treatments-xml/data/5B/56/E8/5B56E872FFA74A63FF328843FC49EEF6.xml
2024-06-21 12:37:31 +02:00

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26 KiB
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<document id="781E76B64DA93C3506563870DCED2BD7" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FFA74A63FF328843FC49EEF6" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole kasparii Longino 2019, new species" docType="treatment" docVersion="22" lastPageNumber="41" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="40" updateTime="1698726215157" updateUser="ExternalLinkService">
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<mods:title id="FB46B2A3A6F4C3A2D72FE11ABAFB6FFE">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
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<mods:namePart id="EA423312D23EA84684B3B8B1F7E5AAC6">Longino, John T.</mods:namePart>
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<mods:title id="9F501178AB9278DA54B2092A47D50388">Zootaxa</mods:title>
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<mods:part id="DBA3ACD6D853CF4C05E4605C018DD225">
<mods:date id="5FF9D89C3CD1807E434D35838A69B6E5">2019</mods:date>
<mods:detail id="177F1FBA619049C3D1E0254555DBDF94" type="pubDate">
<mods:number id="F1C551EB21617C65D94611F3D67593A7">2019-05-06</mods:number>
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<mods:number id="64415BBABC79701613D934923BAB2941">4599</mods:number>
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<mods:identifier id="004CE932B5048E701B0827F0BCFE6DD6" type="DOI">10.11646/zootaxa.4599.1.1</mods:identifier>
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<treatment id="5B56E872FFA74A63FF328843FC49EEF6" ID-DOI="http://doi.org/10.5281/zenodo.5586198" ID-GBIF-Taxon="157062613" ID-Zenodo-Dep="5586198" LSID="urn:lsid:plazi:treatment:5B56E872FFA74A63FF328843FC49EEF6" httpUri="http://treatment.plazi.org/id/5B56E872FFA74A63FF328843FC49EEF6" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<subSubSection id="9BE50AEFFFA74A6CFF328843FF02EC3C" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="D3405964FFA74A6CFF328843FDF5EC1D" blockId="39.[151,513,812,871]" box="[151,513,812,838]" pageId="39" pageNumber="40">
<heading id="8808EE08FFA74A6CFF328843FDF5EC1D" bold="true" box="[151,513,812,838]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<emphasis id="E18B8576FFA74A6CFF328843FDF5EC1D" bold="true" box="[151,513,812,838]" pageId="39" pageNumber="40">
<taxonomicName id="14FF22E7FFA74A6CFF328843FE9EEC1D" authority="Longino, 2019" authorityName="Longino" authorityYear="2019" box="[151,362,812,838]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kasparii" status="sp. nov.">
<emphasis id="E18B8576FFA74A6CFF328843FE9EEC1D" bold="true" box="[151,362,812,838]" italics="true" pageId="39" pageNumber="40">Pheidole kasparii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAB8380DFFA74A6CFED48843FDF5EC1D" box="[369,513,812,838]" pageId="39" pageNumber="40" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D3405964FFA74A6CFF328821FF02EC3C" blockId="39.[151,513,812,871]" box="[151,246,846,871]" pageId="39" pageNumber="40">
(
<figureCitation id="4BC445E1FFA74A6CFF3B8821FF19EC3C" box="[158,237,846,871]" captionStart="PLATE 5" captionStartId="71.[151,238,1954,1977]" captionTargetId="figure@71.[202,1384,194,1933]" captionTargetPageId="71" captionText="PLATE 5." httpUri="https://zenodo.org/record/2668237/files/figure.png" pageId="39" pageNumber="40">Plate 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA74A6CFF3288F9FC6FECF4" box="[151,923,918,943]" pageId="39" pageNumber="40" type="description">
<paragraph id="D3405964FFA74A6CFF3288F9FC6FECF4" blockId="39.[151,1437,918,2023]" box="[151,923,918,943]" pageId="39" pageNumber="40">
<taxonomicName id="14FF22E7FFA74A6CFF3288F9FF0FECF4" authorityName="Biolleyi Forel" authorityYear="1908" box="[151,251,918,943]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="E18B8576FFA74A6CFF3288F9FF0FECF4" box="[151,251,918,943]" italics="true" pageId="39" pageNumber="40">Pheidole</emphasis>
</taxonomicName>
JTL-248: morphospecies code previously used on AntWeb.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA74A6CFF6288D4FDDAEBDC" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="D3405964FFA74A6CFF6288D4FDB1EB38" blockId="39.[151,1437,918,2023]" pageId="39" pageNumber="40">
<materialsCitation id="63975339FFA74A6CFF6288D4FE87EB40" ID-GBIF-Occurrence="2237851585" collectingDate="2015-06-25" collectionCode="MUCR" collectorName="J. Longino" country="Costa Rica" elevation="0" location="Santa Maria" pageId="39" pageNumber="40" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
<typeStatus id="0C44E7C6FFA74A6CFF6288D4FEABEC88" box="[199,351,955,979]" pageId="39" pageNumber="40" type="holotype">HOLOTYPE</typeStatus>
: 1 major worker,
<collectingCountry id="ABE819F4FFA74A6CFD8388D4FD50EC88" box="[550,676,954,979]" name="Costa Rica" pageId="39" pageNumber="40">Costa Rica</collectingCountry>
,
<collectingRegion id="113B9786FFA74A6CFD1588D4FCE3EC88" box="[688,791,955,979]" country="Costa Rica" name="San Jose" pageId="39" pageNumber="40">San José</collectingRegion>
:
<quantity id="1407F481FFA74A6CFC8788D5FCA0EC88" box="[802,852,954,979]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="40" unit="km" value="8.0">8km</quantity>
S
<location id="D6200FBFFFA74A6CFCD688D4FC0BEC88" LSID="urn:lsid:plazi:treatment:5B56E872FFA74A63FF328843FC49EEF6:D6200FBFFFA74A6CFCD688D4FC0BEC88" box="[883,1023,954,979]" country="Costa Rica" name="Santa Maria" pageId="39" pageNumber="40" stateProvince="San Jose">Santa Maria</location>
, 9.57666 -83.94911 ±
<elevation id="58D2BE57FFA74A6CFB5288D4FA63EC89" box="[1271,1431,955,979]" isEnumeration="true" pageId="39" pageNumber="40">
<quantity id="1407F481FFA74A6CFB5288D4FAC7EC89" box="[1271,1331,955,979]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="40" unit="m" value="20.0">20 m</quantity>
,
<quantity id="1407F481FFA74A6CFA9A88D4FA63EC89" box="[1343,1431,955,979]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="39" pageNumber="40" unit="m" value="1700.0">1700 m</quantity>
</elevation>
,
<date id="A7417FA4FFA74A6CFF3288B1FED3ECAC" box="[151,295,990,1015]" pageId="39" pageNumber="40" value="2015-06-25">
<collectingDate id="B705864CFFA74A6CFF3288B1FED3ECAC" box="[151,295,990,1015]" pageId="39" pageNumber="40" value="2015-06-25">25-Jun-2015</collectingDate>
</date>
, cloud forest, under epiphytes (
<collectorName id="7E0A3CB2FFA74A6CFD6188B0FCBFECAC" box="[708,843,990,1015]" pageId="39" pageNumber="40">J. Longino</collectorName>
, JTL9356) [
<collectionCode id="B5EEC1A1FFA74A6CFC4F88B0FBB6ECAC" box="[1002,1090,991,1015]" country="Costa Rica" name="Museo de Insectos" pageId="39" pageNumber="40">MUCR</collectionCode>
, unique specimen identifier CASENT0637174]
</materialsCitation>
.
<typeStatus id="0C44E7C6FFA74A6CFED98F6CFDEFEB40" box="[380,539,1027,1051]" pageId="39" pageNumber="40" type="paratype">PARATYPES</typeStatus>
: major, minor worker, dealate queen:
<materialsCitation id="63975339FFA74A6CFC1A8F6CFAD7EB40" ID-GBIF-Occurrence="2237851534" box="[959,1315,1026,1051]" collectingDate="2015-06-25" collectionCode="MUCR" collectorName="J. Longino" country="Costa Rica" elevation="0" location="Santa Maria" pageId="39" pageNumber="40" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
same data as holotype [
<collectionCode id="B5EEC1A1FFA74A6CFB648F6CFAE8EB40" box="[1217,1308,1027,1051]" country="Costa Rica" name="Museo de Insectos" pageId="39" pageNumber="40">MUCR</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="63975339FFA74A6CFA888F6CFDB6EB38" ID-GBIF-Occurrence="2237851465" collectingDate="2015-06-25" collectionCode="CAS, DZUP, JTLC, MCZC, USNM" collectorName="J. Longino" country="Costa Rica" elevation="0" location="Santa Maria" pageId="39" pageNumber="40" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
same data except 9.56895 -83.94756 ±
<elevation id="58D2BE57FFA74A6CFE768F48FD86EB65" box="[467,626,1063,1087]" isEnumeration="true" pageId="39" pageNumber="40">
<quantity id="1407F481FFA74A6CFE768F48FDFBEB65" box="[467,527,1063,1087]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="40" unit="m" value="20.0">20 m</quantity>
,
<quantity id="1407F481FFA74A6CFDBE8F48FD86EB65" box="[539,626,1063,1087]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="39" pageNumber="40" unit="m" value="1770.0">1770 m</quantity>
</elevation>
,
<date id="A7417FA4FFA74A6CFDDA8F49FCE4EB64" box="[639,784,1062,1087]" pageId="39" pageNumber="40" value="2015-06-25">
<collectingDate id="B705864CFFA74A6CFDDA8F49FCE4EB64" box="[639,784,1062,1087]" pageId="39" pageNumber="40" value="2015-06-25">25-Jun-2015</collectingDate>
</date>
, cloud forest, nest under epiphytes (
<collectorName id="7E0A3CB2FFA74A6CFB028F48FAD6EB64" box="[1191,1314,1062,1087]" pageId="39" pageNumber="40">J. Longino</collectorName>
, JTL9358) [
<collectionCode id="B5EEC1A1FFA74A6CFF3B8F24FF2DEB38" box="[158,217,1099,1123]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="40" type="Museum">CAS</collectionCode>
,
<collectionCode id="B5EEC1A1FFA74A6CFF418F24FED9EB39" box="[228,301,1099,1122]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="39" pageNumber="40" type="University or college">DZUP</collectionCode>
,
<collectionCode id="B5EEC1A1FFA74A6CFE9C8F24FE88EB38" box="[313,380,1099,1123]" pageId="39" pageNumber="40">JTLC</collectionCode>
,
<collectionCode id="B5EEC1A1FFA74A6CFE238F24FE2CEB38" box="[390,472,1099,1123]" pageId="39" pageNumber="40">MCZC</collectionCode>
,
<collectionCode id="B5EEC1A1FFA74A6CFE418F24FDCFEB38" box="[484,571,1099,1123]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="D3405964FFA74A6CFF628F01FDDAEBDC" blockId="39.[151,1437,918,2023]" box="[199,558,1134,1159]" pageId="39" pageNumber="40">
<emphasis id="E18B8576FFA74A6CFF628F01FE51EBDC" bold="true" box="[199,421,1134,1159]" pageId="39" pageNumber="40">Geographic range.</emphasis>
<collectingCountry id="ABE819F4FFA74A6CFE098F00FDDEEBDC" box="[428,554,1134,1159]" name="Costa Rica" pageId="39" pageNumber="40">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA74A6CFF628FFDFC6CEAB5" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="D3405964FFA74A6CFF628FFDFC6CEAB5" blockId="39.[151,1437,918,2023]" pageId="39" pageNumber="40">
<emphasis id="E18B8576FFA74A6CFF628FFDFE6FEBF0" bold="true" box="[199,411,1170,1195]" pageId="39" pageNumber="40">
Diagnosis.
<emphasis id="E18B8576FFA74A6CFEF78FFDFE6FEBF0" bold="true" box="[338,411,1170,1195]" italics="true" pageId="39" pageNumber="40">Minor</emphasis>
</emphasis>
: face smooth and shining; vertex margin shallowly emarginate in full face view; promesonotal groove absent; pronotum entirely smooth and shiny; katepisternum shiny, with varying development of faint foveolation; propodeal spines in form of obtuse to right angles, not spiniform; gaster smooth and shining; abundant erect setae on mesosomal and gastral dorsum; tibiae lack erect setae; color red brown to yellow (color variation can be intracolonial).
<emphasis id="E18B8576FFA74A6CFE5F8E4DFDB5EA60" bold="true" box="[506,577,1314,1339]" italics="true" pageId="39" pageNumber="40">Major</emphasis>
: median hypostomal tooth strongly developed; inner hypostomal teeth very weakly developed, hardly visible, closer to outer hypostomal teeth than to medial tooth, recessed, not on ventral margin of hypostomal bridge; ventral margin of head in profile view convex; scape base terete; face mostly smooth and shiny, with longitudinal carinulae between eye and antennal fossa; propodeal spines in form of short, triangular teeth, &lt;one fifth length of posterior face of propodeum; gastral dorsum smooth and shiny; abundant erect setae on sides of head, mesosomal and gastral dorsum; tibiae lack erect setae.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA74A6CFF628E95FE2AE99C" pageId="39" pageNumber="40" type="description">
<paragraph id="D3405964FFA74A6CFF628E95FD9BE96C" blockId="39.[151,1437,918,2023]" pageId="39" pageNumber="40">
<emphasis id="E18B8576FFA74A6CFF628E95FDD9E948" bold="true" box="[199,557,1530,1555]" pageId="39" pageNumber="40">Measurements, minor worker</emphasis>
: HW 0.41, HL 0.43, SL 0.34, EL 0.10, WL 0.47, PSL 0.01, PTW 0.07, PPW 0.09, CI 96, SI 85, PSLI 2, PPI 129 (n=2).
</paragraph>
<paragraph id="D3405964FFA74A6CFF628D2DFD9BE924" blockId="39.[151,1437,918,2023]" pageId="39" pageNumber="40">
<emphasis id="E18B8576FFA74A6CFF628D2DFDD9E900" bold="true" box="[199,557,1602,1627]" pageId="39" pageNumber="40">Measurements, major worker</emphasis>
: HW 0.79, HL 0.87, SL 0.40, EL 0.12, WL 0.68, PSL 0.02, PTW 0.12, PPW 0.16, CI 91, SI 50, PSLI 3, PPI 133 (n=2).
</paragraph>
<paragraph id="D3405964FFA74A6CFF628DE5FE2AE99C" blockId="39.[151,1437,918,2023]" pageId="39" pageNumber="40">
<emphasis id="E18B8576FFA74A6CFF628DE5FE38E9F8" bold="true" box="[199,460,1674,1699]" pageId="39" pageNumber="40">Measurements, queen</emphasis>
: HW 0.87, HL 0.81, SL 0.48, EL 0.26, WL 1.39, PSL 0.00, PTW 0.27, PPW 0.38, CI 108, SL 55, PSLI 0, PPI 139.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA74A6CFF628DBDFB0AE80D" pageId="39" pageNumber="40" type="biology_ecology">
<paragraph id="D3405964FFA74A6CFF628DBDFB0AE80D" blockId="39.[151,1437,918,2023]" pageId="39" pageNumber="40">
<emphasis id="E18B8576FFA74A6CFF628DBDFED3E9B0" bold="true" box="[199,295,1746,1771]" pageId="39" pageNumber="40">Biology.</emphasis>
This species occurs in cloud forest to high montane oak forest. It is locally abundant around the
<typeStatus id="0C44E7C6FFA74A6CFAC98DBCFA68E9B0" box="[1388,1436,1747,1771]" pageId="39" pageNumber="40">type</typeStatus>
locality. Minor and major workers recruit to ground baits, and workers occur in Winkler samples of sifted litter and rotten wood. Nests have been found under epiphytes on recent branchfalls, under epiphytes on clay banks (road cut), in an acorn amongst ground litter, and in a cavity in a low, moss-covered live stem of
<taxonomicName id="14FF22E7FFA74A6CFB348C50FB0CE80D" box="[1169,1272,1855,1878]" class="Magnoliopsida" family="Siparunaceae" genus="Siparuna" kingdom="Plantae" order="Laurales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
<emphasis id="E18B8576FFA74A6CFB348C50FB0CE80D" box="[1169,1272,1855,1878]" italics="true" pageId="39" pageNumber="40">Siparuna</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA74A63FF628C0DFEE4EED3" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="discussion">
<paragraph id="D3405964FFA74A6CFF628C0DFE0EE8BC" blockId="39.[151,1437,918,2023]" pageId="39" pageNumber="40">
<emphasis id="E18B8576FFA74A6CFF628C0DFEBCE820" bold="true" box="[199,328,1890,1915]" pageId="39" pageNumber="40">Comments</emphasis>
. A complex of very similar species occurs in wet forests of
<collectingCountry id="ABE819F4FFA74A6CFC558C0CFB9BE820" box="[1008,1135,1890,1915]" name="Costa Rica" pageId="39" pageNumber="40">Costa Rica</collectingCountry>
:
<taxonomicName id="14FF22E7FFA74A6CFBDE8C0BFB07E820" authorityName="Longino" authorityYear="2019" box="[1147,1267,1890,1915]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kasparii">
<emphasis id="E18B8576FFA74A6CFBDE8C0BFB07E820" box="[1147,1267,1890,1915]" italics="true" pageId="39" pageNumber="40">P. kasparii</emphasis>
</taxonomicName>
,
<taxonomicName id="14FF22E7FFA74A6CFAA48C0BFA91E820" authorityName="Wilson" authorityYear="2003" box="[1281,1381,1890,1915]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="nitella">
<emphasis id="E18B8576FFA74A6CFAA48C0BFA91E820" box="[1281,1381,1890,1915]" italics="true" pageId="39" pageNumber="40">P. nitella</emphasis>
</taxonomicName>
, and
<taxonomicName id="14FF22E7FFA74A6CFF328CE8FED5E8C5" authorityName="Wilson" authorityYear="2003" box="[151,289,1927,1950]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="E18B8576FFA74A6CFF328CE8FED5E8C5" box="[151,289,1927,1950]" italics="true" pageId="39" pageNumber="40">P. sagittaria</emphasis>
</taxonomicName>
.
<taxonomicName id="14FF22E7FFA74A6CFE8A8CE9FE03E8C4" authorityName="Longino" authorityYear="2019" box="[303,503,1926,1951]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kasparii">
<emphasis id="E18B8576FFA74A6CFE8A8CE9FE03E8C4" box="[303,503,1926,1951]" italics="true" pageId="39" pageNumber="40">Pheidole kasparii</emphasis>
</taxonomicName>
as currently understood is a narrow endemic, known only from the vicinity of the
<typeStatus id="0C44E7C6FFA74A6CFF328CC4FF33E898" box="[151,199,1963,1987]" pageId="39" pageNumber="40">type</typeStatus>
locality, and allopatric to the other two.
<taxonomicName id="14FF22E7FFA74A6CFD2A8CC5FCB5E898" authorityName="Wilson" authorityYear="2003" box="[655,833,1962,1987]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="nitella">
<emphasis id="E18B8576FFA74A6CFD2A8CC5FCB5E898" box="[655,833,1962,1987]" italics="true" pageId="39" pageNumber="40">Pheidole nitella</emphasis>
</taxonomicName>
and
<taxonomicName id="14FF22E7FFA74A6CFCDD8CC3FBF4E898" authorityName="Wilson" authorityYear="2003" box="[888,1024,1964,1987]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="E18B8576FFA74A6CFCDD8CC3FBF4E898" box="[888,1024,1964,1987]" italics="true" pageId="39" pageNumber="40">P. sagittaria</emphasis>
</taxonomicName>
occur on the Atlantic slope of
<collectingCountry id="ABE819F4FFA74A6CFAF98CC4FF39E8BC" name="Costa Rica" pageId="39" pageNumber="40">Costa Rica</collectingCountry>
, where they are sympatric.
</paragraph>
<paragraph id="D3405964FFA84A63FF628BF8FEE4EED3" blockId="40.[151,1437,151,429]" pageId="40" pageNumber="41">
The minor workers of the complex are not readily distinguishable, sharing the habitus of being very small, almost completely smooth and shining, and with very short propodeal spines that are no more than low angles in profile. The major workers are more distinct. The inner hypostomal teeth of
<taxonomicName id="14FF22E7FFA84A63FC438B8EFBA9EFA3" authorityName="Longino" authorityYear="2019" box="[998,1117,223,248]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="kasparii">
<emphasis id="E18B8576FFA84A63FC438B8EFBA9EFA3" box="[998,1117,223,248]" italics="true" pageId="40" pageNumber="41">P. kasparii</emphasis>
</taxonomicName>
are farther apart and smaller than the teeth of
<taxonomicName id="14FF22E7FFA84A63FEC48A6AFE33EE46" authorityName="Wilson" authorityYear="2003" box="[353,455,260,285]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="nitella">
<emphasis id="E18B8576FFA84A63FEC48A6AFE33EE46" box="[353,455,260,285]" italics="true" pageId="40" pageNumber="41">P. nitella</emphasis>
</taxonomicName>
and
<taxonomicName id="14FF22E7FFA84A63FDA18A6AFD64EE47" authorityName="Wilson" authorityYear="2003" box="[516,656,261,284]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="E18B8576FFA84A63FDA18A6AFD64EE47" box="[516,656,261,284]" italics="true" pageId="40" pageNumber="41">P. sagittaria</emphasis>
</taxonomicName>
, and recessed behind the ventral margin of the hypostomal bridge.
<taxonomicName id="14FF22E7FFA84A63FF328A48FE84EE1B" authorityName="Wilson" authorityYear="2003" box="[151,368,295,320]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="E18B8576FFA84A63FF328A48FE84EE1B" box="[151,368,295,320]" italics="true" pageId="40" pageNumber="41">Pheidole sagittaria</emphasis>
</taxonomicName>
has the ventral margin of the head in profile very flat, while on
<taxonomicName id="14FF22E7FFA84A63FBF78A46FB3FEE1B" authorityName="Longino" authorityYear="2019" box="[1106,1227,295,320]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="kasparii">
<emphasis id="E18B8576FFA84A63FBF78A46FB3FEE1B" box="[1106,1227,295,320]" italics="true" pageId="40" pageNumber="41">P. kasparii</emphasis>
</taxonomicName>
and
<taxonomicName id="14FF22E7FFA84A63FAA18A46FA9DEE1B" authorityName="Wilson" authorityYear="2003" box="[1284,1385,295,320]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="nitella">
<emphasis id="E18B8576FFA84A63FAA18A46FA9DEE1B" box="[1284,1385,295,320]" italics="true" pageId="40" pageNumber="41">P. nitella</emphasis>
</taxonomicName>
it is more convex. The head of
<taxonomicName id="14FF22E7FFA84A63FE658A22FDD7EE3E" authorityName="Wilson" authorityYear="2003" box="[448,547,332,357]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="nitella">
<emphasis id="E18B8576FFA84A63FE658A22FDD7EE3E" box="[448,547,332,357]" italics="true" pageId="40" pageNumber="41">P. nitella</emphasis>
</taxonomicName>
is relatively more narrow than the heads of
<taxonomicName id="14FF22E7FFA84A63FBAE8A22FB8AEE3E" authorityName="Longino" authorityYear="2019" box="[1035,1150,332,357]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="kasparii">
<emphasis id="E18B8576FFA84A63FBAE8A22FB8AEE3E" box="[1035,1150,332,357]" italics="true" pageId="40" pageNumber="41">P. kasparii</emphasis>
</taxonomicName>
and
<taxonomicName id="14FF22E7FFA84A63FB128A22FACBEE3F" authorityName="Wilson" authorityYear="2003" box="[1207,1343,333,356]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="sagittaria">
<emphasis id="E18B8576FFA84A63FB128A22FACBEE3F" box="[1207,1343,333,356]" italics="true" pageId="40" pageNumber="41">P. sagittaria</emphasis>
</taxonomicName>
(CI ~84 vs. 90-95).
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA84A63FF628AFBFC49EEF6" box="[199,957,404,429]" pageId="40" pageNumber="41" type="etymology">
<paragraph id="D3405964FFA84A63FF628AFBFC49EEF6" blockId="40.[151,1437,151,429]" box="[199,957,404,429]" pageId="40" pageNumber="41">
<emphasis id="E18B8576FFA84A63FF628AFBFEBCEEF6" bold="true" box="[199,328,404,429]" pageId="40" pageNumber="41">Etymology</emphasis>
: In honor of ant ecologist extraordinaire, Mike Kaspari.
</paragraph>
</subSubSection>
</treatment>
</document>