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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.80.12962" ID-PMC="PMC5543604" ID-Pensoft-Pub="1314-2003-80-33" ID-Pensoft-UUID="FFB61E2CFFBFFF8DFF93FFF3FFEFFFFA" ID-PubMed="28781556" ID-Zenodo-Dep="576387" ModsDocID="1314-2003-80-33" checkinTime="1493205412253" checkinUser="pensoft" docAuthor="Freeman, Craig C." docDate="2017" docId="A50293FCD9275AA1A89F222A26603A77" docLanguage="en" docName="PhytoKeys 80: 33-39" docOrigin="PhytoKeys 80" docPubDate="2017-04-25" docSource="http://dx.doi.org/10.3897/phytokeys.80.12962" docTitle="Penstemon linarioides subsp. var. var. coloradoensis C. C. Freeman 2017, stat. nov." docType="treatment" docVersion="3" id="FFB61E2CFFBFFF8DFF93FFF3FFEFFFFA" lastPageNumber="35" masterDocId="FFB61E2CFFBFFF8DFF93FFF3FFEFFFFA" masterDocTitle="Nomenclatural novelties and notes in Penstemon (Plantaginaceae)" masterLastPageNumber="39" masterPageNumber="33" pageNumber="34" updateTime="1668141960729" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Nomenclatural novelties and notes in Penstemon (Plantaginaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Freeman, Craig C.</mods:namePart>
<mods:affiliation>R. L. McGregor Herbarium, Division of Botany, Biodiversity Research Institute, University of Kansas, Lawrence, Kansas 66047 - 3729, USA</mods:affiliation>
<mods:nameIdentifier type="email">ccfree@ku.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-04-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>33</mods:start>
<mods:end>39</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.80.12962</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.80.12962</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-80-33</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFB61E2CFFBFFF8DFF93FFF3FFEFFFFA</mods:identifier>
<mods:identifier type="Zenodo-Dep">576387</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182226143" LSID="urn:lsid:plazi:treatment:A50293FCD9275AA1A89F222A26603A77" httpUri="http://treatment.plazi.org/id/A50293FCD9275AA1A89F222A26603A77" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
<subSubSection pageId="1" pageNumber="34" type="nomenclature">
<paragraph pageId="1" pageNumber="34">
<taxonomicName LSID="A50293FC-D927-5AA1-A89F-222A26603A77" authority="(A. Nelson) C. C. Freeman" authorityName="C. C. Freeman" authorityYear="2017" baseAuthorityName="A. Nelson" class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon linarioides subsp. var. var. coloradoensis" order="Lamiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="variety" species="linarioides" status="stat. nov." subSpecies="var." variety="coloradoensis">Penstemon linarioides var. coloradoensis (A.Nelson) C.C.Freeman</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="34">stat. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="34" type="basionym">
<paragraph pageId="1" pageNumber="34">Basionym.</paragraph>
<paragraph pageId="1" pageNumber="34">
<taxonomicName class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon coloradoensis" order="Lamiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="species" species="coloradoensis">
<emphasis italics="true" pageId="1" pageNumber="34">Penstemon coloradoensis</emphasis>
</taxonomicName>
A. Nelson, Bull. Torrey Bot. Club 26: 355. 1899;
<taxonomicName authorityName="C. C. Freeman" authorityYear="2017" baseAuthorityName="A. Nelson" class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon linarioides subsp. coloradoensis" order="Lamiales" pageId="1" pageNumber="34" phylum="Tracheophyta" rank="subSpecies" species="linarioides" subSpecies="coloradoensis">Penstemon linarioides subsp. coloradoensis</taxonomicName>
(A. Nelson) D.D. Keck.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34" type="materials_examined">
<paragraph pageId="1" pageNumber="34">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="1" pageNumber="34">
<materialsCitation accessionNumber="NDG45904, NDG45695" collectingDate="1898-06-23" collectorName="La Platas, C. F. Baker, F. S. Earle, S. M. Tracy" country="USA" location="COLORADO" specimenCount="1" stateProvince="COLORADO" typeStatus="lectotype">
<collectingCountry name="United States of America">USA</collectingCountry>
.
<collectingRegion country="United States of America" name="Colorado">COLORADO</collectingRegion>
: Mancos, common on sage plains and foothills throughout the
<collectorName>La Platas</collectorName>
,
<collectingDate value="1898-06-23">23 Jun 1898</collectingDate>
,
<collectorName>C.F. Baker</collectorName>
,
<collectorName>F.S. Earle</collectorName>
, and
<collectorName>S.M. Tracy</collectorName>
70 (
<typeStatus>lectotype</typeStatus>
: RM, RM0004187 [JSTOR image]; isolectotypes: CAS, CAS0006620 [CAS website image], F, F0072589F [JSTOR image], GH, GH00091215 [GH website image], K, K000979702 [K website image], KSC, MICH, MICH1108038 [MICH website image], MO, MO-100230852, NDG,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/NDG45904">NDG45904</accessionNumber>
[JSTOR image], NEB, NEB-v-0000619 [JSTOR image], NY, NY00091049 [NY website image], OS, OS0000264 [JSTOR image], RM, RM0004188 [JSTOR image],
<collectingCountry isEnumeration="true">RSA, RSA</collectingCountry>
0006119 [JSTOR image],
<collectingCountry name="South Africa">RSA</collectingCountry>
0006120 [JSTOR image],
<collectingCountry isEnumeration="true">US, US</collectingCountry>
00122246 [
<collectingCountry name="United States of America">US</collectingCountry>
website image]; probable isolectotype: NDG,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/NDG45695">NDG45695</accessionNumber>
[JSTOR image])
</materialsCitation>
.
</paragraph>
<paragraph pageId="2" pageNumber="35">
<pageBreakToken pageId="2" pageNumber="35" start="start">In</pageBreakToken>
describing
<taxonomicName class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon coloradoensis" order="Lamiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="coloradoensis">
<emphasis italics="true" pageId="2" pageNumber="35">Penstemon coloradoensis</emphasis>
</taxonomicName>
,
<bibRefCitation DOI="https://doi.org/10.2307/2478041" author="Nelson, A" journalOrPublisher="Club" pageId="5" pageNumber="38" pagination="350 - 358" refId="B4" refString="Nelson, A, 1899. New plants from Wyoming.-IX. Bull. Torrey Bot. Club 26: 350 - 358, DOI: https://doi.org/10.2307/2478041" title="New plants from Wyoming. - IX. Bull. Torrey Bot." url="https://doi.org/10.2307/2478041" volume="26" year="1899">Nelson (1899)</bibRefCitation>
stated &quot;
<emphasis italics="true" pageId="2" pageNumber="35">
I have before me specimens from two collections made near Mancos, Colo., by Messrs. Baker, Earle, and Tracy, 1898 and distributed as P.
<taxonomicName class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="caespitosus" order="Lamiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="caespitosus">caespitosus</taxonomicName>
Nutt. Also from two collections by Professor Crandall, from Hotchkiss, Colo., 1892, and from Durango, 1898, both distributed as P.
<taxonomicName class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="linarioides" order="Lamiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="linarioides">linarioides</taxonomicName>
Sileri Gray.
</emphasis>
&quot;
<bibRefCitation DOI="https://doi.org/10.2307/2478041" author="Nelson, A" journalOrPublisher="Club" pageId="5" pageNumber="38" pagination="350 - 358" refId="B4" refString="Nelson, A, 1899. New plants from Wyoming.-IX. Bull. Torrey Bot. Club 26: 350 - 358, DOI: https://doi.org/10.2307/2478041" title="New plants from Wyoming. - IX. Bull. Torrey Bot." url="https://doi.org/10.2307/2478041" volume="26" year="1899">Nelson (1899)</bibRefCitation>
did not designate a
<typeStatus>holotype</typeStatus>
; the collections he cited are
<typeStatus>syntypes</typeStatus>
.
<bibRefCitation DOI="https://doi.org/10.2307/2481121" author="Keck, DD" journalOrPublisher="Club" pageId="5" pageNumber="38" pagination="357 - 381" refId="B3" refString="Keck, DD, 1937. Studies in Penstemon IV. The section Ericopsis. Bull. Torrey Bot. Club 64: 357 - 381, DOI: https://doi.org/10.2307/2481121" title="Studies in Penstemon IV. The section Ericopsis. Bull. Torrey Bot." url="https://doi.org/10.2307/2481121" volume="64" year="1937">Keck (1937)</bibRefCitation>
designated C.F. Baker, F.S. Earle, and S.M. Tracy 70 (RM) as the
<typeStatus>lectotype</typeStatus>
, of which there currently are two sheets, suggesting a second-step lectotypification might be necessary. Accession labels on the two sheets indicate they were received from the herbaria of F.S. Earle in 1899 (RM0004187; accession number 13703) and George E. Osterhout in 1938 (RM0004188; accession number 165282), subsequent to
<normalizedToken originalValue="Kecks">Keck's</normalizedToken>
<typeStatus>lectotype</typeStatus>
designation. RM0004187 bears annotations by A. Nelson (
<taxonomicName class="Magnoliopsida" family="Plantaginaceae" genus="Penstemon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Penstemon coloradoensis" order="Lamiales" pageId="2" pageNumber="35" phylum="Tracheophyta" rank="species" species="coloradoensis">
<emphasis italics="true" pageId="2" pageNumber="35">Penstemon coloradoensis</emphasis>
</taxonomicName>
A. N. n. sp., without date), F.W. Pennell (
<taxonomicName lsidName="P. coloradoensis" pageId="2" pageNumber="35" rank="species" species="coloradoensis">
<emphasis italics="true" pageId="2" pageNumber="35">P. coloradoensis</emphasis>
</taxonomicName>
, type, 1916), and D.D. Keck (
<taxonomicName lsidName="P. linarioides subsp. coloradoensis" pageId="2" pageNumber="35" rank="subSpecies" species="linarioides" subSpecies="coloradoensis">P. linarioides A. Gray subsp. coloradoensis</taxonomicName>
(A. Nelson) D.D. Keck, type, 1936). RM0004188, the later accession, was not annotated by Nelson, Pennell, or Keck. Whether Nelson had another sheet at RM that was lost, destroyed, or sent elsewhere, or had access to a sheet from some other herbarium, is not known. Of the two sheets currently at RM, only RM0004187 appears to have been there when Nelson described
<taxonomicName lsidName="P. coloradoensis" pageId="2" pageNumber="35" rank="species" species="coloradoensis">
<emphasis italics="true" pageId="2" pageNumber="35">P. coloradoensis</emphasis>
</taxonomicName>
and later seen by Keck when he lectotypified the name. RM0004187 is taken as the
<typeStatus>lectotype</typeStatus>
,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/RM004188">RM004188</accessionNumber>
is taken as an isolectotype, and no second-step lectotypification is needed. As an aside, the sheet at KSC is
<taxonomicName authority="A. Nelson var. crandallii" lsidName="P. crandallii subsp. var. var. crandallii" pageId="2" pageNumber="35" rank="variety" species="crandallii" subSpecies="var." variety="crandallii">P. crandallii A. Nelson var. crandallii</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>