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124 lines
10 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.352.6052" ID-GBIF-Dataset="99647020-9754-4dd3-8bf9-0db3657c1b5f" ID-PMC="PMC3837398" ID-Pensoft-Pub="1313-2970-352-51" ID-PubMed="24294092" ID-ZBK="FD8FE06F82C341D89C9512943B957BC6" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-352-51" ModsDocOrigin="ZooKeys 352" ModsDocTitle="A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus" checkinTime="1451246663146" checkinUser="pensoft" docAuthor="Sokolov, Igor M." docDate="2013" docId="FB21D1151E5F080EF381C9017CA8A7C5" docLanguage="en" docName="ZooKeys 352: 51-92" docOrigin="ZooKeys 352" docSource="http://dx.doi.org/10.3897/zookeys.352.6052" docTitle="Zapotecanillus oaxacanus Sokolov, 2013, sp. n." docType="treatment" docUuid="86234081-8CB0-45D0-861D-721A0C05174A" docUuidSource="ZooBank" docVersion="4" lastPageNumber="59" masterDocId="FFFCE9690B65E958B312FF9EFFA8B926" masterDocTitle="A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus" masterLastPageNumber="92" masterPageNumber="51" pageNumber="57" updateTime="1668156898856" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>352</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>51</mods:start>
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<mods:end>92</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.352.6052</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.352.6052</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-352-51</mods:identifier>
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<mods:identifier type="ZBK">FD8FE06F82C341D89C9512943B957BC6</mods:identifier>
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<mods:identifier type="ZooBank">FD8FE06F82C341D89C9512943B957BC6</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152049650" LSID="urn:lsid:zoobank.org:act:86234081-8CB0-45D0-861D-721A0C05174A" httpUri="http://treatment.plazi.org/id/FB21D1151E5F080EF381C9017CA8A7C5" lastPageId="8" lastPageNumber="59" pageId="6" pageNumber="57">
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<subSubSection pageId="6" pageNumber="57" type="nomenclature">
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<paragraph pageId="6" pageNumber="57">
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<taxonomicName LSID="http://zoobank.org/86234081-8CB0-45D0-861D-721A0C05174A" class="Insecta" family="Carabidae" genus="Zapotecanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zapotecanillus oaxacanus" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="oaxacanus">Zapotecanillus oaxacanus</taxonomicName>
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<taxonomicNameLabel pageId="6" pageNumber="57">sp. n.</taxonomicNameLabel>
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Figs 1, 3, 5, 7, 9-13, 15-18, 22, 24, 32, 36-38, 60, 68, 72, 77, 90-92, 94
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57" type="type material">
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<paragraph pageId="6" pageNumber="57">Type material.</paragraph>
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<paragraph lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57">
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HOLOTYPE, male, in NMNH, point-mounted, labeled: \MEXICO. Oaxaca 18.7mi S Valle Nacional 5200' 17 Aug.1973\ A.Newton Collector \ Loan from
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<pageBreakToken pageId="7" pageNumber="58" start="start">NMNH</pageBreakToken>
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2051867\. PARATYPES (8 ex., 4♂2♀ were dissected), labeled same as a holotype, except two specimens, which have an additional label: \
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<taxonomicName class="Insecta" family="Carabidae" genus="Geocharidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geocharidius" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Geocharidius</taxonomicName>
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n. sp. det. T.L.Erwin 76\ each, where italicized font means handwritten (deposited in CAS, NMNH).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="58" type="specific epithet">
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<paragraph pageId="7" pageNumber="58">Specific epithet.</paragraph>
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<paragraph pageId="7" pageNumber="58">The specific epithet is a Latinized adjective in the masculine form based on Oaxaca, the state of Mexico from which the new species is described.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="58" type="type locality">
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<paragraph pageId="7" pageNumber="58">Type locality.</paragraph>
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<paragraph pageId="7" pageNumber="58">Mexico, Oaxaca, 18.7mi S from Valle Nacional.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="58" type="recognition">
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<paragraph pageId="7" pageNumber="58">Recognition.</paragraph>
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<paragraph pageId="7" pageNumber="58">Adults of this new species can be distinguished easily from those of other species of the genus by the following combination of external characters: bicolored and robust appearance, comparatively small head and distinctly transverse pronotum.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58" type="description">
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<paragraph pageId="7" pageNumber="58">Description.</paragraph>
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<paragraph pageId="7" pageNumber="58">
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Size. Medium-sized for the genus (SBL range 1.30-1.36 mm, mean 1.33
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.049 mm, n=8).
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</paragraph>
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<paragraph pageId="7" pageNumber="58">
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Habitus. Body form (Fig. 24) moderately convex, slightly elongate (WE/SBL 0.41
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.09), head narrow for genus compared to pronotum (WH/WPm 0.69
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.015), pronotum wide compared to elytra (WPm/WE 0.81
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.017).
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</paragraph>
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<paragraph pageId="7" pageNumber="58">Color. Body bicolored: head and pronotum brunneorufous, elytra rufotestaceous, appendages testaceous.</paragraph>
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<paragraph pageId="7" pageNumber="58">Microsculpture. Disc of pronotum with well-developed microsculpture.</paragraph>
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<paragraph pageId="7" pageNumber="58">
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Prothorax. Pronotum (Fig. 32) relatively long (LP/LE 0.44
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.006) and markedly transverse (WPm/LP 1.33
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.022), with lateral margins straight and moderately constricted posteriorly (WPm/WPp 1.32
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.034). Basal margin straight. Contour of posterior angles nearly rectangular (100-110°) with 1-2 small denticles in front of the angles.
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</paragraph>
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<paragraph pageId="7" pageNumber="58">
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Elytra (Fig. 3). Convex, not depressed along suture, comparatively wide (WE/LE 0.73
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.013). Margins rounded, slightly divergent in basal half, evenly rounded to apex in apical half, maximal width of elytra at midpoint.
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</paragraph>
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<paragraph pageId="7" pageNumber="58">Legs. 1st male protarsomere only slightly dilated (Fig. 22).</paragraph>
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<paragraph pageId="8" pageNumber="59">
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<pageBreakToken pageId="8" pageNumber="59" start="start">Male</pageBreakToken>
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genitalia. Median lobe of aedeagus (Fig. 36), with short and transverse apex, broadly rounded at tip. Dorsal plate 1 long, with apical pointed attenuation of moderate length. Dorsal plate 2 joined to plate 1 at its middle ventrally, where it forms a distinct protuberance. Ventral sclerites elongate, with subparallel sides and obliquely stretched from dorsal plates towards ventral margin of median lobe. Right paramere short and moderately wide (Fig. 38). Left paramere with distinct apical constriction (Fig. 37). Ring sclerite with long handle-like extension, widely rounded apically (Fig. 60).
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</paragraph>
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<paragraph pageId="8" pageNumber="59">Female genitalia. Gonocoxite 2 rather long, with slightly curved blade and rounded apex (Fig. 68). Laterotergite with 7-8 setae. Spermatheca standard for genus (Fig. 72).</paragraph>
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<paragraph pageId="8" pageNumber="59">
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Geographical distribution. This species is known only from the type locality in the Sierra de
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<normalizedToken originalValue="Juárez">Juarez</normalizedToken>
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Range, a part of the Sierra Madre de Oaxaca, within the high course of the Rio de Valle Nacional (Figs 77 and 94, black quadrangles).
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</paragraph>
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<paragraph pageId="8" pageNumber="59">Way of life. Specimens of this species were collected at an altitude of 5200 feet (1600 m).</paragraph>
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<paragraph pageId="8" pageNumber="59">
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Relationships. The armature of internal sac of
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<taxonomicName class="Insecta" family="Carabidae" genus="Zapotecanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zapotecanillus oaxacanus" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="oaxacanus">Zapotecanillus oaxacanus</taxonomicName>
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males is nearly indistinguishable from that of
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<taxonomicName class="Insecta" family="Carabidae" genus="Zapotecanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zapotecanillus nanus" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="nanus">Zapotecanillus nanus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Carabidae" genus="Zapotecanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zapotecanillus ixtlanus" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="ixtlanus">Zapotecanillus ixtlanus</taxonomicName>
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males, described below, clearly suggesting both of them as closest relatives. The former species is sympatric with
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<taxonomicName class="Insecta" family="Carabidae" genus="Zapotecanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zapotecanillus oaxacanus" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="oaxacanus">Zapotecanillus oaxacanus</taxonomicName>
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and, based on the same label data, may also be syntopic (i.e., their members may occur together in the same habitat). See also Fig. 90 for cladistic affinities.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |