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<document ID-DOI="http://doi.org/10.5281/zenodo.4754875" ID-GBIF-Dataset="a60c41ef-ab6a-406c-af1d-a0cf0face281" ID-GBIF-Taxon="182067539" ID-ISSN="1854-0392" ID-Zenodo-Dep="4754875" approvalRequired="8" approvalRequired_for_textStreams="4" approvalRequired_for_treatments="4" checkinTime="1620844466829" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Sivec, Ignac" docDate="2008" docId="747887DCFFAC9E45F84EFBCBF244F995" docLanguage="en" docName="Illiesia.4.17.161-175.pdf" docOrigin="Illiesia 4 (17)" docStyle="DocumentStyle:1DEF1196F206C1C297CF0540A5C6768F.1:Illesia.2005-2008.journal_article" docStyleId="1DEF1196F206C1C297CF0540A5C6768F" docStyleName="Illesia.2005-2008.journal_article" docStyleVersion="1" docTitle="Kiotina quadrituberculata Wu 1948" docType="treatment" docVersion="6" lastPageNumber="167" masterDocId="8841FFA4FFAA9E43F962FFF1F660FFE9" masterDocTitle="Systematic Notes On Kiotina Klapálek And Hemacroneuria Enderlein (Plecoptera: Perlidae), With Description Of Four New Species" masterLastPageNumber="175" masterPageNumber="161" pageId="6" pageNumber="167" updateTime="1632007339879" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematic Notes On Kiotina Klapálek And Hemacroneuria Enderlein (Plecoptera: Perlidae), With Description Of Four New Species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>and Ignac Sivec &amp; Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sivec, Ignac</mods:namePart>
<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>17</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>161</mods:start>
<mods:end>175</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4754771</mods:identifier>
<mods:identifier type="GBIF-Dataset">a60c41ef-ab6a-406c-af1d-a0cf0face281</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4754771</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4754875" ID-GBIF-Taxon="182067539" ID-Zenodo-Dep="4754875" LSID="urn:lsid:plazi:treatment:747887DCFFAC9E45F84EFBCBF244F995" httpUri="http://treatment.plazi.org/id/747887DCFFAC9E45F84EFBCBF244F995" lastPageNumber="167" pageId="6" pageNumber="167">
<subSubSection box="[300,659,1082,1107]" pageId="6" pageNumber="167" type="nomenclature">
<paragraph blockId="6.[300,659,1082,1107]" box="[300,659,1082,1107]" pageId="6" pageNumber="167">
<heading bold="true" box="[300,659,1082,1107]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="167" reason="2">
<taxonomicName authority="Wu" authorityName="Wu" authorityYear="1948" box="[300,659,1082,1107]" class="Insecta" family="Perlidae" genus="Kiotina" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="quadrituberculata">
<emphasis bold="true" box="[300,659,1082,1107]" pageId="6" pageNumber="167">
<emphasis bold="true" box="[300,604,1082,1106]" italics="true" pageId="6" pageNumber="167">Kiotina quadrituberculata</emphasis>
Wu
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[409,550,1117,1141]" pageId="6" pageNumber="167" type="description">
<paragraph blockId="6.[409,550,1117,1141]" box="[409,550,1117,1141]" pageId="6" pageNumber="167">
(
<figureCitation box="[417,543,1117,1141]" captionStart="Figs" captionStartId="6.[184,234,979,1003]" captionTargetBox="[263,1348,422,893]" captionTargetId="figure-388@6.[252,1354,415,902]" captionTargetPageId="6" captionText="Figs. 20-22. Kiotina quadrituberculata structures. 20. Male terminalia, dorsal, 21. Male paraproct, caudoventral aspect, 22. Male sternum 9." figureDoi="http://doi.org/10.5281/zenodo.4754785" httpUri="https://zenodo.org/record/4754785/files/figure.png" pageId="6" pageNumber="167">Figs. 2022</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="167" type="reference_group">
<paragraph blockId="6.[221,775,1185,1270]" box="[221,632,1186,1207]" pageId="6" pageNumber="167">
<treatmentCitationGroup box="[221,632,1186,1207]" pageId="6" pageNumber="167">
<treatmentCitation author="Wu, C. F." box="[221,628,1186,1207]" page="148" pageId="6" pageNumber="167" year="1948">
<taxonomicName authority="Wu, 1948: 148" authorityName="Wu" authorityPageNumber="148" authorityYear="1948" box="[221,628,1186,1207]" class="Insecta" family="Perlidae" genus="Kiotina" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="167" phylum="Arthropoda" rank="species" species="quadrituberculata">
<emphasis box="[221,471,1186,1207]" italics="true" pageId="6" pageNumber="167">Kiotina quadrituberculata</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111197307" box="[483,628,1186,1207]" country="China" county="Fukien" location="Yenching University Collection" municipality="Shao-wu" pageId="6" pageNumber="167" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="holotype">
<bibRefCitation author="Wu, C. F." box="[483,628,1186,1207]" pageId="6" pageNumber="167" pagination="145 - 151" refId="ref6233" refString="Wu, C. F. 1948. Fifth supplement to the stoneflies of China (Order Plecoptera). Peking Natural History Bulletin, 17: 145 - 151." type="journal article" year="1948">Wu, 1948:148</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="6.[221,775,1185,1270]" pageId="6" pageNumber="167">
<materialsCitation ID-GBIF-Occurrence="3111847301" country="China" county="Fukien" location="Tachu-luan" municipality="Shao-wu" pageId="6" pageNumber="167" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="holotype">
<typeStatus box="[643,741,1186,1208]" pageId="6" pageNumber="167">Holotype</typeStatus>
<specimenCount box="[752,770,1185,1207]" pageId="6" pageNumber="167" type="male"></specimenCount>
, presumed lost (Yenching University Collection).
<location LSID="urn:lsid:plazi:treatment:747887DCFFAC9E45F84EFBCBF244F995:F90E6011FFAC9E45FB87FB30F75BFB1F" country="China" county="Fukien" municipality="Shao-wu" name="Tachu-luan" pageId="6" pageNumber="167" stateProvince="Fujian">Tachu-luan</location>
,
<collectingMunicipality box="[324,417,1248,1270]" pageId="6" pageNumber="167">Shao-wu</collectingMunicipality>
,
<collectingCounty box="[428,500,1248,1270]" pageId="6" pageNumber="167">Fukien</collectingCounty>
(
<collectingRegion box="[512,581,1248,1270]" country="China" name="Fujian" pageId="6" pageNumber="167">Fujian</collectingRegion>
),
<collectingCountry box="[597,659,1248,1270]" name="China" pageId="6" pageNumber="167">China</collectingCountry>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="167" type="materials_examined">
<paragraph blockId="6.[184,776,1315,1961]" pageId="6" pageNumber="167">
<emphasis bold="true" box="[184,410,1315,1339]" pageId="6" pageNumber="167">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111197328" collectingDate="1948-05-01" collectingDateMax="1948-05-05" collectingDateMin="1948-05-01" collectionCode="USNM" collectorName="J. Fu" country="China" county="Fukien" elevation="1372" location="Kuatun Mountain" municipality="Province" pageId="6" pageNumber="167" specimenCount="2" specimenCount-male="2" stateProvince="Fujian">
<collectingCountry box="[414,485,1315,1339]" name="China" pageId="6" pageNumber="167">China</collectingCountry>
:
<collectingCounty box="[494,573,1315,1339]" pageId="6" pageNumber="167">Fukien</collectingCounty>
(
<collectingRegion box="[584,657,1315,1339]" country="China" name="Fujian" pageId="6" pageNumber="167">Fujian</collectingRegion>
)
<collectingMunicipality box="[669,771,1315,1339]" pageId="6" pageNumber="167">Province</collectingMunicipality>
, Ta-chu-laun, foot of
<location LSID="urn:lsid:plazi:treatment:747887DCFFAC9E45F84EFBCBF244F995:F90E6011FFAC9E45F8FCFAB7F40DFAB4" box="[414,621,1349,1373]" country="China" county="Fukien" municipality="Province" name="Kuatun Mountain" pageId="6" pageNumber="167" stateProvince="Fujian">Kuatun Mountain</location>
,
<quantity box="[631,731,1349,1373]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" pageId="6" pageNumber="167" unit="ft" value="4500.0">
<elevation box="[631,731,1349,1373]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" pageId="6" pageNumber="167" unit="ft" value="4500.0">4500 feet</elevation>
</quantity>
,
<date pageId="6" pageNumber="167" value="1948-05-01" valueMax="1948-05-05" valueMin="1948-05-01">
<collectingDate pageId="6" pageNumber="167" value="1948-05-01" valueMax="1948-05-05" valueMin="1948-05-01">1-5 May 1948</collectingDate>
</date>
,
<collectorName box="[307,361,1385,1408]" pageId="6" pageNumber="167">J. Fu</collectorName>
,
<specimenCount box="[371,410,1384,1408]" pageId="6" pageNumber="167" type="male">2 ♂</specimenCount>
(
<collectionCode box="[424,509,1384,1408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="167" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="6.[184,776,1315,1961]" pageId="6" pageNumber="167">
<emphasis bold="true" box="[184,353,1419,1443]" pageId="6" pageNumber="167">Adult habitus.</emphasis>
Obscured by specimen condition but general color dark brown to black. Head black, pronotum black with pale lateral margins. Costal area of wings pale, contrasting with dark brown on rest of wing membrane. Antennae and legs dark brown to black.
</paragraph>
<paragraph blockId="6.[184,776,1315,1961]" lastBlockId="6.[812,1404,1082,1660]" pageId="6" pageNumber="167">
<emphasis bold="true" box="[184,250,1626,1650]" pageId="6" pageNumber="167">Male.</emphasis>
Forewing length
<quantity box="[444,557,1626,1650]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="6" pageNumber="167" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15-16 mm</quantity>
. Tergum 10 bearing a pair of short, triangular spines positioned on either side of epiproct sclerite, and a pair of darkened, blunt projections on posterior margin above cercal bases (
<figureCitation box="[192,271,1765,1789]" captionStart="Figs" captionStartId="6.[184,234,979,1003]" captionTargetBox="[263,1348,422,893]" captionTargetId="figure-388@6.[252,1354,415,902]" captionTargetPageId="6" captionText="Figs. 20-22. Kiotina quadrituberculata structures. 20. Male terminalia, dorsal, 21. Male paraproct, caudoventral aspect, 22. Male sternum 9." figureDoi="http://doi.org/10.5281/zenodo.4754785" httpUri="https://zenodo.org/record/4754785/files/figure.png" pageId="6" pageNumber="167">Fig. 20</figureCitation>
). Epiproct sclerite Y-shaped with short stem and truncate posterior margin. Paraprocts narrowly triangular with outer margins sclerotized and inner margins pale (
<figureCitation box="[373,475,1868,1892]" captionStart="Figs" captionStartId="6.[184,234,979,1003]" captionTargetBox="[263,1348,422,893]" captionTargetId="figure-388@6.[252,1354,415,902]" captionTargetPageId="6" captionText="Figs. 20-22. Kiotina quadrituberculata structures. 20. Male terminalia, dorsal, 21. Male paraproct, caudoventral aspect, 22. Male sternum 9." figureDoi="http://doi.org/10.5281/zenodo.4754785" httpUri="https://zenodo.org/record/4754785/files/figure.png" pageId="6" pageNumber="167">Figs. 21</figureCitation>
); suture along base of paraprocts dark. Hammer transversely oval (
<figureCitation box="[688,764,1903,1927]" captionStart="Figs" captionStartId="6.[184,234,979,1003]" captionTargetBox="[263,1348,422,893]" captionTargetId="figure-388@6.[252,1354,415,902]" captionTargetPageId="6" captionText="Figs. 20-22. Kiotina quadrituberculata structures. 20. Male terminalia, dorsal, 21. Male paraproct, caudoventral aspect, 22. Male sternum 9." figureDoi="http://doi.org/10.5281/zenodo.4754785" httpUri="https://zenodo.org/record/4754785/files/figure.png" pageId="6" pageNumber="167">Fig. 22</figureCitation>
). Aedeagus not everted, but apparently a membranous, multilobed, wide sac armed with microtrichia.
</paragraph>
<paragraph blockId="6.[812,1404,1082,1660]" box="[812,1032,1152,1176]" pageId="6" pageNumber="167">
<emphasis bold="true" box="[812,904,1152,1176]" pageId="6" pageNumber="167">Female.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="6.[812,1404,1082,1660]" box="[812,1014,1186,1211]" pageId="6" pageNumber="167">
<emphasis bold="true" box="[812,885,1187,1211]" pageId="6" pageNumber="167">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="167" type="discussion">
<paragraph blockId="6.[812,1404,1082,1660]" pageId="6" pageNumber="167">
<emphasis bold="true" box="[812,924,1221,1245]" pageId="6" pageNumber="167">Remarks.</emphasis>
The type material for this species is presumed lost or destroyed, and to our knowledge, no
<typeStatus box="[848,940,1291,1315]" pageId="6" pageNumber="167">neotype</typeStatus>
has been proposed. Although these
<specimenCount pageId="6" pageNumber="167" type="male">two males</specimenCount>
are in poor condition, they are apparently from a site very near the type locality and were taken in the same time frame as the
<typeStatus box="[1124,1229,1394,1418]" pageId="6" pageNumber="167">holotype</typeStatus>
. Consequently, we regard these specimens as topotype representatives for this species known only from
<collectingRegion box="[812,1000,1498,1522]" country="China" name="Fujian" pageId="6" pageNumber="167">Fujian Province</collectingRegion>
,
<collectingCountry box="[1016,1088,1498,1522]" name="China" pageId="6" pageNumber="167">China</collectingCountry>
. Fresh material, including gravid females from the type locality would be highly desirable. In the absence of fresh material from the type locality, one of these specimens would be a suitable
<typeStatus box="[961,1057,1636,1660]" pageId="6" pageNumber="167">neotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>