525 lines
54 KiB
XML
525 lines
54 KiB
XML
<document ID-DOI="10.1080/00222933.2022.2152750" ID-GBIF-Dataset="53db727e-224e-4dbc-82fa-67760d26f0e3" ID-ISSN="1464-5262" ID-Zenodo-Dep="7560931" checkinTime="1674200384013" checkinUser="plazi" docAuthor="Páll-Gergely, Barna & White, Tom S." docDate="2023" docId="2050F30FBA4BFFC5FE89254EFCDCF92A" docLanguage="en" docName="JNATHIST.56.45-48.2011-2029.pdf" docOrigin="Journal of Natural History 56 (45 - 48)" docSource="http://dx.doi.org/10.1080/00222933.2022.2152750" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Bensonella Pilsbry and Vanatta 1900" docType="treatment" docVersion="3" lastPageNumber="2016" masterDocId="DC698B77BA48FFC0FFF9214FFF93FFB7" masterDocTitle="Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)" masterLastPageNumber="2029" masterPageNumber="2011" pageNumber="2014" updateTime="1674475451874" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Páll-Gergely, Barna</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>White, Tom S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>56</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>45 - 48</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>2011</mods:start>
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<mods:end>2029</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1080/00222933.2022.2152750</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/00222933.2022.2152750</mods:identifier>
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<mods:identifier type="GBIF-Dataset">53db727e-224e-4dbc-82fa-67760d26f0e3</mods:identifier>
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<mods:identifier type="ISSN">1464-5262</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7560931</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7560933" ID-Zenodo-Dep="7560933" LSID="urn:lsid:plazi:treatment:2050F30FBA4BFFC5FE89254EFCDCF92A" httpUri="http://treatment.plazi.org/id/2050F30FBA4BFFC5FE89254EFCDCF92A" lastPageId="5" lastPageNumber="2016" pageId="3" pageNumber="2014">
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<subSubSection box="[368,948,1025,1052]" pageId="3" pageNumber="2014" type="nomenclature">
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||
<paragraph blockId="3.[368,948,1025,1052]" box="[368,948,1025,1052]" pageId="3" pageNumber="2014">
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||
<heading bold="true" box="[368,948,1025,1052]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="2014" reason="3">
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||
<emphasis bold="true" box="[368,948,1025,1052]" pageId="3" pageNumber="2014">
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Genus
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<taxonomicName ID-CoL="7NSZW" authority="Pilsbry and Vanatta, 1900" authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[458,948,1025,1052]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis bold="true" box="[458,598,1025,1052]" italics="true" pageId="3" pageNumber="2014">Bensonella</emphasis>
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<bibRefCitation author="Pilsbry HA & Vanatta EG" box="[606,948,1025,1052]" pageId="3" pageNumber="2014" pagination="582 - 611" refId="ref7771" refString="Pilsbry HA, Vanatta EG. 1900. A partial revision of the Pupae of the United States. Proc Acad Nat Sci Phila. 52: 582 - 611." type="journal article" year="1900">Pilsbry and Vanatta, 1900</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="2014" type="reference_group">
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<paragraph blockId="3.[192,962,1079,1173]" box="[192,768,1079,1104]" pageId="3" pageNumber="2014">
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<treatmentCitationGroup box="[192,768,1079,1104]" pageId="3" pageNumber="2014">
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<treatmentCitation author="Pilsbry HA & Vanatta EG" box="[192,764,1079,1104]" page="591" pageId="3" pageNumber="2014" year="1900">
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<taxonomicName authority="Pilsbry and Vanatta 1900: 591" authorityName="Pilsbry and Vanatta" authorityPageNumber="591" authorityYear="1900" box="[192,764,1079,1104]" class="Gastropoda" family="Gastrocoptidae" genus="Bifidaria" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="subGenus" subGenus="Bensonella">
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<emphasis box="[192,282,1079,1104]" italics="true" pageId="3" pageNumber="2014">Bifidaria</emphasis>
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(
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<emphasis box="[297,413,1079,1104]" italics="true" pageId="3" pageNumber="2014">Bensonella</emphasis>
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)
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<bibRefCitation author="Pilsbry HA & Vanatta EG" box="[428,764,1079,1104]" pageId="3" pageNumber="2014" pagination="582 - 611" refId="ref7771" refString="Pilsbry HA, Vanatta EG. 1900. A partial revision of the Pupae of the United States. Proc Acad Nat Sci Phila. 52: 582 - 611." type="journal article" year="1900">Pilsbry and Vanatta 1900: 591</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="3.[192,962,1079,1173]" box="[192,901,1114,1139]" pageId="3" pageNumber="2014">
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<taxonomicName box="[192,421,1114,1139]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="subGenus" subGenus="Bensonella">
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<emphasis box="[192,282,1114,1139]" italics="true" pageId="3" pageNumber="2014">Boysidia</emphasis>
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(
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<emphasis box="[297,413,1114,1139]" italics="true" pageId="3" pageNumber="2014">Bensonella</emphasis>
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)
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</taxonomicName>
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– Pilsbry 1917 (in
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<bibRefCitation author="Pilsbry HA" box="[628,766,1114,1139]" pageId="3" pageNumber="2014" refId="ref7592" refString="Pilsbry HA. 1916 - 1918. Manual of conchology, structural and systematic, with illustrations of the species. Ser. 2, Pulmonata. Vol. 24: Pupillidae (Gastrocoptinae). pp 1 - 380, pls 1 - 49. Philadelphia, published by the Conchological Section, Academy of Natural Sciences. [pp. 1 - 112, Pl. 1 - 13, 18 Dec 1916; pp. 113 - 176, pl. 14 - 29, 18 July 1917; pp. 177 - 256, pl. 30 - 38, 09 Nov 1917; pp. 257 - 380, pl. 39 - 49, 24 April 1918]." type="book" year="1916">Pilsbry 1916</bibRefCitation>
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–1918): 198.
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</paragraph>
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</subSubSection>
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<subSubSection box="[192,962,1148,1173]" pageId="3" pageNumber="2014" type="description">
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<paragraph blockId="3.[192,962,1079,1173]" box="[192,962,1148,1173]" pageId="3" pageNumber="2014">
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<taxonomicName authority="Pilsbry, 1917" authorityName="Pilsbry" authorityYear="1917" box="[192,593,1148,1173]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="subGenus" subGenus="Paraboysidia">
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<emphasis box="[192,282,1148,1173]" italics="true" pageId="3" pageNumber="2014">Boysidia</emphasis>
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(
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<emphasis box="[297,437,1148,1173]" italics="true" pageId="3" pageNumber="2014">Paraboysidia</emphasis>
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) Pilsbry, 1917
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</taxonomicName>
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(in
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<bibRefCitation author="Pilsbry HA" box="[636,774,1148,1173]" pageId="3" pageNumber="2014" refId="ref7592" refString="Pilsbry HA. 1916 - 1918. Manual of conchology, structural and systematic, with illustrations of the species. Ser. 2, Pulmonata. Vol. 24: Pupillidae (Gastrocoptinae). pp 1 - 380, pls 1 - 49. Philadelphia, published by the Conchological Section, Academy of Natural Sciences. [pp. 1 - 112, Pl. 1 - 13, 18 Dec 1916; pp. 113 - 176, pl. 14 - 29, 18 July 1917; pp. 177 - 256, pl. 30 - 38, 09 Nov 1917; pp. 257 - 380, pl. 39 - 49, 24 April 1918]." type="book" year="1916">Pilsbry 1916</bibRefCitation>
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–1918): 174, 201.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="2014" type="type_taxon">
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<paragraph blockId="3.[160,971,1217,1277]" box="[160,307,1217,1243]" pageId="3" pageNumber="2014">
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<emphasis bold="true" box="[160,307,1217,1243]" italics="true" pageId="3" pageNumber="2014">
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<typeStatus box="[160,216,1217,1243]" pageId="3" pageNumber="2014">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph blockId="3.[160,971,1217,1277]" box="[160,971,1252,1277]" pageId="3" pageNumber="2014">
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<taxonomicName authority="Benson, 1849" authorityName="Benson" authorityYear="1849" box="[160,475,1252,1277]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="species" species="plicidens">
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<emphasis box="[160,215,1252,1277]" italics="true" pageId="3" pageNumber="2014">Pupa</emphasis>
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<emphasis box="[223,317,1252,1277]" italics="true" pageId="3" pageNumber="2014">plicidens</emphasis>
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<bibRefCitation author="Benson WH" box="[324,475,1252,1277]" pageId="3" pageNumber="2014" pagination="125 - 128" refId="ref6320" refString="Benson WH. 1849. Descriptions of four new Asiatic species of the genus Pupa of Draparnaud. Ann Mag Nat Hist. 4 (20): 125 - 128. doi: 10.1080 / 03745486009496158." type="journal article" year="1849">Benson, 1849</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation box="[489,551,1252,1277]" captionStart="Figure 3" captionStartId="4.[159,224,1608,1631]" captionTargetBox="[212,1099,138,1575]" captionTargetId="figure-3@4.[211,1101,137,1577]" captionTargetPageId="4" captionText="Figure 3. Bensonella plicidens (Benson, 1849) and Bensonella paviei (Bavay and Dautzenberg, 1912). a–e: Lectotype of Bensonella plicidens (Benson, 1849), UMZC I.103325.A; f–k: Syntype of Bensonella paviei (Bavay and Dautzenberg, 1912), MNHN-IM-2000-35,158. Scale refers to a–e; f–k are not to scale (could not measure properly with hand callipers, ca. 2 mm). Photos: B. Páll-Gergely (f–k); G. Kevin Webb, NHM Imaging Unit, © Trustees of the Natural History Museum, London (a–e)." figureDoi="http://doi.org/10.5281/zenodo.7560939" httpUri="https://zenodo.org/record/7560939/files/figure.png" pageId="3" pageNumber="2014">Fig. 3</figureCitation>
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(a–e) herein), by original designation.
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</paragraph>
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</subSubSection>
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||
<subSubSection lastPageId="5" lastPageNumber="2016" pageId="3" pageNumber="2014" type="discussion">
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<paragraph blockId="3.[160,1156,1321,1728]" box="[160,262,1321,1347]" pageId="3" pageNumber="2014">
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<emphasis bold="true" box="[160,262,1321,1347]" italics="true" pageId="3" pageNumber="2014">Remarks</emphasis>
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</paragraph>
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<paragraph blockId="3.[160,1156,1321,1728]" pageId="3" pageNumber="2014">
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According to Pilsbry̾s (1917) original description of
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<taxonomicName authorityName="Pilsbry" authorityYear="1917" box="[740,880,1357,1382]" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[740,880,1357,1382]" italics="true" pageId="3" pageNumber="2014">Paraboysidia</emphasis>
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</taxonomicName>
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, this genus (at that time a subgenus of
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<taxonomicName authorityName="Ancey" authorityYear="1881" box="[327,417,1391,1416]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[327,417,1391,1416]" italics="true" pageId="3" pageNumber="2014">Boysidia</emphasis>
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</taxonomicName>
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) differs from
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<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[569,685,1391,1416]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[569,685,1391,1416]" italics="true" pageId="3" pageNumber="2014">Bensonella</emphasis>
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</taxonomicName>
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by having normal (not hooked) apertural barriers. However, this claim is incorrect because the
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<typeStatus box="[781,831,1426,1451]" pageId="3" pageNumber="2014">type</typeStatus>
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species of
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<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[965,1081,1426,1451]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[965,1081,1426,1451]" italics="true" pageId="3" pageNumber="2014">Bensonella</emphasis>
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</taxonomicName>
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(
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<emphasis italics="true" pageId="3" pageNumber="2014">
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Pupa
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<taxonomicName authorityName="Benson" authorityYear="1849" box="[160,254,1461,1486]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="species" species="plicidens">plicidens</taxonomicName>
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</emphasis>
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) has normal barriers. The
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<typeStatus box="[545,595,1461,1486]" pageId="3" pageNumber="2014">type</typeStatus>
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species of
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<taxonomicName authorityName="Pilsbry" authorityYear="1917" box="[719,859,1461,1486]" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[719,859,1461,1486]" italics="true" pageId="3" pageNumber="2014">Paraboysidia</emphasis>
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</taxonomicName>
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(
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<taxonomicName authority="Bavay and Dautzenberg, 1912" authorityName="Bavay and Dautzenberg" authorityYear="1912" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="species" species="paviei">
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<emphasis box="[872,1033,1461,1486]" italics="true" pageId="3" pageNumber="2014">Boysidia paviei</emphasis>
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<bibRefCitation author="Bavay A & Dautzenberg P" pageId="3" pageNumber="2014" pagination="1 - 54" refId="ref6290" refString="Bavay A, Dautzenberg P. 1912. Description de coquilles nouvelles de l ̾ Indo-Chine. Journal de Conchyliologie. 60 (1): 1 - 54." type="journal article" year="1912">Bavay and Dautzenberg, 1912</bibRefCitation>
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</taxonomicName>
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, fig. 3(f–k)) is very similar to that of
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<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[819,935,1495,1520]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[819,935,1495,1520]" italics="true" pageId="3" pageNumber="2014">Bensonella</emphasis>
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</taxonomicName>
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(
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<emphasis box="[954,1115,1495,1520]" italics="true" pageId="3" pageNumber="2014">
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Pupa
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<taxonomicName authorityName="Benson" authorityYear="1849" box="[1021,1115,1495,1520]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="species" species="plicidens">plicidens</taxonomicName>
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</emphasis>
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) in general shell shell and aperture shape, and in the arrangement of the apertural barriers. The main difference is the presence of some additional ̍supplementary̾ apertural folds (i.e. tiny plicae between main plicae) in
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<taxonomicName authorityName="Benson" authorityYear="1849" box="[618,741,1599,1624]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="species" species="plicidens">
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<emphasis box="[618,741,1599,1624]" italics="true" pageId="3" pageNumber="2014">P. plicidens</emphasis>
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</taxonomicName>
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, which does not seem to justify the distinction of two genera. Therefore, herein
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<taxonomicName authorityName="Pilsbry" authorityYear="1917" box="[649,789,1634,1659]" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[649,789,1634,1659]" italics="true" pageId="3" pageNumber="2014">Paraboysidia</emphasis>
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</taxonomicName>
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is treated as a junior synonym of
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<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[160,276,1669,1694]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[160,276,1669,1694]" italics="true" pageId="3" pageNumber="2014">Bensonella</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Gittenberger E & Gyeltshen C & Leda P & Sherub S" box="[288,566,1669,1694]" pageId="3" pageNumber="2014" pagination="69 - 90" refId="ref6713" refString="Gittenberger E, Gyeltshen C, Leda P, Sherub S. 2021. The superfamilies Pupilloidea and Enoidea (Gastropoda, Eupulmonata) in Bhutan. Folia Malacol. 29 (2): 69 - 90. doi: 10.12657 / folmal. 029.009." type="journal article" year="2021">Gittenberger et al. (2021)</bibRefCitation>
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also treated
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<taxonomicName authorityName="Pilsbry" authorityYear="1917" box="[709,849,1669,1694]" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[709,849,1669,1694]" italics="true" pageId="3" pageNumber="2014">Paraboysidia</emphasis>
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</taxonomicName>
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as a synonym of
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<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[1039,1155,1669,1694]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="2014" phylum="Mollusca" rank="genus">
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<emphasis box="[1039,1155,1669,1694]" italics="true" pageId="3" pageNumber="2014">Bensonella</emphasis>
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</taxonomicName>
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but did not explain their decision to do so.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7560939" ID-Zenodo-Dep="7560939" httpUri="https://zenodo.org/record/7560939/files/figure.png" pageId="4" pageNumber="2015" startId="4.[159,224,1608,1631]" targetBox="[212,1099,138,1575]" targetPageId="4">
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||
<paragraph blockId="4.[159,1154,1608,1749]" pageId="4" pageNumber="2015">
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<emphasis bold="true" box="[159,250,1608,1631]" pageId="4" pageNumber="2015">Figure 3.</emphasis>
|
||
<taxonomicName authority="(Benson, 1849)" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[259,607,1608,1631]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="2015" phylum="Mollusca" rank="species" species="plicidens">
|
||
<emphasis box="[259,451,1608,1631]" italics="true" pageId="4" pageNumber="2015">Bensonella plicidens</emphasis>
|
||
(
|
||
<bibRefCitation author="Benson WH" box="[465,600,1608,1631]" pageId="4" pageNumber="2015" pagination="125 - 128" refId="ref6320" refString="Benson WH. 1849. Descriptions of four new Asiatic species of the genus Pupa of Draparnaud. Ann Mag Nat Hist. 4 (20): 125 - 128. doi: 10.1080 / 03745486009496158." type="journal article" year="1849">Benson, 1849</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="(Bavay and Dautzenberg, 1912)" baseAuthorityName="Bavay and Dautzenberg" baseAuthorityYear="1912" box="[660,1150,1608,1631]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="2015" phylum="Mollusca" rank="species" species="paviei">
|
||
<emphasis box="[660,827,1608,1631]" italics="true" pageId="4" pageNumber="2015">Bensonella paviei</emphasis>
|
||
(
|
||
<bibRefCitation author="Bavay A & Dautzenberg P" box="[840,1142,1608,1631]" pageId="4" pageNumber="2015" pagination="1 - 54" refId="ref6290" refString="Bavay A, Dautzenberg P. 1912. Description de coquilles nouvelles de l ̾ Indo-Chine. Journal de Conchyliologie. 60 (1): 1 - 54." type="journal article" year="1912">Bavay and Dautzenberg, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. a–e: Lectotype of
|
||
<taxonomicName authority="(Benson, 1849)" baseAuthorityName="Benson" baseAuthorityYear="1849" box="[342,694,1637,1660]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="2015" phylum="Mollusca" rank="species" species="plicidens">
|
||
<emphasis box="[342,535,1637,1660]" italics="true" pageId="4" pageNumber="2015">Bensonella plicidens</emphasis>
|
||
(
|
||
<bibRefCitation author="Benson WH" box="[550,685,1637,1660]" pageId="4" pageNumber="2015" pagination="125 - 128" refId="ref6320" refString="Benson WH. 1849. Descriptions of four new Asiatic species of the genus Pupa of Draparnaud. Ann Mag Nat Hist. 4 (20): 125 - 128. doi: 10.1080 / 03745486009496158." type="journal article" year="1849">Benson, 1849</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, UMZC I.103325.A; f–k: Syntype of
|
||
<taxonomicName authority="(Bavay and Dautzenberg, 1912)" baseAuthorityName="Bavay and Dautzenberg" baseAuthorityYear="1912" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="2015" phylum="Mollusca" rank="species" species="paviei">
|
||
<emphasis italics="true" pageId="4" pageNumber="2015">Bensonella paviei</emphasis>
|
||
(
|
||
<bibRefCitation author="Bavay A & Dautzenberg P" box="[226,521,1667,1690]" pageId="4" pageNumber="2015" pagination="1 - 54" refId="ref6290" refString="Bavay A, Dautzenberg P. 1912. Description de coquilles nouvelles de l ̾ Indo-Chine. Journal de Conchyliologie. 60 (1): 1 - 54." type="journal article" year="1912">Bavay and Dautzenberg, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, MNHN-IM-2000-35,158. Scale refers to a–e; f–k are not to scale (could not measure properly with hand callipers, ca. 2 mm). Photos: B. Páll-Gergely (f–k); G. Kevin Webb, NHM Imaging Unit, © Trustees of the Natural History Museum, London (a–e).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="5.[160,1157,142,1693]" pageId="5" pageNumber="2016">
|
||
<taxonomicName box="[192,402,142,167]" pageId="5" pageNumber="2016">
|
||
<emphasis box="[192,311,142,167]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with hooked and normal barriers in the Himalaya (i.e.
|
||
<taxonomicName authority="Páll-Gergely & White, 2023" authorityName="Páll-Gergely & White" authorityYear="2023" box="[1027,1116,142,167]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
|
||
<emphasis box="[1027,1116,142,167]" italics="true" pageId="5" pageNumber="2016">B. hooki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="5" pageNumber="2016" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName baseAuthorityName="Benson" baseAuthorityYear="1849" box="[280,411,177,202]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="plicidens">
|
||
<emphasis box="[280,411,177,202]" italics="true" pageId="5" pageNumber="2016">B. plicidens</emphasis>
|
||
</taxonomicName>
|
||
) are very similar to each other in all other shell characters. Moreover, hooked and normal apertural barriers are characteristic of otherwise extremely similar and close-occurring species in the genera
|
||
<taxonomicName authority="F. G. Thompson and Upatham 1997" authorityName="F. G. Thompson and Upatham" authorityYear="1997" class="Gastropoda" family="Gastrocoptidae" genus="Acinolaemus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[786,927,246,271]" italics="true" pageId="5" pageNumber="2016">Acinolaemus</emphasis>
|
||
F.G.
|
||
<bibRefCitation author="Thompson FG & Upatham S" pageId="5" pageNumber="2016" pagination="221 - 245" refId="ref7890" refString="Thompson FG, Upatham S. 1997. Vertiginid land snails from Thailand (Gastropoda, Pulmonata, Pupilloidea). Bull Fla Mus Nat Hist. 39 (7): 221 - 245." type="journal article" year="1997">Thompson and Upatham 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName authority="Pilsbry, 1917" authorityName="Pilsbry" authorityYear="1917" box="[349,620,281,306]" class="Gastropoda" family="Gastrocoptidae" genus="Anauchen" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[349,460,281,306]" italics="true" pageId="5" pageNumber="2016">Anauchen</emphasis>
|
||
Pilsbry, 1917
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName authority="Benson, 1856" authorityName="Benson" authorityYear="1856" box="[690,1013,281,306]" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[690,845,281,306]" italics="true" pageId="5" pageNumber="2016">Hypselostoma</emphasis>
|
||
<bibRefCitation author="Benson WH" box="[856,1013,281,306]" pageId="5" pageNumber="2016" pagination="342 - 343" refId="ref6358" refString="Benson WH. 1856. Remarks on the genera Tanystoma, Nematura, and Anaulus. Ann Mag Nat Hist. 17 (2): 342 - 343. doi: 10.1080 / 00222935608697520." type="journal article" year="1856">Benson, 1856</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Vermeulen JJ & Phung LC & Truong QT" pageId="5" pageNumber="2016" pagination="81 - 92" refId="ref8118" refString="Vermeulen JJ, Phung LC, Truong QT. 2007. New species of terrestrial molluscs (Caenogastropoda, Pupinidae & Pulmonata, Vertiginidae) of the Hon Chong - Ha Tien limestone hills, Southern Vietnam. Basteria. 71: 81 - 92." type="journal article" year="2007">Vermeulen et al. 2007</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Vermeulen JJ & Luu HT & Theary K & Anker K" box="[308,364,316,341]" pageId="5" pageNumber="2016" pagination="167 - 177" refId="ref8040" refString="Vermeulen JJ, Luu HT, Theary K, Anker K. 2019. Land snail fauna of the Mekong Delta Limestone Hills (Cambodia, Vietnam): notharinia Vermeulen, Phung et Truong, 2007, and a note on Plectostoma A. Adams, 1865 (Mollusca: Gastropoda: Caenogastropoda: Diplommatinidae). Folia Malacol. 27 (3): 167 - 177. doi: 10.12657 / folmal. 027.015." type="journal article" year="2019">2019</bibRefCitation>
|
||
). This suggests that the hooked or normal apertural barriers are subjects of rapid, probably adaptive (see
|
||
<bibRefCitation author="Wada S & Chiba S" box="[636,888,350,375]" pageId="5" pageNumber="2016" pagination="54123" refId="ref8164" refString="Wada S, Chiba S. 2013. The dual protection of a micro land snail against a micro predatory snail. PLoS ONE. 8 (1): e 54123. doi: 10.1371 / journal. pone. 0054123." type="journal article" year="2013">
|
||
Wada and
|
||
<collectingRegion box="[759,825,350,375]" country="Japan" name="Chiba" pageId="5" pageNumber="2016">Chiba</collectingRegion>
|
||
2013
|
||
</bibRefCitation>
|
||
) evolutionary changes. Since it is probable that the hooked apertural barriers have been evolved multiple times in different groups within the
|
||
<taxonomicName authorityName="Zilch" authorityYear="1959" box="[511,734,420,445]" class="Gastropoda" family="Hypselostomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="family">Hypselostomatidae</taxonomicName>
|
||
, it is unlikely that species with apertural barriers would form a monophyletic unit. Therefore, it is not recommended to use this character state to distinguish between genera. We thus classify species with both hooked (
|
||
<taxonomicName authority="Páll-Gergely & White, 2023" authorityName="Páll-Gergely & White" authorityYear="2023" box="[261,349,524,549]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="hooki" status="sp. nov.">
|
||
<emphasis box="[261,349,524,549]" italics="true" pageId="5" pageNumber="2016">B. hooki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[357,447,524,549]" pageId="5" pageNumber="2016" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authority="Hwang, 2014" authorityName="Hwang" authorityYear="2014" box="[457,760,524,549]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="lakainguta">
|
||
<emphasis box="[457,603,524,549]" italics="true" pageId="5" pageNumber="2016">B. lakainguta</emphasis>
|
||
<bibRefCitation author="Hwang C-C" box="[610,760,524,549]" pageId="5" pageNumber="2016" pagination="15 - 26" refId="ref6907" refString="Hwang C-C. 2014. A new subspecies of land snail Bensonella plicidens lakainguta (Gastropoda, Vertiginidae) from southern Taiwan. Bull Malacol Taiwan 貝類學報. 37: 15 - 26." type="journal article" year="2014">Hwang, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
) and normal apertural barriers (i.e.
|
||
<taxonomicName baseAuthorityName="Benson" baseAuthorityYear="1849" box="[160,284,558,583]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="plicidens">
|
||
<emphasis box="[160,284,558,583]" italics="true" pageId="5" pageNumber="2016">B. plicidens</emphasis>
|
||
</taxonomicName>
|
||
) in
|
||
<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[328,447,558,583]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[328,447,558,583]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[160,1157,142,1693]" pageId="5" pageNumber="2016">
|
||
In the last three decades, several
|
||
<taxonomicName box="[574,804,593,618]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="subGenus" subGenus="Bensonella">
|
||
<emphasis box="[574,664,593,618]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[680,796,593,618]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species from
|
||
<collectingCountry box="[964,1028,593,618]" name="China" pageId="5" pageNumber="2016">China</collectingCountry>
|
||
have been newly described or transferred into the (sub)genus:
|
||
<taxonomicName authorityName="D. -N. Chen, Y. -H. Liu & W. -X. Wu" authorityYear="1995" box="[743,964,628,653]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="dilamellaris">
|
||
<emphasis box="[743,964,628,653]" italics="true" pageId="5" pageNumber="2016">Boysidia dilamellaris</emphasis>
|
||
</taxonomicName>
|
||
D.-N. Chen, Y.-H. Liu and W.-X. Wu, 1995 (transferred to
|
||
<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[653,773,662,687]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[653,773,662,687]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Chen D-N & Wu M & Zhang G-Q" box="[829,1035,662,687]" pageId="5" pageNumber="2016" pagination="124 - 135" refId="ref6572" refString="Chen D-N, Wu M, Zhang G-Q. 1999. Studies on the genus Boysidia Ancey from China with descriptions of six new species (Pulmonata: Stylommatophora: Pupillidae). Acta Zootaxonomica Sinica. 24 (2): 124 - 135." type="journal article" year="1999">Chen et al. 1999</bibRefCitation>
|
||
);
|
||
<taxonomicName authority="M. Tian, B. Fao" authorityName="M. Tian, B. Fao" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="jinpingensis" subGenus="Bensonella">
|
||
<emphasis box="[1063,1155,662,687]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[168,284,697,722]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis box="[301,428,697,722]" italics="true" pageId="5" pageNumber="2016">jinpingensis</emphasis>
|
||
M. Tian, B. Fao
|
||
</taxonomicName>
|
||
and Y.-X. Chen, 2015;
|
||
<taxonomicName authorityName="Z. -L. Zhang, W. -H. Zhang & T. -C. Luo" authorityYear="2011" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="nanjiangensis" subGenus="Bensonella">
|
||
<emphasis box="[865,955,697,722]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[971,1087,697,722]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis italics="true" pageId="5" pageNumber="2016">nanjiangensis</emphasis>
|
||
</taxonomicName>
|
||
Z.-L. Zhang, W.-H. Zhang and T.-C. Luo, 2011;
|
||
<taxonomicName authority="F. Fang, J. Wang and Y. Chen, 2015" authorityName="F. Fang, J. Wang and Y. Chen" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="qingliangfengensis" subGenus="Bensonella">
|
||
<emphasis box="[804,894,732,757]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[911,1027,732,757]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis italics="true" pageId="5" pageNumber="2016">qingliangfengensis</emphasis>
|
||
F. Fang, J. Wang and Y. Chen, 2015
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName authorityName="D. -N. Chen, M. Wu & G. -Q. Zhang" authorityYear="1999" box="[676,1021,766,791]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="shilinensis" subGenus="Bensonella">
|
||
<emphasis box="[676,766,766,791]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[781,897,766,791]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis box="[912,1021,766,791]" italics="true" pageId="5" pageNumber="2016">shilinensis</emphasis>
|
||
</taxonomicName>
|
||
D.-N. Chen, M. Wu and G.-Q. Zhang, 1999;
|
||
<taxonomicName authorityName="T. -C. Luo, D. -N. Chen & G. -Q. Zhang" authorityYear="2000" box="[498,921,801,826]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="tianxingqiaoensis" subGenus="Bensonella">
|
||
<emphasis box="[498,588,801,826]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[602,718,801,826]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis box="[732,921,801,826]" italics="true" pageId="5" pageNumber="2016">tianxingqiaoensis</emphasis>
|
||
</taxonomicName>
|
||
T.-C. Luo, D.-N. Chen and G.-Q. Zhang, 2000 (
|
||
<bibRefCitation author="Luo T-C & Chen D-N & Zhang G-Q" box="[432,604,836,861]" pageId="5" pageNumber="2016" pagination="147 - 151" refId="ref7061" refString="Luo T-C, Chen D-N, Zhang G-Q. 2000. Two new species of Pupillidae From China (Pulmonata: Stylommatophora: Pupillidae). Acta Zootaxonomica Sinica. 25 (2): 147 - 151." type="journal article" year="2000">Luo et al. 2000</bibRefCitation>
|
||
);
|
||
<taxonomicName authorityName="D.-N. Chen & W.-H. Zhang" authorityYear="2002" box="[626,1024,836,861]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="tongguanensis" subGenus="Bensonella">
|
||
<emphasis box="[626,716,836,861]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[732,848,836,861]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis box="[865,1024,836,861]" italics="true" pageId="5" pageNumber="2016">tongguanensis</emphasis>
|
||
</taxonomicName>
|
||
D.-N. Chen and W.-H. Zhang, 2002;
|
||
<taxonomicName authorityName="Y. -H. Guo, W. -C. Zhou & T. -C. Luo" authorityYear="2006" box="[431,799,870,895]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="xingyinensis" subGenus="Bensonella">
|
||
<emphasis box="[431,521,870,895]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[536,652,870,895]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis box="[668,799,870,895]" italics="true" pageId="5" pageNumber="2016">xingyinensis</emphasis>
|
||
</taxonomicName>
|
||
Y.-H. Guo, W.-C. Zhou and T.-C. Luo, 2006 (
|
||
<bibRefCitation author="Guo Y-H & Zhou W-C & Luo T-C" box="[284,458,905,930]" pageId="5" pageNumber="2016" pagination="541 - 543" refId="ref6802" refString="Guo Y-H, Zhou W-C, Luo T-C. 2006. One new species of the genus Boysidia (Pulmonata: Stylommatophora: Pupillidae). Acta Zootaxonomica Sinica. 31 (3): 541 - 543." type="journal article" year="2006">Guo et al. 2006</bibRefCitation>
|
||
); and
|
||
<taxonomicName authorityName="W. -H. Zhang, T. -C. Luo & W. -C. Zhou" authorityYear="2010" box="[530,896,905,930]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="xiuwenensis" subGenus="Bensonella">
|
||
<emphasis box="[530,620,905,930]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[635,751,905,930]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis box="[767,896,905,930]" italics="true" pageId="5" pageNumber="2016">xiuwenensis</emphasis>
|
||
</taxonomicName>
|
||
W.-H. Zhang, T.-C. Luo and W.-C. Zhou, 2010.
|
||
</paragraph>
|
||
<paragraph blockId="5.[160,1157,142,1693]" pageId="5" pageNumber="2016">
|
||
Of these,
|
||
<emphasis box="[321,1096,974,999]" italics="true" pageId="5" pageNumber="2016">
|
||
<taxonomicName authorityName="D. -N. Chen, Y. -H. Liu & W. -X. Wu" authorityYear="1995" box="[321,488,974,999]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="dilamellaris">B. dilamellaris</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Z. -L. Zhang, W. -H. Zhang & T. -C. Luo" authorityYear="2011" box="[509,702,974,999]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="nanjiangensis">B. nanjiangensis</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="D. -N. Chen, M. Wu & G. -Q. Zhang" authorityYear="1999" box="[722,874,974,999]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="shilinensis">B. shilinensis</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="D.-N. Chen & W.-H. Zhang" authorityYear="2002" box="[895,1096,974,999]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="tongguanensis">B. tongguanensis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authorityName="W. -H. Zhang, T. -C. Luo & W. -C. Zhou" authorityYear="2010" box="[160,317,1009,1034]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="xiuwenensis">
|
||
<emphasis box="[160,317,1009,1034]" italics="true" pageId="5" pageNumber="2016">B. xiuwenensis</emphasis>
|
||
</taxonomicName>
|
||
were classified as within
|
||
<taxonomicName authority="Ancey, 1881" authorityName="Ancey" authorityYear="1881" box="[609,845,1009,1034]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[609,699,1009,1034]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
<bibRefCitation author="Ancey CF" box="[707,845,1009,1034]" pageId="5" pageNumber="2016" pagination="373 - 374" refId="ref6240" refString="Ancey CF. 1881. Description de mollusques terrestres nouveaux. Le Naturaliste. 3 (47): 373 - 374." type="journal article" year="1881">Ancey, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
(without subgeneric classification) by
|
||
<bibRefCitation author="Zhang W-H & Chen D-N & Zhou W-C" box="[290,498,1044,1069]" pageId="5" pageNumber="2016" pagination="570 - 575" refId="ref8207" refString="Zhang W-H, Chen D-N, Zhou W-C. 2014. Two new species of the genus Boysidia from China, with a preliminary discussion of its geography (Pulmonata: Stylommatophora: Pupillidae). Zool Syst. 39 (4): 570 - 575." type="journal article" year="2014">Zhang et al. (2014)</bibRefCitation>
|
||
. We agree with this classification because these species are similar in shell shape and apertural dentition (i.e. fused angular and palatal lamella) to the
|
||
<typeStatus box="[160,210,1113,1138]" pageId="5" pageNumber="2016">type</typeStatus>
|
||
species,
|
||
<emphasis box="[313,460,1113,1138]" italics="true" pageId="5" pageNumber="2016">Pupa dorsata</emphasis>
|
||
<bibRefCitation author="Ancey CF" box="[467,605,1113,1138]" pageId="5" pageNumber="2016" pagination="373 - 374" refId="ref6240" refString="Ancey CF. 1881. Description de mollusques terrestres nouveaux. Le Naturaliste. 3 (47): 373 - 374." type="journal article" year="1881">Ancey, 1881</bibRefCitation>
|
||
. We further classify
|
||
<taxonomicName box="[834,989,1113,1138]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="jinpingensis">
|
||
<emphasis box="[834,989,1113,1138]" italics="true" pageId="5" pageNumber="2016">B. jinpingensis</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName authorityName="Ancey" authorityYear="1881" box="[1026,1116,1113,1138]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[1026,1116,1113,1138]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
</taxonomicName>
|
||
for the same reason (i.e. that species is a member of
|
||
<taxonomicName authorityName="Ancey" authorityYear="1881" box="[730,820,1148,1173]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[730,820,1148,1173]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
</taxonomicName>
|
||
and not
|
||
<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[927,1043,1148,1173]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[927,1043,1148,1173]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="T. -C. Luo, D. -N. Chen & G. -Q. Zhang" authorityYear="2000" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="tianxingqiaoensis" subGenus="Bensonella">
|
||
<emphasis box="[1066,1156,1148,1173]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[168,284,1182,1207]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis box="[302,491,1182,1207]" italics="true" pageId="5" pageNumber="2016">tianxingqiaoensis</emphasis>
|
||
</taxonomicName>
|
||
(treated as a
|
||
<taxonomicName box="[658,840,1182,1207]" pageId="5" pageNumber="2016">
|
||
<emphasis box="[658,748,1182,1207]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Zhang W-H & Chen D-N & Zhou W-C" box="[889,1095,1182,1207]" pageId="5" pageNumber="2016" pagination="570 - 575" refId="ref8207" refString="Zhang W-H, Chen D-N, Zhou W-C. 2014. Two new species of the genus Boysidia from China, with a preliminary discussion of its geography (Pulmonata: Stylommatophora: Pupillidae). Zool Syst. 39 (4): 570 - 575." type="journal article" year="2014">Zhang et al. 2014</bibRefCitation>
|
||
) and
|
||
<taxonomicName authorityName="Y. -H. Guo, W. -C. Zhou & T. -C. Luo" authorityYear="2006" box="[160,528,1217,1242]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="xingyinensis" subGenus="Bensonella">
|
||
<emphasis box="[160,250,1217,1242]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[265,381,1217,1242]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
<emphasis box="[397,528,1217,1242]" italics="true" pageId="5" pageNumber="2016">xingyinensis</emphasis>
|
||
</taxonomicName>
|
||
are here both transferred to
|
||
<taxonomicName authority="Tomlin, 1930" authorityName="Tomlin" authorityYear="1930" box="[859,1155,1217,1242]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[859,1002,1217,1242]" italics="true" pageId="5" pageNumber="2016">Gyliotrachela</emphasis>
|
||
<bibRefCitation author="Tomlin JRB" box="[1010,1155,1217,1242]" pageId="5" pageNumber="2016" pagination="22 - 24" refId="ref7969" refString="Tomlin JRB. 1930. Some preoccupied generic names. - II. Proc Malacol Soc Lond. 19: 22 - 24." type="journal article" year="1930">Tomlin, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
due to the free and keeled last whorl and the separate parietal and angular lamella all being similar to those of the
|
||
<typeStatus box="[491,541,1286,1311]" pageId="5" pageNumber="2016">type</typeStatus>
|
||
species of that genus:
|
||
<taxonomicName authority="Mollendorff, 1891" authorityName="Mollendorff" authorityYear="1891" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="hungerfordianum">
|
||
<emphasis box="[804,1155,1286,1311]" italics="true" pageId="5" pageNumber="2016">Hypselostoma hungerfordianum</emphasis>
|
||
<bibRefCitation author="Mollendorff OF" box="[160,359,1321,1346]" pageId="5" pageNumber="2016" pagination="330 - 348" refId="ref7178" refString="Mollendorff OF von. 1891. On the land and freshwater shells of Perak. Proc Zool Soc Lond. 1891: 330 - 348." type="journal article" year="1891">Möllendorff, 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[160,1157,142,1693]" pageId="5" pageNumber="2016">
|
||
<taxonomicName authorityName="D. -N. Chen, Y. -H. Liu & W. -X. Wu" authorityYear="1995" box="[192,571,1356,1381]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="fengxianensis" subGenus="Boysidia">
|
||
<emphasis box="[192,284,1356,1381]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[306,398,1356,1381]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
)
|
||
<emphasis box="[421,571,1356,1381]" italics="true" pageId="5" pageNumber="2016">fengxianensis</emphasis>
|
||
</taxonomicName>
|
||
D.-N. Chen, Y.-H. Liu and W.-X. Wu, 1995 was transferred to
|
||
<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[332,451,1390,1415]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[332,451,1390,1415]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Schileyko AA" box="[502,693,1390,1415]" pageId="5" pageNumber="2016" pagination="129 - 261" refId="ref7802" refString="Schileyko AA. 1998. Treatise on recent terrestrial pulmonate molluscs. Part 2. Gastrocoptidae, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica. Supplement 2: 129 - 261." type="journal article" year="1998">Schileyko (1998)</bibRefCitation>
|
||
and to
|
||
<taxonomicName box="[794,1031,1390,1415]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="subGenus" subGenus="Bensonella">
|
||
<emphasis box="[794,885,1390,1415]" italics="true" pageId="5" pageNumber="2016">Boysidia</emphasis>
|
||
(
|
||
<emphasis box="[904,1023,1390,1415]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Zhang W-H & Chen D-N & Zhou W-C" pageId="5" pageNumber="2016" pagination="570 - 575" refId="ref8207" refString="Zhang W-H, Chen D-N, Zhou W-C. 2014. Two new species of the genus Boysidia from China, with a preliminary discussion of its geography (Pulmonata: Stylommatophora: Pupillidae). Zool Syst. 39 (4): 570 - 575." type="journal article" year="2014">Zhang et al. (2014)</bibRefCitation>
|
||
. Judging as best we can from the poor-quality drawing in the original description (mostly the three parietal lamellae), we agree that this species belongs to
|
||
<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[160,279,1494,1519]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[160,279,1494,1519]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[160,1157,142,1693]" pageId="5" pageNumber="2016">
|
||
Lastly, the Sumatran
|
||
<taxonomicName authority="Maassen, 1999" authorityName="Maassen" authorityYear="1999" box="[444,858,1529,1554]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="karoensis">
|
||
<emphasis box="[444,676,1529,1554]" italics="true" pageId="5" pageNumber="2016">Bensonella karoensis</emphasis>
|
||
<bibRefCitation author="Maassen WJM" box="[688,858,1529,1554]" pageId="5" pageNumber="2016" pagination="121 - 126" refId="ref7100" refString="Maassen WJM. 1999. Some new records of Streptaxidae and Hypselostomatidae from Sumatra, Indonesia, with descriptions of three new species. Basteria. 63: 121 - 126." type="journal article" year="1999">Maassen, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
is retained
|
||
<emphasis box="[1005,1155,1529,1554]" italics="true" pageId="5" pageNumber="2016">
|
||
in
|
||
<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[1037,1155,1529,1554]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">Bensonella</taxonomicName>
|
||
</emphasis>
|
||
herein, due to the arrangement of the apertural barriers (separate angular and parietal lamella, two palatal lamellae, a basal and a columellar lamella) that largely match with the situation in
|
||
<emphasis box="[290,345,1633,1658]" italics="true" pageId="5" pageNumber="2016">Pupa</emphasis>
|
||
<taxonomicName authority="Benson, 1849" authorityName="Benson" authorityYear="1849" box="[351,599,1633,1658]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="plicidens">
|
||
<emphasis box="[351,445,1633,1658]" italics="true" pageId="5" pageNumber="2016">plicidens</emphasis>
|
||
<bibRefCitation author="Benson WH" box="[451,599,1633,1658]" pageId="5" pageNumber="2016" pagination="125 - 128" refId="ref6320" refString="Benson WH. 1849. Descriptions of four new Asiatic species of the genus Pupa of Draparnaud. Ann Mag Nat Hist. 4 (20): 125 - 128. doi: 10.1080 / 03745486009496158." type="journal article" year="1849">Benson, 1849</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Bavay and Dautzenberg, 1912" authorityName="Bavay and Dautzenberg" authorityYear="1912" box="[653,1150,1633,1658]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="species" species="paviei">
|
||
<emphasis box="[653,812,1633,1658]" italics="true" pageId="5" pageNumber="2016">Boysidia paviei</emphasis>
|
||
<bibRefCitation author="Bavay A & Dautzenberg P" box="[818,1150,1633,1658]" pageId="5" pageNumber="2016" pagination="1 - 54" refId="ref6290" refString="Bavay A, Dautzenberg P. 1912. Description de coquilles nouvelles de l ̾ Indo-Chine. Journal de Conchyliologie. 60 (1): 1 - 54." type="journal article" year="1912">Bavay and Dautzenberg, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus box="[204,254,1668,1693]" pageId="5" pageNumber="2016">type</typeStatus>
|
||
species of
|
||
<taxonomicName authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[380,496,1668,1693]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[380,496,1668,1693]" italics="true" pageId="5" pageNumber="2016">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Pilsbry" authorityYear="1917" box="[555,695,1668,1693]" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="2016" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[555,695,1668,1693]" italics="true" pageId="5" pageNumber="2016">Paraboysidia</emphasis>
|
||
</taxonomicName>
|
||
, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |