171 lines
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171 lines
12 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.37.6499" ID-PMC="PMC4023333" ID-Pensoft-Pub="1314-2003-37-25" ID-Pensoft-UUID="FFE8FFACFF84FFA95573FFFECD03F742" ID-PubMed="24843297" ID-Zenodo-Dep="576215" ModsDocID="1314-2003-37-25" checkinTime="1451251747243" checkinUser="pensoft" docAuthor="Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C." docDate="2014" docId="1BA205E8CE845246B045785B87853DBB" docLanguage="en" docName="PhytoKeys 37: 25-101" docOrigin="PhytoKeys 37" docPubDate="2014-05-13" docSource="http://dx.doi.org/10.3897/phytokeys.37.6499" docTitle="Acilepis squarrosa D. Don, Prodr. Fl. Nepal.: 169. 1825." docType="treatment" docVersion="3" id="FFE8FFACFF84FFA95573FFFECD03F742" lastPageNumber="47" masterDocId="FFE8FFACFF84FFA95573FFFECD03F742" masterDocTitle="Revisions and key to the Vernonieae (Compositae) of Thailand" masterLastPageNumber="101" masterPageNumber="25" pageNumber="46" updateTime="1668141106797" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revisions and key to the Vernonieae (Compositae) of Thailand</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bunwong, Sukhonthip</mods:namePart>
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<mods:affiliation>Maejo University Phrae Campus, Mae Sai, Rong Kwang, Phrae 54140, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chantaranothai, Pranom</mods:namePart>
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<mods:affiliation>Applied Taxonomic Research Center, Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Keeley, Sterling C.</mods:namePart>
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<mods:affiliation>Department of Botany, University of Hawaii, Honolulu, HI 96816 USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-05-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>37</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>25</mods:start>
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<mods:end>101</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.37.6499</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.37.6499</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-37-25</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFE8FFACFF84FFA95573FFFECD03F742</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576215</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182228688" LSID="urn:lsid:plazi:treatment:1BA205E8CE845246B045785B87853DBB" httpUri="http://treatment.plazi.org/id/1BA205E8CE845246B045785B87853DBB" lastPageId="22" lastPageNumber="47" pageId="21" pageNumber="46">
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<subSubSection pageId="21" pageNumber="46" type="nomenclature">
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<paragraph pageId="21" pageNumber="46">
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<taxonomicName LSID="1BA205E8-CE84-5246-B045-785B87853DBB" authority="D. Don, Prodr. Fl. Nepal.: 169. 1825." authorityName="D. Don, Prodr. Fl. Nepal.: 169. 1825." class="Magnoliopsida" family="Asteraceae" genus="Acilepis" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Acilepis squarrosa" order="Asterales" pageId="21" pageNumber="46" phylum="Tracheophyta" rank="species" species="squarrosa">Acilepis squarrosa D.Don, Prodr. Fl. Nepal.: 169. 1825.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="46" type="reference_group">
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<paragraph pageId="21" pageNumber="46">
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<taxonomicName class="Insecta" family="Philopteridae" genus="Vernonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Phthiraptera" pageId="21" pageNumber="46" phylum="Arthropoda" rank="species" species="teres">Vernonia teres</taxonomicName>
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Wall., Numer. List [Wallich] no. 2926, comp. no. 36,
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<emphasis italics="true" pageId="21" pageNumber="46">nom. nud.</emphasis>
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</paragraph>
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<paragraph pageId="21" pageNumber="46">
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<taxonomicName class="Insecta" family="Philopteridae" genus="Vernonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Phthiraptera" pageId="21" pageNumber="46" phylum="Arthropoda" rank="species" species="squarrosa">Vernonia squarrosa</taxonomicName>
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(D.Don) Less., Linnaea 6: 678. 1831.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="46" type="type">
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<paragraph pageId="21" pageNumber="46">Type.</paragraph>
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<paragraph pageId="21" pageNumber="46">
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Nepal,
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<emphasis italics="true" pageId="21" pageNumber="46">Hamilton</emphasis>
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s.n. (not seen).
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Vernonieae in Thailand 1. A Acilepis attenuata B Acilepis divergens C Acilepis namnaoensis D Acilepis ngaoensis E Acilepis peguensis F Acilepis principis G Acilepis saligna H Acilepis silhetensis I Acilepis squarrosa." httpUri="https://binary.pensoft.net/fig/10646" pageId="21" pageNumber="46">Fig. 5I</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="46" type="description">
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<paragraph pageId="21" pageNumber="46">Description.</paragraph>
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<paragraph pageId="21" pageNumber="46">Perennial herbs, 30-80 cm tall. Stems erect, inconspicuously ribbed, villose. Leaves 3-10 by 1-3 cm, oblanceolate, margin serrate, apex acute, base cuneate, coriaceous; both surfaces scabrous with whip-shaped hairs and capitate glands; lateral veins 5-10-paired; petioles up to 5 mm long. Capitulescences terminal and axillary, solitary. Capitula campanulate, 15-20 mm long, sessile or subsessile. Receptacle flat, 4.5-5.5 mm in diam., glabrous. Involucres campanulate, in 12-13 series, 15-20 mm long, 10-15 mm in diam., herbaceous. Phyllaries light green or purple apically, margin piliferous, outer surface arachnoid and lacking glands; the outer and the middle ones ovate or lanceolate, apex acuminate or apiculate; the inner ones ovate-lanceolate to oblong, apex acuminate or apiculate. Florets 50-80; corollas funnelform, purple, puberulous, glands capitate; corolla tubes 7-10 mm long; corolla lobes 4-6 mm long. Anthers 3-3.5 mm long, apical appendage acute, base obtuse. Styles purple, 9-11 mm long, branches 2-3 mm long. Achenes subterete, 2.8-3.2 mm long, 10-ribbed, pubescent without glands. Pappus bristles, the inner ones 7-11 mm long.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="46" type="distribution">
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<paragraph pageId="21" pageNumber="46">Distribution.</paragraph>
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<paragraph pageId="21" pageNumber="46">Thailand: Mae Hong Son, Chiang Mai, Chiang Rai, Lamphun, Lampang, Tak, Phetchabun, Loei, Sakon Nakhon, Mukdahan, Kalasin, Khon Kaen, Chaiyaphum, Nakhon Ratchasima, Surin, Uthai Thani, Kanchanaburi, Lop Buri, Prachin Buri, Chon Buri. China (Yunnan), India, Bhutan, Myanmar, Laos.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="22" lastPageNumber="47" pageId="21" pageNumber="46" type="specimens examined">
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<paragraph pageId="21" pageNumber="46">Specimens examined.</paragraph>
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<paragraph lastPageId="22" lastPageNumber="47" pageId="21" pageNumber="46">
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Thailand, Phetchabun, Nam Nao national park,
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<geoCoordinate degrees="16" direction="north" minutes="44.29" orientation="latitude" precision="9" value="16.738167">16°44.29'N</geoCoordinate>
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,
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<geoCoordinate degrees="101" direction="east" minutes="34.19" orientation="longitude" precision="9" value="101.56983">101°34.19'E</geoCoordinate>
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, 26 Dec 2007,
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<emphasis italics="true" pageId="22" pageNumber="47">
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<pageBreakToken pageId="22" pageNumber="47" start="start">S</pageBreakToken>
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. Bunwong
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</emphasis>
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374 (KKU, US); Mae Hong Son, Pai, 16 Jan 1983,
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<emphasis italics="true" pageId="22" pageNumber="47">H. Koyama & H. Terao & Th. Wongprasert</emphasis>
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T-32609 (BKF); Pai, 16 Jan 1786,
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<emphasis italics="true" pageId="22" pageNumber="47">Y. Paisooksantivatana</emphasis>
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1780-86 (BK); Pai, 23 Nov 1989,
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<emphasis italics="true" pageId="22" pageNumber="47">Y. Paisooksantivatana</emphasis>
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2548-89 (BK); Chiang Mai, Hod, 13 Jan 1983,
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<emphasis italics="true" pageId="22" pageNumber="47">H. Koyama & H. Terao & Th. Wongprasert</emphasis>
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T-32454 (BKF); Hod, 13 Nov 1965,
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<emphasis italics="true" pageId="22" pageNumber="47">Prayad</emphasis>
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44 (BK); Hod, 4 Dec 1975,
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<emphasis italics="true" pageId="22" pageNumber="47">J. Sadakorn</emphasis>
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664 (BK); Lamphun, Maeli, 3 Nov 1925,
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<emphasis italics="true" pageId="22" pageNumber="47">Winit</emphasis>
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1515 (BKF, K); Lampang, Doi Khun Tan, 28 Dec 1984,
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<emphasis italics="true" pageId="22" pageNumber="47">H. Koyama & C. Phengklai</emphasis>
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T-39155 (BKF, L); Tak, Maesod, 13 Jan 1989,
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<emphasis italics="true" pageId="22" pageNumber="47">Y. Paisooksantivatana</emphasis>
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2306-89 (BK); Phetchabun, Nam Nao national park, 18 Nov 1973,
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<emphasis italics="true" pageId="22" pageNumber="47">J.F. Maxwell</emphasis>
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73-630 (AAU, BK); Nam Nao national park, 23 Jun 1975,
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<emphasis italics="true" pageId="22" pageNumber="47">J.F. Maxwell</emphasis>
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75-626 (AAU, BK, L); Loei, Paa See Than, 2 Sep 1991,
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<emphasis italics="true" pageId="22" pageNumber="47">Dee</emphasis>
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321 (BKF); Sakon Nakhon, 23 Nov 1962,
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<emphasis italics="true" pageId="22" pageNumber="47">Adisai</emphasis>
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155 (BK); Sakon Nakhon, Phu Phan, 15 Nov 1984,
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<emphasis italics="true" pageId="22" pageNumber="47">G. Murata & C. Phengklai</emphasis>
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50372 (BKF); Mukdahan, route number 212, 12 Dec 1982,
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<emphasis italics="true" pageId="22" pageNumber="47">H. Koyama & H. Terao & Th. Wongprasert</emphasis>
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T-32161 (BKF); Kalasin, Phu Sing, Sahadsakhan, 22 Oct 1975,
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<emphasis italics="true" pageId="22" pageNumber="47">S. Sutheesorn</emphasis>
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3503 (BK); Khon Kaen, route to Nam Nao national park, 18 Nov 1979,
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<emphasis italics="true" pageId="22" pageNumber="47">T. Shimitzu, H. Toyokuni, H. Koyama, T. Yahara & C. Niyomdham</emphasis>
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T-22533 (BKF, L); Chaiyaphum, Kang Kraw, 23 Oct 1965,
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<emphasis italics="true" pageId="22" pageNumber="47">S. Sutheesorn</emphasis>
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3048 (BK); Nakhon Ratchasima, Pak Thong Chai, 10 Nov 1963,
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<emphasis italics="true" pageId="22" pageNumber="47">Pradit</emphasis>
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523 (BK); Hui Taleng, 21 Dec 1928,
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<emphasis italics="true" pageId="22" pageNumber="47">Put</emphasis>
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2175 (BK, BM, K); Surin, 3 Dec 1976,
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<emphasis italics="true" pageId="22" pageNumber="47">C. Phengklai et al.</emphasis>
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3595 (BKF); Uthai Thani, 4 Dec 1977,
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<emphasis italics="true" pageId="22" pageNumber="47">C. Phengklai et al.</emphasis>
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3993 (BKF); Kanchanaburi, Thong Pha Phume, 25 Jan 1983,
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<emphasis italics="true" pageId="22" pageNumber="47">H. Koyama & H. Terao & Th. Wongprasert</emphasis>
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T-32909 (BKF); Thong Pha Phume,
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<emphasis italics="true" pageId="22" pageNumber="47">T. Shimitzu, H. Toyokuni, H. Koyama, T. Yahara & C. Niyomdham</emphasis>
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T-21892 (BKF, L).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="47" type="diagnostic characters">
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<paragraph pageId="22" pageNumber="47">Diagnostic characters.</paragraph>
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<paragraph pageId="22" pageNumber="47">
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Acilepis" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="22" pageNumber="47" phylum="Tracheophyta" rank="species" species="squarrosa">
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<emphasis italics="true" pageId="22" pageNumber="47">Acilepis squarrosa</emphasis>
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</taxonomicName>
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is easily recognized by its large capitula (with more than 100 phyllaries) sessile in the axils of the leaves
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="47" type="ecology">
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<paragraph pageId="22" pageNumber="47">Ecology.</paragraph>
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<paragraph pageId="22" pageNumber="47">Dipterocarp, deciduous, hill evergreen or pine-oak forest, alt. 10-1200 m; flowering September to May.</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="47" type="vernacular names">
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<paragraph pageId="22" pageNumber="47">Vernacular names.</paragraph>
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<paragraph pageId="22" pageNumber="47">Kiang Pa Chang (เกี๋ยงพาช้าง), Nat Dhum (หนาดดำ), Nat Khum (หนาดคำ).</paragraph>
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</subSubSection>
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</treatment>
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</document> |