[560,695,177,202]" metricMagnitude="-3" metricUnit="m" metricValue="8.55" metricValueMax="9.3" metricValueMin="7.8" pageId="57" pageNumber="1784" unit="mm" value="8.55" valueMax="9.3" valueMin="7.8">7.89.3 mm</quantity>
in width) and is referred to above as
<taxonomicName id="81D3B288FFD9FFC2D954FF60FB04CCB6" authority="' Cape" authorityName="Cape" box="[1136,1152,177,202]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD9FFC2D954FF60FB04CCB6" box="[1136,1152,177,202]" italics="true" pageId="57" pageNumber="1784">P</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="74A71519FFD9FFC2D9A5FF60FF65CC91" italics="true" pageId="57" pageNumber="1784">. kreffti</emphasis>
Cape York.
<materialsCitation id="F6BBC356FFD9FFC2DC53FF05FC93CD4E" ID-GBIF-Occurrence="3978770344" location="The" municipality="Near Western Torres Strait Islands" pageId="57" pageNumber="1784" specimenCount="1">
<location id="430C9FD0FFD9FFC2DC53FF05FE25CC91" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DC53FF05FE25CC91" box="[375,417,212,237]" municipality="Near Western Torres Strait Islands" name="The" pageId="57" pageNumber="1784">The</location>
other is much larger (shells of 5.35.8 whorls are
<quantity id="812B64EEFFD9FFC2DEC5FF05FB00CC91" box="[993,1156,212,237]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.53" metricValueMin="1.29" pageId="57" pageNumber="1784" unit="mm" value="14.1" valueMax="15.3" valueMin="12.9">12.915.3 mm</quantity>
in width) and is conspecific with a group found primarily on the Inner and
<collectingMunicipality id="A6085371FFD9FFC2D969FF27FE7DCD4E" pageId="57" pageNumber="1784">Near Western Torres Strait Islands</collectingMunicipality>
, referred to above as
<taxonomicName id="81D3B288FFD9FFC2DE23FEC8FC93CD4E" authority="' Inner" authorityName="Inner" box="[775,791,281,306]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD9FFC2DE23FEC8FC93CD4E" box="[775,791,281,306]" italics="true" pageId="57" pageNumber="1784">P</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="74A71519FFD9FFC2DE33FEC8FC9FCD4E" box="[791,795,281,306]" italics="true" pageId="57" pageNumber="1784">.</emphasis>
<materialsCitation id="F6BBC356FFD9FFC2DE01FEC8FE9ECD0B" ID-GBIF-Occurrence="3978770352" collectionCode="AM" collectorName="Cape York &amp; Utingu" location="Western Islands'" municipality="Simpson's Bay" pageId="57" pageNumber="1784" specimenCount="1">
<emphasis id="74A71519FFD9FFC2DE01FEC8FCE2CD4E" box="[805,870,281,306]" italics="true" pageId="57" pageNumber="1784">kreffti</emphasis>
Inner/
<location id="430C9FD0FFD9FFC2DE9CFEC8FB05CD4E" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DE9CFEC8FB05CD4E" box="[952,1153,281,306]" municipality="Simpson's Bay" name="Western Islands'" pageId="57" pageNumber="1784">Western Islands</location>
; only a single lot from
<collectorName id="EB26ACDDFFD9FFC2DC87FEEDFD9DCD29" box="[419,537,316,341]" pageId="57" pageNumber="1784">Cape York</collectorName>
is known (
<collectionCode id="20C251CEFFD9FFC2DFB3FEEDFD3BCD29" box="[663,703,316,341]" country="Australia" name="Australian Museum" pageId="57" pageNumber="1784" type="Museum">AM</collectionCode>
C.425474 from
<collectingMunicipality id="A6085371FFD9FFC2DE5EFEEDFBA7CD29" box="[890,1059,316,341]" pageId="57" pageNumber="1784">Simpsons Bay</collectingMunicipality>
,
<collectorName id="EB26ACDDFFD9FFC2D90AFEEDFB04CD29" box="[1070,1152,316,341]" pageId="57" pageNumber="1784">Utingu</collectorName>
, coll. 1912)
</materialsCitation>
.
<materialsCitation id="F6BBC356FFD9FFC2DC02FE8FFED0CDE6" ID-GBIF-Occurrence="3978770348" location="Cox's" pageId="57" pageNumber="1784" specimenCount="1">
<location id="430C9FD0FFD9FFC2DC02FE8FFEE7CD0B" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DC02FE8FFEE7CD0B" box="[294,355,350,375]" name="Cox's" pageId="57" pageNumber="1784">Coxs</location>
description, dimensions and figure match the larger species and we accept this as
<taxonomicName id="81D3B288FFD9FFC2DC60FE50FED0CDE6" box="[324,340,385,410]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD9FFC2DC60FE50FED0CDE6" box="[324,340,385,410]" italics="true" pageId="57" pageNumber="1784">P</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="74A71519FFD9FFC2DC70FE50FE1BCDE6" box="[340,415,385,410]" italics="true" pageId="57" pageNumber="1784">. kreffti</emphasis>
.
<materialsCitation id="F6BBC356FFD9FFC2DC88FE50FD40CDC1" ID-GBIF-Occurrence="3978770310" collectionCode="AM, NHMUK" collectorName="The" pageId="57" pageNumber="1784" specimenCount="1" typeStatus="holotype">
<collectorName id="EB26ACDDFFD9FFC2DC88FE50FE52CDE6" box="[428,470,385,410]" pageId="57" pageNumber="1784">The</collectorName>
<typeStatus id="996877A9FFD9FFC2DCF9FE50FDC1CDE6" box="[477,581,385,410]" pageId="57" pageNumber="1784" type="holotype">holotype</typeStatus>
was recorded as lost by
<bibRefCitation id="2242B4FAFFD9FFC2DE45FE50FC71CDE6" author="Smith BJ" box="[865,1013,385,410]" pageId="57" pageNumber="1784" pagination="1 - 408" refId="ref31733" refString="Smith BJ 1992. Non-marine Mollusca. In: Houston W, editor. Zoological catalogue of Australia (Vol. 8). Canberra: Australian Government Publishing Service; pp. 1 - 408." type="book chapter" year="1992">Smith (1992)</bibRefCitation>
, and cannot be found in the collections of
<collectionCode id="20C251CEFFD9FFC2DF36FE75FDBCCDC1" box="[530,568,420,445]" country="Australia" name="Australian Museum" pageId="57" pageNumber="1784" type="Museum">AM</collectionCode>
or
<collectionCode id="20C251CEFFD9FFC2DF42FE75FD40CDC1" box="[614,708,420,445]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="57" pageNumber="1784" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
<materialsCitation id="F6BBC356FFD9FFC2DFF5FE75FB04CDC1" ID-GBIF-Occurrence="3978770302" box="[721,1152,420,445]" location="To" pageId="57" pageNumber="1784" specimenCount="1" typeStatus="holotype">
<location id="430C9FD0FFD9FFC2DFF5FE75FD69CDC1" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DFF5FE75FD69CDC1" box="[721,749,420,445]" name="To" pageId="57" pageNumber="1784">To</location>
clarify the taxonomic status of
<taxonomicName id="81D3B288FFD9FFC2D954FE75FB04CDC1" box="[1136,1152,420,445]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD9FFC2D954FE75FB04CDC1" box="[1136,1152,420,445]" italics="true" pageId="57" pageNumber="1784">P</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="74A71519FFD9FFC2D9A5FE75FB01CDC1" box="[1153,1157,420,445]" italics="true" pageId="57" pageNumber="1784">.</emphasis>
<materialsCitation id="F6BBC356FFD9FFC2DD84FE17FE87CE7E" ID-GBIF-Occurrence="3978770320" collectionCode="QM" location="Waiben Island" municipality="Art" pageId="57" pageNumber="1784" specimenCode="MO86003" specimenCount="1" typeStatus="neotype">
<emphasis id="74A71519FFD9FFC2DD84FE17FF65CDA3" box="[160,225,454,479]" italics="true" pageId="57" pageNumber="1784">kreffti</emphasis>
, we designate
<collectionCode id="20C251CEFFD9FFC2DCBFFE17FE47CDA3" box="[411,451,454,479]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="57" pageNumber="1784" type="Museum">QM</collectionCode>
<specimenCode id="16756170FFD9FFC2DCEAFE17FDB9CDA3" box="[462,573,454,479]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="57" pageNumber="1784" type="Museum">MO86003</specimenCode>
from
<location id="430C9FD0FFD9FFC2DFADFE17FCB4CDA3" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DFADFE17FCB4CDA3" box="[649,816,454,479]" municipality="Art" name="Waiben Island" pageId="57" pageNumber="1784">Waiben Island</location>
as a
<typeStatus id="996877A9FFD9FFC2DE50FE17FC50CDA3" box="[884,980,454,479]" pageId="57" pageNumber="1784" type="neotype">neotype</typeStatus>
in accordance with
<collectingMunicipality id="A6085371FFD9FFC2DDF9FE38FE87CE7E" box="[221,259,489,514]" pageId="57" pageNumber="1784">Art</collectingMunicipality>
</materialsCitation>
.
<materialsCitation id="F6BBC356FFD9FFC2DC2BFE38FBC3CE7E" ID-GBIF-Occurrence="3978770327" box="[271,1095,489,514]" location="International Code of Zoological Nomenclature" pageId="57" pageNumber="1784" specimenCount="1" typeStatus="neotype">
75 of the
<location id="430C9FD0FFD9FFC2DCA1FE38FC36CE7E" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DCA1FE38FC36CE7E" box="[389,946,489,514]" name="International Code of Zoological Nomenclature" pageId="57" pageNumber="1784">International Code of Zoological Nomenclature</location>
(
<bibRefCitation id="2242B4FAFFD9FFC2DEE6FE38FBB9CE7E" author="ICZN" box="[962,1085,489,514]" pageId="57" pageNumber="1784" refId="ref30554" refString="ICZN. 1999. The international code of zoological nomenclature. 4 th ed. London: The International Trust for Zoological Nomenclature." type="book" year="1999">ICZN 1999</bibRefCitation>
)
</materialsCitation>
. This specimen was dissected and sequenced and used as a basis for our taxonomic description.
<materialsCitation id="F6BBC356FFD9FFC2DC12FDFFFB04CE16" ID-GBIF-Occurrence="3978770321" location="It" pageId="57" pageNumber="1784" specimenCount="1" typeStatus="holotype">
<location id="430C9FD0FFD9FFC2DC12FDFFFEC2CE3B" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DC12FDFFFEC2CE3B" box="[310,326,558,583]" name="It" pageId="57" pageNumber="1784">It</location>
matches the dimensions of the
<typeStatus id="996877A9FFD9FFC2DFFEFDFFFCC1CE3B" box="[730,837,558,583]" pageId="57" pageNumber="1784" type="holotype">holotype</typeStatus>
, and among the available ethanol-preserved specimens, is geographically closest to the original type locality
</materialsCitation>
.
<materialsCitation id="F6BBC356FFD9FFC2DD84FDA5FE45CEF1" ID-GBIF-Occurrence="3978770307" box="[160,449,628,653]" collectorName="Art" location="In" pageId="57" pageNumber="1784" specimenCount="1" typeStatus="holotype">
<location id="430C9FD0FFD9FFC2DD84FDA5FF32CEF1" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DD84FDA5FF32CEF1" box="[160,182,628,653]" name="In" pageId="57" pageNumber="1784">In</location>
accordance with
<collectorName id="EB26ACDDFFD9FFC2DCBEFDA5FE45CEF1" box="[410,449,628,653]" pageId="57" pageNumber="1784">Art</collectorName>
</materialsCitation>
.
<materialsCitation id="F6BBC356FFD9FFC2DCF5FDA5FDFBCED3" ID-GBIF-Occurrence="3978770328" country="Iceland" county="Waiben" location="Code" municipality="Thursday" pageId="57" pageNumber="1784" specimenCount="1" typeStatus="holotype">
76 of the
<location id="430C9FD0FFD9FFC2DF76FDA5FD17CEF1" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DF76FDA5FD17CEF1" box="[594,659,628,653]" country="Iceland" county="Waiben" municipality="Thursday" name="Code" pageId="57" pageNumber="1784">Code</location>
,
<location id="430C9FD0FFD9FFC2DF86FDA5FCA4CEF1" LSID="urn:lsid:plazi:treatment:CE7A781DFFD7FFC2DCC6FEB5FDCDC833:430C9FD0FFD9FFC2DF86FDA5FCA4CEF1" box="[674,800,628,653]" country="Iceland" county="Waiben" municipality="Thursday" name="Green Hill" pageId="57" pageNumber="1784">Green Hill</location>
,
<collectingMunicipality id="A6085371FFD9FFC2DE14FDA5FC19CEF1" box="[816,925,628,653]" pageId="57" pageNumber="1784">Thursday</collectingMunicipality>
(=
<collectingCounty id="AF0DB187FFD9FFC2DEEBFDA5FBAFCEF1" box="[975,1067,628,653]" pageId="57" pageNumber="1784">Waiben</collectingCounty>
) Island becomes the type locality of this species
</materialsCitation>
.
</paragraph>
<paragraph id="466CC90BFFD9FFC2DDE4FD68FDCDC833" blockId="57.[160,1157,142,1103]" pageId="57" pageNumber="1784">
The distribution of
<taxonomicName id="81D3B288FFD9FFC2DC83FD68FDE3CEAE" box="[423,615,697,722]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="kreffti">
<emphasis id="74A71519FFD9FFC2DC83FD68FDE3CEAE" box="[423,615,697,722]" italics="true" pageId="57" pageNumber="1784">Pravonitor kreffti</emphasis>
</taxonomicName>
was previously recorded from Weipa, east to Portland Roads and north to Bamaga on Cape York, and also the Torres Strait Islands (
<bibRefCitation id="2242B4FAFFD9FFC2DC27FD2FFE6ACF6B" author="Stanisic J &amp; Shea M &amp; Potter D &amp; Griffiths O" box="[259,494,766,791]" pageId="57" pageNumber="1784" refId="ref32007" refString="Stanisic J, Shea M, Potter D, Griffiths O. 2010. Australian land snails. 1. A field guide to eastern Australian species. Riviere des Anguilles (Mauritius): Bioculture Press." type="book" year="2010">Stanisic et al. 2010</bibRefCitation>
). It is now limited to the very tip of Cape York, and the Near Western and Inner Islands of the Torres Strait (
<figureCitation id="DEE8D58EFFD9FFC2DE1CFCF0FC1DCF46" box="[824,921,801,826]" captionStart="Figure 4" captionStartId="27.[160,225,1613,1636]" captionTargetBox="[255,1060,142,1581]" captionTargetId="figure-3@27.[254,1061,142,1582]" captionTargetPageId="27" captionText="Figure 4. Distribution map. Grey symbols: Geminitor. Blue symbols: Pravonitor. Red symbols: Elatonitor. Biogeographic barriers are marked in black; A = Normanby Basin, B = Black Mountain Corridor, and C = Atherton Tableland." figureDoi="http://doi.org/10.5281/zenodo.7380691" httpUri="https://zenodo.org/record/7380691/files/figure.png" pageId="57" pageNumber="1784">Figure 4</figureCitation>
), including Waiben, Muralug, Mua and Badu Islands, and probably numerous additional islands including Ngurupai (= Horn), Maururra (= Wednesday), Gealug (= Friday) and Kiriri (= Hammond) Islands. Populations from islands in the Central and Top Western groups and the southern coast of New
<collectingCountry id="3EC4899BFFD9FFC2DF85FC7DFD70CFB9" box="[673,756,940,965]" name="Guinea-Bissau" pageId="57" pageNumber="1784">Guinea</collectingCountry>
are provisionally referred to
<taxonomicName id="81D3B288FFD9FFC2D954FC7DFF7FCF9B" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="annulus">
<emphasis id="74A71519FFD9FFC2D954FC7DFF7FCF9B" italics="true" pageId="57" pageNumber="1784">P. annulus</emphasis>
</taxonomicName>
, and populations from the Eastern Islands (including Mer Island) are placed in
<emphasis id="74A71519FFD9FFC2DD9BFC20FD99C876" box="[191,541,1009,1034]" italics="true" pageId="57" pageNumber="1784">
<taxonomicName id="81D3B288FFD9FFC2DD9BFC20FED4C876" box="[191,336,1009,1034]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="insularum">P. insularum</taxonomicName>
.
<taxonomicName id="81D3B288FFD9FFC2DC79FC20FD99C876" box="[349,541,1009,1034]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="kreffti">Pravonitor kreffti</taxonomicName>
</emphasis>
can be distinguished from
<taxonomicName id="81D3B288FFD9FFC2DE42FC20FC59C876" box="[870,989,1009,1034]" class="Gastropoda" family="Helicarionidae" genus="Pravonitor" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="1784" phylum="Mollusca" rank="species" species="annulus">
<emphasis id="74A71519FFD9FFC2DE42FC20FC59C876" box="[870,989,1009,1034]" italics="true" pageId="57" pageNumber="1784">P. annulus</emphasis>
</taxonomicName>
by its slightly larger size, higher spire and rounded rather than subangulate whorl profile, and by the lack of a supraperipheral band.
</paragraph>
</subSubSection>
</treatment>
</document>