309 lines
31 KiB
XML
309 lines
31 KiB
XML
<document ID-DOI="10.11646/zootaxa.5254.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7732141" ID-ZooBank="96609374-84D0-4032-A3F0-0B19DEAF4E03" checkinTime="1678782987451" checkinUser="plazi" docAuthor="Chandramouli, S. R., Adhikari, Omkar D., Amarasinghe, A. A. Thasun & Abinawanto, A." docDate="2023" docId="1B6487E70F31371CFF02FBA2E8D4FAE7" docLanguage="en" docName="zootaxa.5254.4.3.pdf" docOrigin="Zootaxa 5254 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5254.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bronchocela cyanopalpebra Chandramouli & Adhikari & Amarasinghe & Abinawanto 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="499" masterDocId="E75DFF9F0F35371AFF95FFC0EC41FFD6" masterDocTitle="A review of the genus Bronchocela Kaup, 1827 (Reptilia: Agamidae) in the Nicobar Archipelago with the description of two new species" masterLastPageNumber="516" masterPageNumber="493" pageNumber="497" updateTime="1678812247699" updateUser="tatiana" zenodo-license-document="CLOSED">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A review of the genus Bronchocela Kaup, 1827 (Reptilia: Agamidae) in the Nicobar Archipelago with the description of two new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Chandramouli, S. R.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0003-0626-0527</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Ecology and Environmental Sciences, School of Life Sciences, Pondicherry University, Puducherry 605014, India findthesnakeman @ gmail. com; https: // orcid. org / 0000 - 0003 - 0626 - 0527</mods:affiliation>
|
||
<mods:nameIdentifier type="email">findthesnakeman@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Adhikari, Omkar D.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0003-2921-5593</mods:nameIdentifier>
|
||
<mods:affiliation>Natural History Collections Department, Bombay Natural History Society, Hornbill House, Opp. Lion Gate, S. B. S. Road, Fort, Mumbai 400001, India. proahaetulla @ gmail. com; https: // orcid. org / 0000 - 0003 - 2921 - 5593</mods:affiliation>
|
||
<mods:nameIdentifier type="email">proahaetulla@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Amarasinghe, A. A. Thasun</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0002-4151-1806</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Indonesia, Kampus UI, Depok 16424, Indonesia & Herpetology Lab, Research Center for Biosystematics and Evolution, The National Research and Innovation Agency (BRIN), Government of Indonesia, Cibinong 16911, Indonesia thasun. amarasinghe @ ui. ac. id; https: // orcid. org / 0000 - 0002 - 4151 - 1806</mods:affiliation>
|
||
<mods:nameIdentifier type="email">thasun.amarasinghe@ui.ac.id</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Abinawanto, A.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0003-0181-9336</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Biology, Faculty of Mathematics and Natural Sciences, Universitas Indonesia, Kampus UI, Depok 16424, Indonesia & abinawanto. ms @ sci. ui. ac. id; https: // orcid. org / 0000 - 0003 - 0181 - 9336</mods:affiliation>
|
||
<mods:nameIdentifier type="email">abinawanto.ms@sci.ui.ac.id</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-03-14</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5254</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>493</mods:start>
|
||
<mods:end>516</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/zootaxa.5254.4.3</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.5254.4.3</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7732141</mods:identifier>
|
||
<mods:identifier type="ZooBank">96609374-84D0-4032-A3F0-0B19DEAF4E03</mods:identifier>
|
||
</mods:mods>
|
||
<treatment LSID="urn:lsid:plazi:treatment:1B6487E70F31371CFF02FBA2E8D4FAE7" httpUri="http://treatment.plazi.org/id/1B6487E70F31371CFF02FBA2E8D4FAE7" lastPageId="6" lastPageNumber="499" pageId="4" pageNumber="497">
|
||
<subSubSection box="[151,592,1122,1149]" pageId="4" pageNumber="497" type="nomenclature">
|
||
<paragraph blockId="4.[151,592,1122,1185]" box="[151,592,1122,1149]" pageId="4" pageNumber="497">
|
||
<heading bold="true" box="[151,592,1122,1149]" fontSize="11" level="1" pageId="4" pageNumber="497" reason="1">
|
||
<emphasis bold="true" box="[151,592,1122,1149]" pageId="4" pageNumber="497">
|
||
<taxonomicName authority="Chandramouli & Adhikari & Amarasinghe & Abinawanto, 2023" authorityName="Chandramouli & Adhikari & Amarasinghe & Abinawanto" authorityYear="2023" box="[151,491,1122,1149]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="cyanopalpebra" status="sp. nov.">
|
||
<emphasis bold="true" box="[151,491,1122,1149]" italics="true" pageId="4" pageNumber="497">Bronchocela cyanopalpebra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[498,592,1123,1149]" pageId="4" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,236,1159,1185]" pageId="4" pageNumber="497" type="description">
|
||
<paragraph blockId="4.[151,592,1122,1185]" box="[151,236,1159,1185]" pageId="4" pageNumber="497">
|
||
(
|
||
<figureCitation box="[159,227,1159,1185]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURE 2. Bronchocela cyanopalpebra sp. nov. (top: Holotype DOSMB 05054, middle: live male, below: live female)." figureDoi="http://doi.org/10.5281/zenodo.7732145" httpUri="https://zenodo.org/record/7732145/files/figure.png" pageId="4" pageNumber="497">Fig. 2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="497" type="reference_group">
|
||
<paragraph blockId="4.[151,1214,1227,1374]" box="[151,965,1227,1251]" pageId="4" pageNumber="497">
|
||
<taxonomicName authorityName="Fitzinger" authorityYear="1860" box="[151,444,1227,1251]" class="Reptilia" family="Agamidae" genus="Pseudocalotes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="archiducissae">
|
||
<emphasis box="[151,444,1227,1251]" italics="true" pageId="4" pageNumber="497">Pseudocalotes archiducissae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[458,601,1227,1251]" italics="true" pageId="4" pageNumber="497">nomen nudum</emphasis>
|
||
)—Fitzinger in
|
||
<bibRefCitation author="Steindachner, F." box="[759,965,1227,1251]" pageId="4" pageNumber="497" refId="ref11736" refString="Steindachner, F. (1867) Reise der Osterreichischen Fregatte Novara um die Erdein den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Zoologischer Theil. Erster Band (Reptilien). Kaiserlich-Koniglichen Hof- und Staatsdruckerei, Wien, 98 pp., 3 pls." type="book" year="1867">Steindachner (1867)</bibRefCitation>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1214,1227,1374]" box="[151,1089,1258,1282]" pageId="4" pageNumber="497">
|
||
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[151,348,1258,1282]" class="Reptilia" family="Agamidae" genus="Calotes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="cristatellus">
|
||
<emphasis box="[151,348,1258,1282]" italics="true" pageId="4" pageNumber="497">Calotes cristatellus</emphasis>
|
||
</taxonomicName>
|
||
(non
|
||
<bibRefCitation author="Kuhl, H." box="[407,520,1258,1282]" pageId="4" pageNumber="497" pagination="1" refId="ref11512" refString="Kuhl, H. (1820) Beitrage zur Zoologie und vergleichenden Anatomie. Erste Abtheilung. Beitrage zur Zoologie. Hermannschen Buchhandlung, Frankfurt am Main, [ii] + 151 + [1] pp." type="journal article" year="1820">Kuhl, 1820</bibRefCitation>
|
||
)—
|
||
<bibRefCitation author="Biswas, S. & Sanyal, D. P." box="[554,805,1258,1282]" pageId="4" pageNumber="497" pagination="107 - 124" refId="ref10540" refString="Biswas, S. & Sanyal, D. P. (1977) Notes on the Reptilia collection from the Great Nicobar Island during the Great Nicobar Expedition in 1966. Records of the Zoological Survey of India, 72, 107 - 124. https: // doi. org / 10.26515 / rzsi / v 72 / i 1 - 4 / 1977 / 161927" type="journal article" year="1977">Biswas & Sanyal (1977)</bibRefCitation>
|
||
, (1980);
|
||
<bibRefCitation author="Biswas, S." box="[896,1043,1258,1282]" pageId="4" pageNumber="497" pagination="476 - 481" refId="ref10608" refString="Biswas, S. (1984) Some notes on the reptiles of the Andaman and Nicobar Islands. Journal of the Bombay Natural History Society, 81, 476 - 481." type="journal article" year="1984">Biswas (1984)</bibRefCitation>
|
||
part
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1214,1227,1374]" box="[151,981,1288,1313]" pageId="4" pageNumber="497">
|
||
<taxonomicName baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[151,389,1288,1312]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="cristatella">
|
||
<emphasis box="[151,389,1288,1312]" italics="true" pageId="4" pageNumber="497">Bronchocela cristatella</emphasis>
|
||
</taxonomicName>
|
||
(non
|
||
<bibRefCitation author="Kuhl, H." box="[448,561,1288,1313]" pageId="4" pageNumber="497" pagination="1" refId="ref11512" refString="Kuhl, H. (1820) Beitrage zur Zoologie und vergleichenden Anatomie. Erste Abtheilung. Beitrage zur Zoologie. Hermannschen Buchhandlung, Frankfurt am Main, [ii] + 151 + [1] pp." type="journal article" year="1820">Kuhl, 1820</bibRefCitation>
|
||
)—
|
||
<bibRefCitation author="Das, I." box="[595,712,1289,1313]" pageId="4" pageNumber="497" pagination="43 - 77" refId="ref10932" refString="Das, I. (1999 b) Biogeography of the amphibians and reptiles of the Andaman and Nicobar Islands. In: Ota, H. (Ed.), Tropical Island Herpetofauna. Origin, Current Diversity and Conservation. Developments in Animal and Veterinary Sciences, 29, pp. 43 - 77." type="journal article" year="1999">Das (1999)</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Das, I. & Gemel, R." box="[722,935,1288,1313]" pageId="4" pageNumber="497" pagination="55 - 58" refId="ref10990" refString="Das, I. & Gemel, R. (2000) Nomenclatural status of Fitzinger's (1861) Pseudocalotes archiducissae, and confirmation of Bronchocela cristatella (Kuhl, 1820) from the Nicobar Archipelago (Squamata: Sauria: Agamidae). Herpetozoa, 13 (1 - 2), 55 - 58." type="journal article" year="2000">Das & Gemel (2000)</bibRefCitation>
|
||
part
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1214,1227,1374]" box="[151,545,1319,1343]" pageId="4" pageNumber="497">
|
||
<taxonomicName box="[151,311,1319,1343]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis box="[151,280,1319,1343]" italics="true" pageId="4" pageNumber="497">Bronchocela</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation author="Vijayakumar, S. P." box="[336,545,1319,1343]" pageId="4" pageNumber="497" refId="ref11959" refString="Vijayakumar, S. P. (2005) Status and distribution of Amphibians and Reptiles of the Nicobar Islands, India. Final Report. Rufford Foundation / Madras Crocodile Bank / Wildlife Institute of India, 48 pp." type="book" year="2005">Vijayakumar (2005)</bibRefCitation>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1214,1227,1374]" box="[151,1214,1350,1374]" pageId="4" pageNumber="497">
|
||
<taxonomicName baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[151,421,1350,1374]" class="Reptilia" family="Agamidae" genus="Bronchocela" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="cristatella">
|
||
<emphasis box="[151,280,1350,1374]" italics="true" pageId="4" pageNumber="497">Bronchocela</emphasis>
|
||
cf.
|
||
<emphasis box="[318,421,1350,1374]" italics="true" pageId="4" pageNumber="497">cristatella</emphasis>
|
||
</taxonomicName>
|
||
(non
|
||
<bibRefCitation author="Kuhl, H." box="[480,593,1350,1374]" pageId="4" pageNumber="497" pagination="1" refId="ref11512" refString="Kuhl, H. (1820) Beitrage zur Zoologie und vergleichenden Anatomie. Erste Abtheilung. Beitrage zur Zoologie. Hermannschen Buchhandlung, Frankfurt am Main, [ii] + 151 + [1] pp." type="journal article" year="1820">Kuhl, 1820</bibRefCitation>
|
||
)—Harikrishnan
|
||
<emphasis box="[767,818,1350,1374]" italics="true" pageId="4" pageNumber="497">et al.</emphasis>
|
||
(2009) part; Chandramouli (2020) part
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="497" type="materials_examined">
|
||
<paragraph blockId="4.[151,1437,1420,2023]" pageId="4" pageNumber="497">
|
||
<materialsCitation collectingDate="2017-06" collectorName="S. R. Chandramouli" country="India" elevation="33" latitude="9.21358" location="Nicobar Islands" longLatPrecision="1" longitude="92.79647" pageId="4" pageNumber="497" specimenCount="1" typeStatus="holotype">
|
||
<emphasis bold="true" box="[151,269,1420,1446]" pageId="4" pageNumber="497">
|
||
<typeStatus box="[151,264,1420,1446]" pageId="4" pageNumber="497">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
DOSMB 05054, an adult male collected from Chukchuka (
|
||
<geoCoordinate box="[976,1100,1420,1447]" degrees="9.21358" direction="north" orientation="latitude" pageId="4" pageNumber="497" precision="1" value="9.21358">9.21358ºN</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1115,1249,1420,1446]" degrees="92.79647" direction="east" orientation="longitude" pageId="4" pageNumber="497" precision="1" value="92.79647">92.79647ºE</geoCoordinate>
|
||
,
|
||
<elevation box="[1265,1372,1420,1447]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="497" unit="m" value="33.0">
|
||
<quantity box="[1265,1326,1420,1447]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="497" unit="m" value="33.0">33 m</quantity>
|
||
asl
|
||
</elevation>
|
||
.) Car Nicobar,
|
||
<collectingCountry box="[254,433,1456,1482]" name="India" pageId="4" pageNumber="497">Nicobar Islands</collectingCountry>
|
||
,
|
||
<collectingCountry box="[445,503,1456,1482]" name="India" pageId="4" pageNumber="497">India</collectingCountry>
|
||
by
|
||
<collectorName box="[544,760,1456,1482]" pageId="4" pageNumber="497">S.R. Chandramouli</collectorName>
|
||
in
|
||
<date box="[796,914,1456,1483]" pageId="4" pageNumber="497" value="2017-06">
|
||
<collectingDate box="[796,914,1456,1483]" pageId="4" pageNumber="497" value="2017-06">June 2017</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1420,2023]" box="[199,1310,1492,1519]" pageId="4" pageNumber="497">
|
||
<emphasis bold="true" box="[199,317,1492,1518]" pageId="4" pageNumber="497">
|
||
<typeStatus box="[199,312,1492,1518]" pageId="4" pageNumber="497">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
DOSMB 05089, an adult male collected along with the
|
||
<typeStatus box="[948,1046,1492,1518]" pageId="4" pageNumber="497">holotype</typeStatus>
|
||
from the same locality.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1420,2023]" pageId="4" pageNumber="497">
|
||
<emphasis bold="true" box="[199,423,1528,1554]" pageId="4" pageNumber="497">Referred material:</emphasis>
|
||
BNHS 1610 (adult male), BNHS 1662 (adult female) collected from Air-force station, Car Nicobar by S. Krishnan in 2004; BNHS 961 (adult male) collected from Car Nicobar by Humayun Abdulali & party in 1966.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="497" type="etymology">
|
||
<paragraph blockId="4.[151,1437,1420,2023]" pageId="4" pageNumber="497">
|
||
<emphasis bold="true" box="[199,338,1636,1662]" pageId="4" pageNumber="497">Etymology:</emphasis>
|
||
The specific epithet
|
||
<taxonomicName authorityName="Chandramouli & Adhikari & Amarasinghe & Abinawanto" authorityYear="2023" box="[582,750,1636,1662]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="cyanopalpebra">
|
||
<emphasis box="[582,750,1636,1662]" italics="true" pageId="4" pageNumber="497">cyanopalpebra</emphasis>
|
||
</taxonomicName>
|
||
is a noun in apposition referring to the bright blue (=cyan) coloured eyelid (=palpebra), a diagnostic character of this species.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1420,2023]" pageId="4" pageNumber="497">
|
||
<emphasis bold="true" box="[199,447,1708,1734]" pageId="4" pageNumber="497">Nomenclatural note:</emphasis>
|
||
As the name
|
||
<taxonomicName box="[606,931,1708,1734]" class="Reptilia" family="Agamidae" genus="Pseudocalotes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="archidussicae">
|
||
<emphasis box="[606,931,1708,1734]" italics="true" pageId="4" pageNumber="497">Pseudocalotes archidussicae</emphasis>
|
||
</taxonomicName>
|
||
, attributed to the Car Nicobar population by Fitzinger in:
|
||
<bibRefCitation author="Steindachner, F." box="[296,523,1744,1770]" pageId="4" pageNumber="497" refId="ref11736" refString="Steindachner, F. (1867) Reise der Osterreichischen Fregatte Novara um die Erdein den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Zoologischer Theil. Erster Band (Reptilien). Kaiserlich-Koniglichen Hof- und Staatsdruckerei, Wien, 98 pp., 3 pls." type="book" year="1867">Steindachner (1867)</bibRefCitation>
|
||
has been shown to be a
|
||
<emphasis box="[796,953,1744,1770]" italics="true" pageId="4" pageNumber="497">nomen nudum</emphasis>
|
||
(
|
||
<bibRefCitation author="Das, I. & Gemel, R." box="[968,1197,1744,1770]" pageId="4" pageNumber="497" pagination="55 - 58" refId="ref10990" refString="Das, I. & Gemel, R. (2000) Nomenclatural status of Fitzinger's (1861) Pseudocalotes archiducissae, and confirmation of Bronchocela cristatella (Kuhl, 1820) from the Nicobar Archipelago (Squamata: Sauria: Agamidae). Herpetozoa, 13 (1 - 2), 55 - 58." type="journal article" year="2000">Das & Gemel, 2000</bibRefCitation>
|
||
), it cannot be applied to this species as it is not an available nomen (ICZN Art. 11.6).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="499" pageId="4" pageNumber="497" type="diagnosis">
|
||
<paragraph blockId="4.[151,1437,1420,2023]" lastBlockId="6.[151,1437,150,1329]" lastPageId="6" lastPageNumber="499" pageId="4" pageNumber="497">
|
||
<emphasis bold="true" box="[199,673,1816,1842]" pageId="4" pageNumber="497">Differential diagnosis and comparisons:</emphasis>
|
||
A species of
|
||
<taxonomicName authorityName="Kaup" authorityYear="1827" box="[822,964,1816,1842]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="genus">
|
||
<emphasis box="[822,964,1816,1842]" italics="true" pageId="4" pageNumber="497">Bronchocela</emphasis>
|
||
</taxonomicName>
|
||
endemic to the northern group of
|
||
<collectingCountry name="India" pageId="4" pageNumber="497">Nicobar Islands</collectingCountry>
|
||
characterized by: moderate to large body size (
|
||
<quantity box="[784,980,1852,1879]" metricMagnitude="-2" metricUnit="m" metricValue="9.241499999999998" metricValueMax="10.36" metricValueMin="8.123" pageId="4" pageNumber="497" unit="mm" value="92.41499999999999" valueMax="103.6" valueMin="81.23">81.23–103.6 mm</quantity>
|
||
SVL); relative tail length (336 % SVL
|
||
<emphasis box="[151,223,1889,1914]" italics="true" pageId="4" pageNumber="497">versus</emphasis>
|
||
330-390 % SVL in
|
||
<taxonomicName baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[464,612,1888,1914]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="cristatella">
|
||
<emphasis box="[464,612,1888,1914]" italics="true" pageId="4" pageNumber="497">B. cristatella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[622,769,1888,1915]" pageId="4" pageNumber="497" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
); 56–78 scale rows around mid-body (
|
||
<emphasis box="[1218,1290,1889,1914]" italics="true" pageId="4" pageNumber="497">versus</emphasis>
|
||
<quantity box="[1300,1402,1888,1914]" metricMagnitude="0" metricUnit="m" metricValue="1.778" metricValueMax="2.0065999999999997" metricValueMin="1.5493999999999999" pageId="4" pageNumber="497" unit="in" value="70.0" valueMax="79.0" valueMin="61.0">61–79 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kuhl" baseAuthorityYear="1820" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="497" phylum="Chordata" rank="species" species="cristatella">
|
||
<emphasis italics="true" pageId="4" pageNumber="497">B. cristatella</emphasis>
|
||
</taxonomicName>
|
||
), of which, the first 3–4 rows on the dorsum oriented posterodorsally, the next 1–3 rows oriented posteriorly, followed by 17–22 rows oriented posteroventrally; 7–12 enlarged and elongated scales forming the nuchal crest; dorsal crest merely a serrated ridge; three postmentals; 9–10 supralabials; 8–10 infralabials; 5–6 canthals; 6–8 supraciliaries; 32–43 subdigital lamellae under toe IV (
|
||
<emphasis box="[824,896,151,176]" italics="true" pageId="6" pageNumber="499">versus</emphasis>
|
||
<quantity box="[903,1003,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="8.254999999999999" metricValueMax="8.636" metricValueMin="7.874" pageId="6" pageNumber="499" unit="in" value="32.5" valueMax="34.0" valueMin="31.0">31–34 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[1011,1157,151,177]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="499" phylum="Chordata" rank="species" species="cristatella">
|
||
<emphasis box="[1011,1157,151,177]" italics="true" pageId="6" pageNumber="499">B. cristatella</emphasis>
|
||
</taxonomicName>
|
||
); 76–85 ventrals (
|
||
<emphasis box="[1365,1437,151,176]" italics="true" pageId="6" pageNumber="499">versus</emphasis>
|
||
<quantity box="[151,251,187,213]" metricMagnitude="0" metricUnit="m" metricValue="1.8542" metricValueMax="2.1844" metricValueMin="1.524" pageId="6" pageNumber="499" unit="in" value="73.0" valueMax="86.0" valueMin="60.0">60–86 in</quantity>
|
||
<taxonomicName baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[258,404,187,213]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="499" phylum="Chordata" rank="species" species="cristatella">
|
||
<emphasis box="[258,404,187,213]" italics="true" pageId="6" pageNumber="499">B. cristatella</emphasis>
|
||
</taxonomicName>
|
||
); bright grass green dorsal colouration with a distinctly cyan coloured ring around the eyes (
|
||
<emphasis box="[161,233,223,248]" italics="true" pageId="6" pageNumber="499">versus</emphasis>
|
||
absent in
|
||
<taxonomicName baseAuthorityName="Kuhl" baseAuthorityYear="1820" box="[346,491,223,249]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="499" phylum="Chordata" rank="species" species="cristatella">
|
||
<emphasis box="[346,491,223,249]" italics="true" pageId="6" pageNumber="499">B. cristatella</emphasis>
|
||
</taxonomicName>
|
||
) in both sexes (
|
||
<tableCitation box="[668,756,223,249]" captionStart="TABLE 2" captionStartId="12.[151,239,764,788]" captionTargetPageId="12" captionText="TABLE 2. Comparison of morphological characters of Bronchocela spp. of the Nicobar Islands and B. cristatella" pageId="6" pageNumber="499">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7732145" ID-Zenodo-Dep="7732145" box="[151,1383,1943,1968]" httpUri="https://zenodo.org/record/7732145/files/figure.png" pageId="5" pageNumber="498" startId="5.[151,250,1943,1967]" targetBox="[179,1408,181,1920]" targetPageId="5">
|
||
<paragraph blockId="5.[151,1383,1943,1968]" box="[151,1383,1943,1968]" pageId="5" pageNumber="498">
|
||
<emphasis bold="true" box="[151,269,1943,1967]" pageId="5" pageNumber="498">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName authority="Chandramouli & Adhikari & Amarasinghe & Abinawanto, 2023" authorityName="Chandramouli & Adhikari & Amarasinghe & Abinawanto" authorityYear="2023" box="[281,568,1944,1968]" class="Reptilia" family="Agamidae" genus="Bronchocela" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="498" phylum="Chordata" rank="species" species="cyanopalpebra" status="sp. nov.">
|
||
<emphasis box="[281,568,1944,1968]" italics="true" pageId="5" pageNumber="498">Bronchocela cyanopalpebra</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[575,655,1944,1968]" pageId="5" pageNumber="498">
|
||
<taxonomicNameLabel box="[575,655,1944,1968]" pageId="5" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(top: Holotype DOSMB 05054, middle: live male, below: live female).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="6" pageNumber="499" type="description">
|
||
<paragraph blockId="6.[151,1437,150,1329]" pageId="6" pageNumber="499">
|
||
<emphasis bold="true" box="[199,537,258,285]" pageId="6" pageNumber="499">
|
||
Description of the
|
||
<typeStatus box="[419,532,259,285]" pageId="6" pageNumber="499">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
An adult male, measuring
|
||
<quantity box="[839,953,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="8.123" pageId="6" pageNumber="499" unit="mm" value="81.23">81.23 mm</quantity>
|
||
SVL; with a very long tail (TaL:SVL 3.39) with a short and robust body (AG:SVL 0.45). Head large (HL:SVL 0.28); longer than broad (HL:HW 1.75) and about as broad as deep (HW:HD 1.04). Snout long (ES:HL 0.45) with a rounded tip. Canthus rostralis sharply defined, with five canthal scales. Eyes relatively small, about 1/5
|
||
<superScript attach="left" box="[780,793,366,380]" fontSize="6" pageId="6" pageNumber="499">th</superScript>
|
||
the length of the head (ED:HL 0.21); tympanum circular, relatively small, about half as large as the orbit (TYH:ED 0.51) and black in colour. Scales between the eye and tympanum slightly enlarged and keeled. Nasal relatively large, with a pointed anterior and rounded posterior ends. Scales on the check below the tympanum slightly enlarged and keeled. Mental semicircular, bounded by three small postmentals and nine infralabials on either sides. Rostral relatively small, bounded by nine supralabials on each side. Nostrils separated from each other by nine small internasals. Nuchal crest composed of nine slightly enlarged, triangular scales projecting posterodorsally. Dorsal body scales relatively small, feebly keeled and homogeneous, in 68 rows around the mid-body. Of the dorsal scales, the first two rows on the dorsum on either sides oriented posteriorly; the next three rows of scales below oriented posterodorsally, with the rest of the scales on the lateral body oriented posteroventrally. Ventrals
|
||
<quantity box="[601,657,691,717]" metricMagnitude="0" metricUnit="m" metricValue="2.032" pageId="6" pageNumber="499" unit="in" value="80.0">80 in</quantity>
|
||
a longitudinal series; much larger (about twice) than the dorsal scales, bearing a strong median keel, projecting posteriorly towards the lower vertex. Limbs relatively robust and elongate; upper arm short and slender (UAL:SVL 0.19); lower arm nearly as long as the upper arm (UAL:LAL 1.05); palm about as long as the lower arm (LAL:PAL 1.06). Fingers long and slender, relative length of fingers IV>III>V>II>I. Thigh long and slender (FEL:SVL 0.25); tibia slightly longer than thighs (FEL:TBL 0.95); foot elongated, slightly longer than thigh (FEL:FOL 0.87). Hindlimbs much longer than half the body length (HLL:SVL 0.82). Relative lengths of toes IV>III>V>II>I; 35 subdigital lamellae under toe IV.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,150,1329]" pageId="6" pageNumber="499">Dorsal coloration verdant green throughout the body; tip of the tail reddish. Ventral side of the head and body of a lighter shade of green than the dorsum. Sides of the lips, cheeks and eyelids bright aqua blue in color. Lips with an orange-red streak; tympanum black. Upper portion of the head slightly more bluish. In preservation, overall body colouration bright green with trunk and posterior edge of the neck being dark grey; tail brownish. Venter greenish brown without any specific pattern on the throat.</paragraph>
|
||
<paragraph blockId="6.[151,1437,150,1329]" pageId="6" pageNumber="499">
|
||
<emphasis bold="true" box="[199,321,1122,1148]" pageId="6" pageNumber="499">Variation:</emphasis>
|
||
Measurements and scale-counts of the male
|
||
<typeStatus box="[836,932,1123,1149]" pageId="6" pageNumber="499">paratype</typeStatus>
|
||
and the referred specimens are presented in
|
||
<tableCitation box="[151,239,1159,1185]" captionStart="TABLE 3" captionStartId="15.[144,232,466,492]" captionTargetId="graphics-31@15.[144,2030,534,1703]" captionTargetPageId="15" captionText="TABLE 3. Measurements and scale counts of Bronchocela spp. of the Nicobar Islands and B. criatatella" pageId="6" pageNumber="499">Table 3</tableCitation>
|
||
. Females slightly smaller than males, with a weekly developed nuchal crest and less legible blue shade around the eyes on the face.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="499" type="distribution">
|
||
<paragraph blockId="6.[151,1437,150,1329]" pageId="6" pageNumber="499">
|
||
<emphasis bold="true" box="[199,599,1230,1257]" pageId="6" pageNumber="499">Distribution and Natural history:</emphasis>
|
||
This species is endemic to Car Nicobar and possibly, the smaller southern island of Batti Malv. It is arboreal and diurnal in habit and is a fairly common species found in an array of habitats ranging from evergreen forests, secondary forests and plantations to home gardens (
|
||
<figureCitation box="[1091,1159,1303,1329]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[218,1369,210,1890]" captionTargetId="figure-21@14.[179,1408,181,1920]" captionTargetPageId="14" captionText="FIGURE 8. Map, showing the distribution of the four Bronchocela species in the Nicobar Archipelago (red triangles indicate type localities)." figureDoi="http://doi.org/10.5281/zenodo.7732157" httpUri="https://zenodo.org/record/7732157/files/figure.png" pageId="6" pageNumber="499">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |