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<document id="0DA365278424A1A9CA08CE1EFDE3546F" ID-CLB-Dataset="300882" ID-DOI="10.37828/em.2022.57.11" ID-GBIF-Dataset="ba0d8cf8-c053-4db7-a222-89cab29291cd" ID-ISSN="2336-9744" ID-Zenodo-Dep="13240772" ID-ZooBank="urn:lsid:zoobank.org:pub:87364D2D-9F82-457A-945C-47BD8F1094CD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722959413189" checkinUser="felipe" docAuthor="Spitsyn, Vitaly M., Spitsyna, Elizaveta A., Kondakov, Alexander V., Tomilova, Alena A., Burchalovskaia, Polina D. &amp; Bolotov, Ivan N." docDate="2022" docId="8C3787F1FFBA6E0ADE8CF984FE10F864" docLanguage="en" docName="EcolMontenegrina.57.118-126.pdf" docOrigin="Ecologica Montenegrina 57" docSource="http://dx.doi.org/10.37828/em.2022.57.11" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Laothoe amurensis subsp. zolotuhini Spitsyn, Bolotov &amp; Spitsyna 2022, ssp. nov." docType="treatment" docVersion="3" lastPageNumber="123" masterDocId="700EFF89FFBE6E0FDE1BFFDBFFB9FFEB" masterDocTitle="Laothoe amurensis zolotuhini ssp. nov. from Kunashir Island, Russia (Lepidoptera: Sphingidae)" masterLastPageNumber="126" masterPageNumber="118" pageNumber="122" updateTime="1722963720222" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="26756DB81953AB596992B1F3862EFAE5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="AE962386C4C1D9964F8A92BD5225B8CB">Laothoe amurensis zolotuhini ssp. nov. from Kunashir Island, Russia (Lepidoptera: Sphingidae)</mods:title>
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<mods:namePart id="B2862ACC3AC1AB680B96C6C8D8294841">Spitsyn, Vitaly M.</mods:namePart>
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<mods:namePart id="1D310F583AF1CD1423825552E6D6D18A">Spitsyna, Elizaveta A.</mods:namePart>
</mods:name>
<mods:name id="06153D1D5EBB6398F39949752A29EA35" type="personal">
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<mods:namePart id="ED14292B53574603C4AD7100BEB270BA">Kondakov, Alexander V.</mods:namePart>
</mods:name>
<mods:name id="127FC80D180DE24B5E250569FB9F4CBE" type="personal">
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<mods:roleTerm id="FDEB92F9618AE1C85DAF4C1E8BB793B7">Author</mods:roleTerm>
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<mods:namePart id="30D5C81EC7BDB754E81FED2069CB3118">Tomilova, Alena A.</mods:namePart>
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<mods:roleTerm id="4754774F9CB70EE65FA010672A0B0539">Author</mods:roleTerm>
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<mods:namePart id="79D4280A439425161C430A7069B4EFFE">Burchalovskaia, Polina D.</mods:namePart>
</mods:name>
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<mods:namePart id="FFBDC1F624DAE4800F9270E98E89B4B0">Bolotov, Ivan N.</mods:namePart>
</mods:name>
<mods:typeOfResource id="095697497FAB9DA38036090EF2C3A67D">text</mods:typeOfResource>
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<mods:title id="F51B7A40DFF72F2E5C4BD68C76EFFD36">Ecologica Montenegrina</mods:title>
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<mods:part id="D316415FB1B06D53233726FC4B6A9148">
<mods:date id="97E823055302E3756B3319E45D372383">2022</mods:date>
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<mods:number id="8CEA2BC3FCA3DB57151DD327391A8036">2022-10-14</mods:number>
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<mods:number id="6E269991B00F487ED3918DD69B20661D">57</mods:number>
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<treatment id="8C3787F1FFBA6E0ADE8CF984FE10F864" LSID="urn:lsid:plazi:treatment:8C3787F1FFBA6E0ADE8CF984FE10F864" httpUri="http://treatment.plazi.org/id/8C3787F1FFBA6E0ADE8CF984FE10F864" lastPageId="5" lastPageNumber="123" pageId="4" pageNumber="122">
<subSubSection id="4C84656CFFBA6E0BDE8CF984FC60F992" box="[151,985,1631,1657]" pageId="4" pageNumber="122" type="nomenclature">
<paragraph id="042136E7FFBA6E0BDE8CF984FC60F992" blockId="4.[151,985,1631,1690]" box="[151,985,1631,1657]" pageId="4" pageNumber="122">
<heading id="5F69818BFFBA6E0BDE8CF984FC60F992" box="[151,985,1631,1657]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="122" reason="2">
<emphasis id="36EAEAF5FFBA6E0BDE8CF984FC60F992" bold="true" box="[151,985,1631,1657]" pageId="4" pageNumber="122">
<taxonomicName id="C39E4D64FFBA6E0BDE8CF984FCD1F992" authority="Spitsyn, Bolotov &amp; Spitsyna" authorityName="Spitsyn, Bolotov &amp; Spitsyna" authorityYear="2022" box="[151,872,1631,1657]" class="Insecta" family="Sphingidae" genus="Laothoe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="122" phylum="Arthropoda" rank="subSpecies" species="amurensis" status="subsp. nov." subSpecies="zolotuhini">
<emphasis id="36EAEAF5FFBA6E0BDE8CF984FDB8F992" bold="true" box="[151,513,1631,1657]" italics="true" pageId="4" pageNumber="122">Laothoe amurensis zolotuhini</emphasis>
Spitsyn, Bolotov &amp; Spitsyna
</taxonomicName>
<taxonomicNameLabel id="2DD9578EFFBA6E0BDD75F984FC60F992" box="[878,985,1631,1657]" pageId="4" pageNumber="122" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4C84656CFFBA6E0BDE8CF9A4FEF1F971" box="[151,328,1663,1690]" pageId="4" pageNumber="122" type="description">
<paragraph id="042136E7FFBA6E0BDE8CF9A4FEF1F971" blockId="4.[151,985,1631,1690]" box="[151,328,1663,1690]" pageId="4" pageNumber="122">
<figureCitation id="9CA52A62FFBA6E0BDE8CF9A4FF63F971" box="[151,218,1663,1690]" captionStart="Figure 1" captionStartId="5.[151,226,1236,1260]" captionTargetBox="[174,1389,211,1199]" captionTargetId="figure-19@5.[151,1424,189,1218]" captionTargetPageId="5" captionText="Figure 1. Laothoe amurensis ssp.: A-B) L. a. amurensis from Primorsky Krai: (A) male; (B) female. (C-F) L. a. zolotuhini Spitsyn, Bolotov &amp; Spitsyna ssp. nov. from Kunashir Island: (C, E) holotype male RMBH Sph0928: (C) upperside; (E) underside; (D, F) paratype female RMBH Sph0927: (D) upperside; (F) underside. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13240774" httpUri="https://zenodo.org/record/13240774/files/figure.png" pageId="4" pageNumber="122">Figs 1</figureCitation>
C-F −2B
</paragraph>
</subSubSection>
<subSubSection id="4C84656CFFBA6E0BDE8CF918FB58F8CA" pageId="4" pageNumber="122" type="materials_examined">
<paragraph id="042136E7FFBA6E0BDE8CF918FB58F8CA" blockId="4.[151,1436,1731,1825]" pageId="4" pageNumber="122">
<emphasis id="36EAEAF5FFBA6E0BDE8CF918FEE9F936" bold="true" box="[151,336,1731,1757]" pageId="4" pageNumber="122">Type material.</emphasis>
<materialsCitation id="B4F63CBAFFBA6E0BDF43F918FAA4F8EB" collectingDate="2021-07-04" collectingDateMax="2021-07-06" collectingDateMin="2021-07-04" collectionCode="RMBH, V" collectorName="E. Spitsyna &amp; Spitsyn" country="Russia" county="Tretyakovo village" latitude="43.986946" location="Sph" longLatPrecision="19" longitude="145.65332" municipality="Kunashir Island" pageId="4" pageNumber="122" specimenCount="1" typeStatus="holotype">
<typeStatus id="DB258845FFBA6E0BDF43F918FE7CF935" box="[344,453,1731,1758]" pageId="4" pageNumber="122" type="holotype">Holotype</typeStatus>
male
<collectionCode id="628FAE22FFBA6E0BDC15F91FFDDDF935" box="[526,612,1732,1758]" pageId="4" pageNumber="122">RMBH</collectionCode>
<location id="0141603CFFBA6E0BDC76F918FD24F935" LSID="urn:lsid:plazi:treatment:8C3787F1FFBA6E0ADE8CF984FE10F864:0141603CFFBA6E0BDC76F918FD24F935" box="[621,669,1731,1758]" country="Russia" county="Tretyakovo village" latitude="43.986946" longLatPrecision="19" longitude="145.65332" municipality="Kunashir Island" name="Sph" pageId="4" pageNumber="122">Sph</location>
0928:
<collectingCountry id="7C897677FFBA6E0BDCFDF918FCE8F935" box="[742,849,1731,1758]" name="Russia" pageId="4" pageNumber="122">RUSSIA</collectingCountry>
,
<collectingMunicipality id="E445AC9DFFBA6E0BDD46F918FBA5F935" box="[861,1052,1731,1758]" pageId="4" pageNumber="122">Kunashir Island</collectingMunicipality>
,
<collectingCounty id="ED404E6BFFBA6E0BDA33F918FAB3F935" box="[1064,1290,1731,1758]" pageId="4" pageNumber="122">Tretyakovo village</collectingCounty>
, coniferousdeciduous forest,
<geoCoordinate id="61AA5020FFBA6E0BDF68F93EFE46F8EB" box="[371,511,1765,1792]" degrees="43" direction="north" minutes="59" orientation="latitude" pageId="4" pageNumber="122" precision="15" seconds="13" value="43.986946">43°59'13&quot;N</geoCoordinate>
,
<geoCoordinate id="61AA5020FFBA6E0BDC09F93EFD13F8EB" box="[530,682,1765,1792]" degrees="145" direction="east" minutes="39" orientation="longitude" pageId="4" pageNumber="122" precision="15" seconds="12" value="145.65332">145°39'12&quot;E</geoCoordinate>
,
<date id="70201027FFBA6E0BDCA7F93EFC2BF8EB" box="[700,914,1765,1792]" pageId="4" pageNumber="122" value="2021-07-04" valueMax="2021-07-06" valueMin="2021-07-04">
<collectingDate id="6064E9CFFFBA6E0BDCA7F93EFC2BF8EB" box="[700,914,1765,1792]" pageId="4" pageNumber="122" value="2021-07-04" valueMax="2021-07-06" valueMin="2021-07-04">04.vii-06.vii.2021</collectingDate>
</date>
,
<collectorName id="A96B5331FFBA6E0BDDBEF93DFB88F8EB" box="[933,1073,1765,1792]" pageId="4" pageNumber="122">E. Spitsyna</collectorName>
&amp;
<collectionCode id="628FAE22FFBA6E0BDA79F93DFBC2F8EB" box="[1122,1147,1766,1792]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="122" type="Museum">V</collectionCode>
.
<collectorName id="A96B5331FFBA6E0BDA97F93EFB5DF8EB" box="[1164,1252,1765,1792]" pageId="4" pageNumber="122">Spitsyn</collectorName>
leg.
</materialsCitation>
<materialsCitation id="B4F63CBAFFBA6E0BDB31F93DFB62F8CA" collectionCode="RMBH" country="Russia" pageId="4" pageNumber="122" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="DB258845FFBA6E0BDB31F93DFA25F8EB" box="[1322,1436,1766,1792]" pageId="4" pageNumber="122" type="paratype">Paratypes</typeStatus>
<collectionCode id="628FAE22FFBA6E0BDE8CF8DCFF54F8CA" box="[151,237,1799,1825]" pageId="4" pageNumber="122">RMBH</collectionCode>
Sph0927, Sph0996−Sph0998: the type locality, the same date and collectors,
<specimenCount id="1298FD6EFFBA6E0BDA99F8DDFB10F8CA" box="[1154,1193,1798,1825]" count="3" pageId="4" pageNumber="122" type="male">3♂</specimenCount>
,
<specimenCount id="1298FD6EFFBA6E0BDAAEF8DDFB62F8CA" box="[1205,1243,1798,1825]" count="1" pageId="4" pageNumber="122" type="female">1♀</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4C84656CFFBA6E0BDE8CF891FEA9F86C" pageId="4" pageNumber="122" type="diagnosis">
<paragraph id="042136E7FFBA6E0BDE8CF891FEA9F86C" blockId="4.[151,1435,1866,1927]" pageId="4" pageNumber="122">
<emphasis id="36EAEAF5FFBA6E0BDE8CF891FEAEF88F" bold="true" box="[151,279,1866,1892]" pageId="4" pageNumber="122">Diagnosis.</emphasis>
The new subspecies differs from the nominate one by having a strongly toothed outer margin of the wings.
</paragraph>
</subSubSection>
<caption id="50E1666FFFBB6E0ADE8CFB0FFAC3FAC2" ID-DOI="http://doi.org/10.5281/zenodo.13240774" ID-Zenodo-Dep="13240774" httpUri="https://zenodo.org/record/13240774/files/figure.png" pageId="5" pageNumber="123" startId="5.[151,226,1236,1260]" targetBox="[174,1389,211,1199]" targetPageId="5" targetType="figure">
<paragraph id="042136E7FFBB6E0ADE8CFB0FFAC3FAC2" blockId="5.[151,1437,1236,1322]" pageId="5" pageNumber="123">
<emphasis id="36EAEAF5FFBB6E0ADE8CFB0FFEB8FB07" bold="true" box="[151,257,1236,1260]" pageId="5" pageNumber="123">Figure 1.</emphasis>
<taxonomicName id="C39E4D64FFBB6E0ADF16FB0FFE65FB00" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[269,476,1236,1260]" class="Insecta" family="Sphingidae" genus="Laothoe" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="123" phylum="Arthropoda" rank="species" species="amurensis">
<emphasis id="36EAEAF5FFBB6E0ADF16FB0FFE65FB00" box="[269,476,1236,1260]" italics="true" pageId="5" pageNumber="123">Laothoe amurensis</emphasis>
</taxonomicName>
ssp.:
<emphasis id="36EAEAF5FFBB6E0ADC39FB0FFDE9FB07" bold="true" box="[546,592,1236,1260]" pageId="5" pageNumber="123">A-B</emphasis>
)
<emphasis id="36EAEAF5FFBB6E0ADC7FFB0EFCA9FB00" box="[612,784,1236,1260]" italics="true" pageId="5" pageNumber="123">
L. a.
<taxonomicName id="C39E4D64FFBB6E0ADCB8FB0FFCA9FB00" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[675,784,1236,1259]" class="Insecta" family="Sphingidae" genus="Laothoe" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="123" phylum="Arthropoda" rank="species" species="amurensis">amurensis</taxonomicName>
</emphasis>
from Primorsky Krai: (
<emphasis id="36EAEAF5FFBB6E0ADA05FB0FFB88FB07" bold="true" box="[1054,1073,1236,1260]" pageId="5" pageNumber="123">A</emphasis>
) male; (
<emphasis id="36EAEAF5FFBB6E0ADA8EFB0FFB1EFB07" bold="true" box="[1173,1191,1236,1260]" pageId="5" pageNumber="123">B</emphasis>
) female. (
<emphasis id="36EAEAF5FFBB6E0ADB33FB0FFAEDFB07" bold="true" box="[1320,1364,1236,1260]" pageId="5" pageNumber="123">C-F</emphasis>
)
<emphasis id="36EAEAF5FFBB6E0ADB73FB0EFEBBFAE1" italics="true" pageId="5" pageNumber="123">L. a. zolotuhini</emphasis>
Spitsyn, Bolotov &amp; Spitsyna
<emphasis id="36EAEAF5FFBB6E0ADC54FB28FD0DFAE0" bold="true" box="[591,692,1267,1291]" pageId="5" pageNumber="123">
<taxonomicNameLabel id="2DD9578EFFBB6E0ADC54FB28FD0DFAE0" box="[591,692,1267,1291]" pageId="5" pageNumber="123" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
from Kunashir Island: (
<emphasis id="36EAEAF5FFBB6E0ADDD9FB28FC41FAE0" bold="true" box="[962,1016,1267,1291]" pageId="5" pageNumber="123">C, E</emphasis>
) holotype male RMBH Sph0928: (
<emphasis id="36EAEAF5FFBB6E0ADB9BFB28FA2AFAE0" bold="true" box="[1408,1427,1267,1291]" pageId="5" pageNumber="123">C</emphasis>
) upperside; (
<emphasis id="36EAEAF5FFBB6E0ADF0DFACAFE91FAC2" bold="true" box="[278,296,1297,1321]" pageId="5" pageNumber="123">E</emphasis>
) underside; (
<emphasis id="36EAEAF5FFBB6E0ADFAEFACAFE5FFAC2" bold="true" box="[437,486,1297,1321]" pageId="5" pageNumber="123">D, F</emphasis>
) paratype female RMBH Sph0927: (
<emphasis id="36EAEAF5FFBB6E0ADD68FACAFC3FFAC2" bold="true" box="[883,902,1297,1321]" pageId="5" pageNumber="123">D</emphasis>
) upperside; (
<emphasis id="36EAEAF5FFBB6E0ADA0EFACAFB9CFAC2" bold="true" box="[1045,1061,1297,1321]" pageId="5" pageNumber="123">F</emphasis>
) underside. Scale bar = 10 mm.
</paragraph>
</caption>
<subSubSection id="4C84656CFFBB6E0ADE8CFAA1FE10F864" pageId="5" pageNumber="123" type="description">
<paragraph id="042136E7FFBB6E0ADE8CFAA1FE10F864" blockId="5.[151,1437,1402,1935]" pageId="5" pageNumber="123">
<emphasis id="36EAEAF5FFBB6E0ADE8CFAA1FE89FA7F" bold="true" box="[151,304,1402,1428]" pageId="5" pageNumber="123">Description.</emphasis>
<emphasis id="36EAEAF5FFBB6E0ADF22FAA1FDAFFA7E" box="[313,534,1402,1429]" italics="true" pageId="5" pageNumber="123">Male morphology:</emphasis>
Wingspan
<quantity id="C3669B02FFBB6E0ADCBBFAA0FC9AFA7E" box="[672,803,1402,1429]" metricMagnitude="-2" metricUnit="m" metricValue="8.05" metricValueMax="8.4" metricValueMin="7.7" pageId="5" pageNumber="123" unit="mm" value="80.5" valueMax="84.0" valueMin="77.0">77–84 mm</quantity>
, forewing length
<quantity id="C3669B02FFBB6E0ADDE3FAA1FBC1FA7E" box="[1016,1144,1402,1429]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="5" pageNumber="123" unit="mm" value="41.0" valueMax="42.0" valueMin="40.0">40–42 mm</quantity>
(
<emphasis id="36EAEAF5FFBB6E0ADA90FAA0FB26FA7F" box="[1163,1183,1403,1428]" italics="true" pageId="5" pageNumber="123">N</emphasis>
= 4). Eye dark grey. Antenna brownish orange on the inner side, creamy on the outer side. Labial palpus dark grey, its length equal to the eye’s diameter. Head, thorax, patagium, and tegula dark grey. Legs grey, outer side of tibia and tarsus creamy grey. Forewing upperside grey, with four wavy lines running from costal margin to anal margin; antemedial stripe dark grey. Outer margin toothed. Marginal area from R4 to Cu1 dark grey. Forewing underside light grey with wavy postmedial and submarginal lines. Marginal area from R4 to Cu1 dark grey. Hindwing upperside grey with two wavy postmedial lines; marginal area from Rs to Cu1 dark grey. Outer margin with a strong ledge in the area of Rs and M1 veins. Hindwing underside repeats the pattern of the upperside; marginal area from Sc+R1 to Cu1 dark grey. Abdomen grey.
<emphasis id="36EAEAF5FFBB6E0ADAFAF953FA24F948" box="[1249,1437,1672,1699]" italics="true" pageId="5" pageNumber="123">Male genitalia:</emphasis>
Tegumen broad; uncus and gnathos form a typical “bird- beak” structure; valva slightly elongated, rounded apically. Saccular process of valva typical for the genus. Aedeagus long, slightly curved. Vesica long, with two fields of spines.
<emphasis id="36EAEAF5FFBB6E0ADF96F936FD3DF8E3" box="[397,644,1773,1800]" italics="true" pageId="5" pageNumber="123">Female morphology:</emphasis>
Wingspan
<quantity id="C3669B02FFBB6E0ADD10F936FCD9F8E3" box="[779,864,1773,1800]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="5" pageNumber="123" unit="mm" value="91.0">91 mm</quantity>
, forewing length
<quantity id="C3669B02FFBB6E0ADA34F936FB3BF8E3" box="[1071,1154,1773,1800]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="123" unit="mm" value="46.0">46 mm</quantity>
(
<emphasis id="36EAEAF5FFBB6E0ADA8FF935FB11F8EC" box="[1172,1192,1774,1799]" italics="true" pageId="5" pageNumber="123">N</emphasis>
= 1). Eye dark grey. Antenna brownish orange on the inner side, creamy on the outer side. Labial palpus dark grey, its length equal to the eye’s diameter. Head, thorax, patagium, and tegula dark grey. Legs grey, outer side of tibia and tarsus creamy grey. Generally, the female’s wings coloration is similar to that of the male but it is noticeably lighter. Abdomen grey.
</paragraph>
</subSubSection>
</treatment>
</document>
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