233 lines
24 KiB
XML
233 lines
24 KiB
XML
<document id="558589C8C661EBF114D59B40F8EDE15F" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FF9E4A54FF328E3DFDA0EC37" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole costaricensis Longino 2019, new species" docType="treatment" docVersion="22" lastPageNumber="32" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="31" updateTime="1698726215157" updateUser="ExternalLinkService">
|
|
<mods:mods id="A5058758EF2A65295A757B1E7B3B0EA6" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="D23128C3F753ECBD7C8902E98237FC85">
|
|
<mods:title id="8E2C3420A662DF821D67297200B55072">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="CBC970DCEF91F37E272B3939FA60F3C8" type="personal">
|
|
<mods:role id="72EBBAF318F8E152CA7266187B4BE756">
|
|
<mods:roleTerm id="3C37B55CF1A5786A3F45AC41B59B338C">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="98962B9E812F0FE4B82FA8F39D47D481">Longino, John T.</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="CB185514B06AD2562ED52351835E90FA">text</mods:typeOfResource>
|
|
<mods:relatedItem id="C7375675E84136E890898A2C436F8423" type="host">
|
|
<mods:titleInfo id="F898907788F4167B58B7C1B48ED0C1C1">
|
|
<mods:title id="D4BD84C669272FC8D61E3A63C307E4EA">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="FA455C9A418899AA2F397090818F899F">
|
|
<mods:date id="ACB166A207019709B94AA281F09FA9D1">2019</mods:date>
|
|
<mods:detail id="AED33416BAAD7C135CE3C0EB25D196AA" type="pubDate">
|
|
<mods:number id="D94E5718240FD8B491ED5C2EBCB9CF29">2019-05-06</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="4DD626CF3F47E889F47A96CF04CD4560" type="volume">
|
|
<mods:number id="D31757CBE6375CB492E8F3EA833DBBEC">4599</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="B86FA9A460D5F04BDD2D2E9FBE81C577" type="issue">
|
|
<mods:number id="9BE164F0162CDC4C8C4025CC6ACF9850">1</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="8FEFE6F58CB62C53EFFF4D7C17C65877" unit="page">
|
|
<mods:start id="6A4902B0461D297DD10805E470E6B2A1">1</mods:start>
|
|
<mods:end id="7769911EC7878D267B61ADE2FBBDA311">126</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="6AA624E94CFEF18195ED4543FFC90B59">journal article</mods:classification>
|
|
<mods:identifier id="253E77CB3B97022B21FEF95AED8E4A41" type="CLB-Dataset">26920</mods:identifier>
|
|
<mods:identifier id="DA6DD25F6366433E5D87049E2B516188" type="DOI">10.11646/zootaxa.4599.1.1</mods:identifier>
|
|
<mods:identifier id="29986132876BE18349C1E05AA779552D" type="GBIF-Dataset">ea727a9b-324b-4c6b-a625-bc7cdd204473</mods:identifier>
|
|
<mods:identifier id="0309243F6EFBE2C4474CE5A7F18C5872" type="ISSN">1175-5326</mods:identifier>
|
|
<mods:identifier id="8EDFE833F958396790072143E4400CCE" type="Zenodo-Dep">2668058</mods:identifier>
|
|
<mods:identifier id="D921831A4978B2CB56464BF51C288F7C" type="ZooBank">7CDD24FF-5BA3-4D06-898D-A3E220515D4C</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="5B56E872FF9E4A54FF328E3DFDA0EC37" ID-DOI="http://doi.org/10.5281/zenodo.5586180" ID-GBIF-Taxon="157062600" ID-Zenodo-Dep="5586180" LSID="urn:lsid:plazi:treatment:5B56E872FF9E4A54FF328E3DFDA0EC37" httpUri="http://treatment.plazi.org/id/5B56E872FF9E4A54FF328E3DFDA0EC37" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
|
|
<subSubSection id="9BE50AEFFF9E4A55FF328E3DFEF1EAD5" pageId="30" pageNumber="31" type="nomenclature">
|
|
<paragraph id="D3405964FF9E4A55FF328E3DFDCDEA36" blockId="30.[151,569,1362,1422]" box="[151,569,1362,1389]" pageId="30" pageNumber="31">
|
|
<heading id="8808EE08FF9E4A55FF328E3DFDCDEA36" bold="true" box="[151,569,1362,1389]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
|
|
<emphasis id="E18B8576FF9E4A55FF328E3DFDCDEA36" bold="true" box="[151,569,1362,1389]" pageId="30" pageNumber="31">
|
|
<taxonomicName id="14FF22E7FF9E4A55FF328E3DFE56EA37" authority="Longino, 2019" authorityName="Longino" authorityYear="2019" box="[151,418,1362,1388]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
|
|
<emphasis id="E18B8576FF9E4A55FF328E3DFE56EA37" bold="true" box="[151,418,1362,1388]" italics="true" pageId="30" pageNumber="31">Pheidole costaricensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="FAB8380DFF9E4A55FE0C8E3CFDCDEA36" box="[425,569,1363,1389]" pageId="30" pageNumber="31" rank="species">new species</taxonomicNameLabel>
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="D3405964FF9E4A55FF328E1AFEF1EAD5" blockId="30.[151,569,1362,1422]" box="[151,261,1397,1422]" pageId="30" pageNumber="31">
|
|
(
|
|
<figureCitation id="4BC445E1FF9E4A55FF3B8E1AFF08EAD5" box="[158,252,1397,1422]" captionStart="PLATE 19" captionStartId="85.[151,238,1954,1977]" captionTargetId="figure@85.[202,1384,194,1933]" captionTargetPageId="85" captionText="PLATE 19." httpUri="https://zenodo.org/record/2668267/files/figure.png" pageId="30" pageNumber="31">Plate 19</figureCitation>
|
|
)
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFF9E4A55FF328ED2FC6FEA8D" box="[151,923,1469,1494]" pageId="30" pageNumber="31" type="description">
|
|
<paragraph id="D3405964FF9E4A55FF328ED2FC6FEA8D" blockId="30.[151,1436,1469,2034]" box="[151,923,1469,1494]" pageId="30" pageNumber="31">
|
|
<taxonomicName id="14FF22E7FF9E4A55FF328ED2FF0FEA8D" authorityName="Biolleyi Forel" authorityYear="1908" box="[151,251,1469,1494]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E18B8576FF9E4A55FF328ED2FF0FEA8D" box="[151,251,1469,1494]" italics="true" pageId="30" pageNumber="31">Pheidole</emphasis>
|
|
</taxonomicName>
|
|
JTL-279: morphospecies code previously used on AntWeb.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFF9E4A55FF628E8DFDDAE9F5" pageId="30" pageNumber="31" type="materials_examined">
|
|
<paragraph id="D3405964FF9E4A55FF628E8DFD16E9D1" blockId="30.[151,1436,1469,2034]" pageId="30" pageNumber="31">
|
|
<materialsCitation id="63975339FF9E4A55FF628E8DFE14E919" ID-GBIF-Occurrence="2237851488" collectingDate="2015-06-19" collectionCode="MUCR" collectorName="J. Longino" country="Costa Rica" elevation="0" location="Cerro Platano" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
|
|
<typeStatus id="0C44E7C6FF9E4A55FF628E8DFEABEAA1" box="[199,351,1506,1530]" pageId="30" pageNumber="31" type="holotype">HOLOTYPE</typeStatus>
|
|
: 1 major worker,
|
|
<collectingCountry id="ABE819F4FF9E4A55FD878E8DFD6BEAA1" box="[546,671,1505,1530]" name="Costa Rica" pageId="30" pageNumber="31">Costa Rica</collectingCountry>
|
|
,
|
|
<collectingRegion id="113B9786FF9E4A55FD0F8E8EFD0DEAA1" box="[682,761,1505,1530]" country="Costa Rica" name="Limon" pageId="30" pageNumber="31">Limón</collectingRegion>
|
|
:
|
|
<location id="D6200FBFFF9E4A55FCA68E8DFC55EAA1" LSID="urn:lsid:plazi:treatment:5B56E872FF9E4A54FF328E3DFDA0EC37:D6200FBFFF9E4A55FCA68E8DFC55EAA1" box="[771,929,1505,1530]" country="Costa Rica" name="Cerro Platano" pageId="30" pageNumber="31" stateProvince="Limon">Cerro Platano</location>
|
|
,
|
|
<quantity id="1407F481FF9E4A55FC0E8E8EFC18EAA1" box="[939,1004,1505,1530]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="30" pageNumber="31" unit="km" value="26.0">26km</quantity>
|
|
WSW
|
|
<collectingRegion id="113B9786FF9E4A55FB988E8EFB7FEAA1" box="[1085,1163,1505,1530]" country="Costa Rica" name="Limon" pageId="30" pageNumber="31">Limón</collectingRegion>
|
|
, 9.86439 -83.24371 ±
|
|
<elevation id="58D2BE57FF9E4A55FA258E8EFEE8E946" isEnumeration="true" pageId="30" pageNumber="31">
|
|
<quantity id="1407F481FF9E4A55FA258E8EFF5BE946" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="m" value="50.0">50 m</quantity>
|
|
,
|
|
<quantity id="1407F481FF9E4A55FF658D6AFEE8E946" box="[192,284,1541,1566]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="31" unit="m" value="1050.0">1050 m</quantity>
|
|
</elevation>
|
|
,
|
|
<date id="A7417FA4FF9E4A55FE888D6AFE49E945" box="[301,445,1541,1566]" pageId="30" pageNumber="31" value="2015-06-19">
|
|
<collectingDate id="B705864CFF9E4A55FE888D6AFE49E945" box="[301,445,1541,1566]" pageId="30" pageNumber="31" value="2015-06-19">19-Jun-2015</collectingDate>
|
|
</date>
|
|
, mature cloud forest, under stone (
|
|
<collectorName id="7E0A3CB2FF9E4A55FCC48D69FC15E945" box="[865,993,1541,1566]" pageId="30" pageNumber="31">J. Longino</collectorName>
|
|
, JTL9332) [
|
|
<collectionCode id="B5EEC1A1FF9E4A55FBD48D69FB3DE945" box="[1137,1225,1542,1566]" country="Costa Rica" name="Museo de Insectos" pageId="30" pageNumber="31">MUCR</collectionCode>
|
|
, unique specimen identifier CASENT0637143]
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="63975339FF9E4A55FE4F8D45FD2AE9D1" ID-GBIF-Occurrence="2237851560" collectingDate="2006-01-03" collectingDateMax="2016-05-03" collectingDateMin="2006-01-03" collectionCode="MCZC, MUCR, CAS, USNM" collectorName="J. Longino" country="Costa Rica" elevation="0" location="Cerro Platano" pageId="30" pageNumber="31" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
|
|
<typeStatus id="0C44E7C6FF9E4A55FE4F8D45FD7DE919" box="[490,649,1578,1602]" pageId="30" pageNumber="31" type="paratype">PARATYPES</typeStatus>
|
|
: major, minor workers: same data as holotype [
|
|
<collectionCode id="B5EEC1A1FF9E4A55FB3D8D45FB19E919" box="[1176,1261,1578,1602]" pageId="30" pageNumber="31">MCZC</collectionCode>
|
|
,
|
|
<collectionCode id="B5EEC1A1FF9E4A55FB5D8D45FAA5E919" box="[1272,1361,1578,1602]" country="Costa Rica" name="Museo de Insectos" pageId="30" pageNumber="31">MUCR</collectionCode>
|
|
]; same data except 9.86621 -83.24203 ±
|
|
<elevation id="58D2BE57FF9E4A55FDB48D22FD41E93E" box="[529,693,1613,1638]" isEnumeration="true" pageId="30" pageNumber="31">
|
|
<quantity id="1407F481FF9E4A55FDB48D22FDBBE93E" box="[529,591,1613,1638]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="m" value="50.0">50 m</quantity>
|
|
,
|
|
<quantity id="1407F481FF9E4A55FDFB8D21FD41E93E" box="[606,693,1614,1638]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="30" pageNumber="31" unit="m" value="1110.0">1110 m</quantity>
|
|
</elevation>
|
|
, at bait (ADMAC, Ba-E-
|
|
<date id="A7417FA4FF9E4A55FC7A8D22FBADE93D" box="[991,1113,1613,1638]" pageId="30" pageNumber="31" value="2006-01-03" valueMax="2006-04-03" valueMin="2006-01-03">
|
|
<collectingDate id="B705864CFF9E4A55FC7A8D22FBADE93D" box="[991,1113,1613,1638]" pageId="30" pageNumber="31" value="2006-01-03" valueMax="2006-04-03" valueMin="2006-01-03">03-1-04-06</collectingDate>
|
|
</date>
|
|
) [
|
|
<collectionCode id="B5EEC1A1FF9E4A55FBD78D21FB44E93D" box="[1138,1200,1614,1638]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="30" pageNumber="31" type="Museum">CAS</collectionCode>
|
|
]; 9.86439 -83.24371 ±
|
|
<elevation id="58D2BE57FF9E4A55FF038D1EFEB0E9D2" box="[166,324,1649,1674]" isEnumeration="true" pageId="30" pageNumber="31">
|
|
<quantity id="1407F481FF9E4A55FF038D1EFF15E9D2" box="[166,225,1649,1674]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="m" value="50.0">50 m</quantity>
|
|
,
|
|
<quantity id="1407F481FF9E4A55FF488D1EFEB0E9D2" box="[237,324,1649,1674]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="31" unit="m" value="1050.0">1050 m</quantity>
|
|
</elevation>
|
|
, at bait (Ba-E-
|
|
<date id="A7417FA4FF9E4A55FE488D1EFD93E9D1" box="[493,615,1649,1674]" pageId="30" pageNumber="31" value="2016-02-03" valueMax="2016-05-03" valueMin="2016-02-03">
|
|
<collectingDate id="B705864CFF9E4A55FE488D1EFD93E9D1" box="[493,615,1649,1674]" pageId="30" pageNumber="31" value="2016-02-03" valueMax="2016-05-03" valueMin="2016-02-03">03-2-05-16</collectingDate>
|
|
</date>
|
|
) [
|
|
<collectionCode id="B5EEC1A1FF9E4A55FDD88D1DFD23E9D1" box="[637,727,1650,1674]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="30" pageNumber="31" type="Museum">USNM</collectionCode>
|
|
]
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="D3405964FF9E4A55FF628DFAFDDAE9F5" blockId="30.[151,1436,1469,2034]" box="[199,558,1685,1710]" pageId="30" pageNumber="31">
|
|
<emphasis id="E18B8576FF9E4A55FF628DFAFE51E9F5" bold="true" box="[199,421,1685,1710]" pageId="30" pageNumber="31">Geographic range.</emphasis>
|
|
<collectingCountry id="ABE819F4FF9E4A55FE098DF9FDDEE9F5" box="[428,554,1685,1710]" name="Costa Rica" pageId="30" pageNumber="31">Costa Rica</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFF9E4A55FF628DD6FCBEE8A9" pageId="30" pageNumber="31" type="diagnosis">
|
|
<paragraph id="D3405964FF9E4A55FF628DD6FCBEE8A9" blockId="30.[151,1436,1469,2034]" pageId="30" pageNumber="31">
|
|
<emphasis id="E18B8576FF9E4A55FF628DD6FE6AE989" bold="true" box="[199,414,1721,1746]" pageId="30" pageNumber="31">
|
|
Diagnosis.
|
|
<emphasis id="E18B8576FF9E4A55FEF08DD6FE6AE989" bold="true" box="[341,414,1721,1746]" italics="true" pageId="30" pageNumber="31">Minor</emphasis>
|
|
</emphasis>
|
|
: face and mesosoma foveolate, anteroventral katepisternum smooth and shining; promesonotum evenly convex, promesonotal groove absent; propodeal spines one fourth length of posterior face of propodeum; gaster smooth and shining; abundant erect setae on mesosomal dorsum and gaster; tibiae with 2-3 erect setae; color orange.
|
|
<emphasis id="E18B8576FF9E4A55FE128C4BFE0AE866" bold="true" box="[439,510,1828,1853]" italics="true" pageId="30" pageNumber="31">Major</emphasis>
|
|
: inner hypostomal teeth stout, closely spaced; scape base terete; face with distinct scrobes, delimited dorsally by frontal carinae and forming concave trough below them, ventral and posterior margins less delimited, surface of scrobe smooth and shiny; head weakly depressed posteriorly; face surface generally smooth and shiny, with sparse, feeble longitudinal carinulae; propodeal spines one fourth length of posterior face of propodeum; gastral dorsum smooth and shining; sides of head with abundant erect setae; abundant flexuous erect setae on mesosomal dorsum, tibiae, and gaster.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFF9F4A54FF628BF8FD9BEE44" pageId="31" pageNumber="32" type="description">
|
|
<paragraph id="D3405964FF9F4A54FF628BF8FD89EF8E" blockId="31.[151,1437,151,876]" pageId="31" pageNumber="32">
|
|
<emphasis id="E18B8576FF9F4A54FF628BF8FDD9EFEB" bold="true" box="[199,557,151,176]" pageId="31" pageNumber="32">Measurements, minor worker</emphasis>
|
|
: HW 0.48, HL 0.51, SL 0.48, EL 0.11, WL 0.63, PSL 0.04, PTW 0.09, PPW 0.14, CI 93, SI 102, PSLI 7, PPI 161 (n=4).
|
|
</paragraph>
|
|
<paragraph id="D3405964FF9F4A54FF628B8EFD9BEE44" blockId="31.[151,1437,151,876]" pageId="31" pageNumber="32">
|
|
<emphasis id="E18B8576FF9F4A54FF628B8EFDD9EFA1" bold="true" box="[199,557,225,250]" pageId="31" pageNumber="32">Measurements, major worker</emphasis>
|
|
: HW 1.03, HL 1.24, SL 0.48, EL 0.14, WL 0.98, PSL 0.07, PTW 0.18, PPW 0.36, CI 83, SI 47, PSLI 7, PPI 200 (n=3).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFF9F4A54FF628A44FA6CEE3C" pageId="31" pageNumber="32" type="biology_ecology">
|
|
<paragraph id="D3405964FF9F4A54FF628A44FA6CEE3C" blockId="31.[151,1437,151,876]" pageId="31" pageNumber="32">
|
|
<emphasis id="E18B8576FF9F4A54FF628A44FED3EE1F" bold="true" box="[199,295,299,324]" pageId="31" pageNumber="32">Biology.</emphasis>
|
|
This species occurs in lowland rain forest to lower cloud forest habitats. Minor and occasional major workers recruit to ground baits. Workers occur in Winkler samples. One nest was observed in soil beneath a stone.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFF9F4A54FF628A1BFA8BEC1D" pageId="31" pageNumber="32" type="discussion">
|
|
<paragraph id="D3405964FF9F4A54FF628A1BFA8BEC1D" blockId="31.[151,1437,151,876]" pageId="31" pageNumber="32">
|
|
<emphasis id="E18B8576FF9F4A54FF628A1BFEBCEED6" bold="true" box="[199,328,372,397]" pageId="31" pageNumber="32">Comments</emphasis>
|
|
. This species remains poorly delimited. The
|
|
<typeStatus id="0C44E7C6FF9F4A54FC9B8A1AFC9AEED6" box="[830,878,373,397]" pageId="31" pageNumber="32">type</typeStatus>
|
|
locality is a cloud forest site southwest of
|
|
<collectingRegion id="113B9786FF9F4A54FAEF8A1BFA6CEED6" box="[1354,1432,372,397]" country="Costa Rica" name="Limon" pageId="31" pageNumber="32">Limón</collectingRegion>
|
|
, where minor workers were definitively associated with major workers. At La Selva Biological Station and the lower Barva transect above it, many minor workers that have been collected at baits and in Winkler samples are tentatively identified as
|
|
<taxonomicName id="14FF22E7FF9F4A54FE028A8BFDA2EEA0" authorityName="Longino" authorityYear="2019" box="[423,598,484,507]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="costaricensis">
|
|
<emphasis id="E18B8576FF9F4A54FE028A8BFDA2EEA0" box="[423,598,484,507]" italics="true" pageId="31" pageNumber="32">P. costaricensis</emphasis>
|
|
</taxonomicName>
|
|
. However, no major workers have been collected from this area. Three of the minor workers have DNA barcodes and form a tight cluster, near the cluster of
|
|
<taxonomicName id="14FF22E7FF9F4A54FBCE8966FAEDED7B" authorityName="Longino" authorityYear="2019" box="[1131,1305,519,544]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="corniclypeus">
|
|
<emphasis id="E18B8576FF9F4A54FBCE8966FAEDED7B" box="[1131,1305,519,544]" italics="true" pageId="31" pageNumber="32">P. corniclypeus</emphasis>
|
|
</taxonomicName>
|
|
. Given the inability to morphologically distinguish minor workers of
|
|
<taxonomicName id="14FF22E7FF9F4A54FCBA8941FC38ED1F" authorityName="Longino" authorityYear="2019" box="[799,972,557,581]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="costaricensis">
|
|
<emphasis id="E18B8576FF9F4A54FCBA8941FC38ED1F" box="[799,972,557,581]" italics="true" pageId="31" pageNumber="32">P. costaricensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="14FF22E7FF9F4A54FBA48941FB5FED1E" authorityName="Longino" authorityYear="2019" box="[1025,1195,556,581]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="corniclypeus">
|
|
<emphasis id="E18B8576FF9F4A54FBA48941FB5FED1E" box="[1025,1195,556,581]" italics="true" pageId="31" pageNumber="32">P. corniclypeus</emphasis>
|
|
</taxonomicName>
|
|
, and the proximity of true
|
|
<taxonomicName id="14FF22E7FF9F4A54FF6F893DFE82ED31" authorityName="Longino" authorityYear="2019" box="[202,374,593,618]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="corniclypeus">
|
|
<emphasis id="E18B8576FF9F4A54FF6F893DFE82ED31" box="[202,374,593,618]" italics="true" pageId="31" pageNumber="32">P. corniclypeus</emphasis>
|
|
</taxonomicName>
|
|
, there is the possibility that both species could occur on the Barva transect. A population was sampled on the Pacific slope of
|
|
<collectingCountry id="ABE819F4FF9F4A54FDB78918FD66EDD4" box="[530,658,630,655]" name="Costa Rica" pageId="31" pageNumber="32">Costa Rica</collectingCountry>
|
|
, in lower montane forest above the Rio Savegre, and minors were associated with major workers. These major workers are somewhat intermediate between
|
|
<taxonomicName id="14FF22E7FF9F4A54FB3889F3FAB9EDE8" authorityName="Longino" authorityYear="2019" box="[1181,1357,668,691]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="costaricensis">
|
|
<emphasis id="E18B8576FF9F4A54FB3889F3FAB9EDE8" box="[1181,1357,668,691]" italics="true" pageId="31" pageNumber="32">P. costaricensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="14FF22E7FF9F4A54FA2289F3FED2ED83" authorityName="Longino" authorityYear="2019" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="corniclypeus">
|
|
<emphasis id="E18B8576FF9F4A54FA2289F3FED2ED83" italics="true" pageId="31" pageNumber="32">P. corniclypeus</emphasis>
|
|
</taxonomicName>
|
|
, with more pronounced facial carinae, and small bulges on the lateral clypeus where the large teeth are on
|
|
<taxonomicName id="14FF22E7FF9F4A54FF418989FE7BEDA6" authorityName="Longino" authorityYear="2019" box="[228,399,740,765]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="corniclypeus">
|
|
<emphasis id="E18B8576FF9F4A54FF418989FE7BEDA6" box="[228,399,740,765]" italics="true" pageId="31" pageNumber="32">P. corniclypeus</emphasis>
|
|
</taxonomicName>
|
|
. Finally, BOLD has a tight cluster of specimens from the lowland dry forest of
|
|
<collectingRegion id="113B9786FF9F4A54FAB1898AFA6CEDA6" box="[1300,1432,741,765]" country="Costa Rica" name="Guanacaste" pageId="31" pageNumber="32">Guanacaste</collectingRegion>
|
|
, close to the clusters for
|
|
<taxonomicName id="14FF22E7FF9F4A54FE018865FDBBEC79" authorityName="Longino" authorityYear="2019" box="[420,591,777,802]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="corniclypeus">
|
|
<emphasis id="E18B8576FF9F4A54FE018865FDBBEC79" box="[420,591,777,802]" italics="true" pageId="31" pageNumber="32">P. corniclypeus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="14FF22E7FF9F4A54FD238865FCC1EC7A" authorityName="Longino" authorityYear="2019" box="[646,821,778,801]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="costaricensis">
|
|
<emphasis id="E18B8576FF9F4A54FD238865FCC1EC7A" box="[646,821,778,801]" italics="true" pageId="31" pageNumber="32">P. costaricensis</emphasis>
|
|
</taxonomicName>
|
|
. Images of these specimens include a lateral view of a major worker, and there is no evidence of a clypeal bulge or tooth. See additional comments under
|
|
<taxonomicName id="14FF22E7FF9F4A54FB488840FA8CEC1C" authorityName="Longino" authorityYear="2019" box="[1261,1400,814,839]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="andersoni">
|
|
<emphasis id="E18B8576FF9F4A54FB488840FA8CEC1C" box="[1261,1400,814,839]" italics="true" pageId="31" pageNumber="32">P. andersoni</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFF9F4A54FF62883CFDA0EC37" box="[199,596,851,876]" pageId="31" pageNumber="32" type="etymology">
|
|
<paragraph id="D3405964FF9F4A54FF62883CFDA0EC37" blockId="31.[151,1437,151,876]" box="[199,596,851,876]" pageId="31" pageNumber="32">
|
|
<emphasis id="E18B8576FF9F4A54FF62883CFEBCEC37" bold="true" box="[199,328,851,876]" pageId="31" pageNumber="32">Etymology</emphasis>
|
|
: From the
|
|
<typeStatus id="0C44E7C6FF9F4A54FE66883BFE07EC37" box="[451,499,852,876]" pageId="31" pageNumber="32">type</typeStatus>
|
|
locality.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |