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<mods:title id="44DE27B7A4BB86956F7EFD2F18C88C02">Updated catalog and generic keys of the Leiodidae (Insecta: Coleoptera) of the Neotropical region (“ Latin America ”: Mexico, the West Indies, and Central and South America)</mods:title>
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<paragraph id="C659EA29FFBA8719FF75CD93FE0BF0A6" blockId="65.[151,476,678,705]" box="[151,476,678,705]" pageId="65" pageNumber="66">
<heading id="9D115D45FFBA8719FF75CD93FE0BF0A6" bold="true" box="[151,476,678,705]" fontSize="11" level="1" pageId="65" pageNumber="66" reason="1">
<emphasis id="F492363BFFBA8719FF75CD93FE0BF0A6" bold="true" box="[151,476,678,705]" pageId="65" pageNumber="66">
Genus
<taxonomicName id="01E691AAFFBA8719FF0FCD92FE0BF0A6" ID-CoL="8VRFW" authority="Panzer" authorityName="Panzer" box="[237,476,678,705]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FF0FCD92FEABF0A6" bold="true" box="[237,380,678,705]" italics="true" pageId="65" pageNumber="66">Agathidium</emphasis>
Panzer
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8EFCB9A2FFBA8719FF75CDDAFD3CF422" pageId="65" pageNumber="66" type="reference_group">
<paragraph id="C659EA29FFBA8719FF75CDDAFC6AF136" blockId="65.[151,1437,750,1785]" pageId="65" pageNumber="66">
<taxonomicName id="01E691AAFFBA8719FF75CDDAFE68F16E" authority="Panzer, 1797" authorityName="Panzer" authorityYear="1797" box="[151,447,750,777]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FF75CDDAFEF7F16E" bold="true" box="[151,288,750,777]" italics="true" pageId="65" pageNumber="66">Agathidium</emphasis>
<bibRefCitation id="A27797D8FFBA8719FECACDDBFE68F16E" author="Panzer, G. W." box="[296,447,751,777]" pageId="65" pageNumber="66" refId="ref103189" refString="Panzer, G. W. (1797) Fauna insectorum Germanicae initia oder deutschlands Insekten. Heft 37. Felssecker, Nurnberg." type="book" year="1797">Panzer, 1797</bibRefCitation>
</taxonomicName>
: no. 13.
<typeStatus id="195D548BFFBA8719FDC3CDDBFD8EF16E" box="[545,601,751,777]" pageId="65" pageNumber="66">Type</typeStatus>
species:
<taxonomicName id="01E691AAFFBA8719FD20CDC4FBECF16E" authority="Herbst [1792]" authorityName="Herbst" authorityYear="1792" box="[706,1083,750,777]" class="Insecta" family="Tetratomidae" genus="Tetratoma" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="F492363BFFBA8719FD20CDC4FC42F16E" box="[706,917,751,777]" italics="true" pageId="65" pageNumber="66">Tetratoma globosa</emphasis>
Herbst [1792]
</taxonomicName>
(=
<taxonomicName id="01E691AAFFBA8719FB86CDDBFEF8F14A" authority="Linnaeus [1758]" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="seminulum">
<emphasis id="F492363BFFBA8719FB86CDDBFAFAF16E" box="[1124,1325,751,777]" italics="true" pageId="65" pageNumber="66">Silpha seminulum</emphasis>
Linnaeus [1758]
</taxonomicName>
, syn. in ?? [already by Gyllenhal 1810: 573] (by monotypy). Note: Proposed as nomen protectum by provision of
<bibRefCitation id="A27797D8FFBA8719FE90CC03FE21F136" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[370,502,823,849]" pageId="65" pageNumber="66" refId="ref101125" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Fourth edition. London. International Trust for Zoological Nomenclature. Available from: http: // www. iczn. org / iczn / index. jsp (accessed 24 September 2019)" type="url" year="1999">ICZN 1999</bibRefCitation>
: Art. 23.9 (
<bibRefCitation id="A27797D8FFBA8719FD90CC03FC67F136" author="Wheeler, Q. D. &amp; Miller, K. B." box="[626,944,822,849]" pageId="65" pageNumber="66" pagination="1 - 95" refId="ref110046" refString="Wheeler, Q. D. &amp; Miller, K. B. (2005) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part I. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 290, 1 - 95. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 290 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Wheeler &amp; Miller, 2005: 34</bibRefCitation>
).
</paragraph>
<paragraph id="C659EA29FFBA8719FF75CC68FBF8F186" blockId="65.[151,1437,750,1785]" pageId="65" pageNumber="66">
<treatmentCitationGroup id="E6F6CD07FFBA8719FF75CC68FBF8F186" pageId="65" pageNumber="66">
<emphasis id="F492363BFFBA8719FF75CC68FF7DF112" box="[151,170,860,885]" italics="true" pageId="65" pageNumber="66">=</emphasis>
<treatmentCitation id="4747CC38FFBA8719FF50CC6FFDD1F112" author="Kugelann, J. G." box="[178,518,858,885]" page="535" pageId="65" pageNumber="66" year="1794">
<taxonomicName id="01E691AAFFBA8719FF50CC6FFDD1F112" ID-CoL="9LKHG" authority="Kugelann, 1794: 535" authorityName="Kugelann" authorityPageNumber="535" authorityYear="1794" box="[178,518,858,885]" class="Insecta" family="Leiodidae" genus="Volvoxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FF50CC6FFED8F112" box="[178,271,859,885]" italics="true" pageId="65" pageNumber="66">Volvoxis</emphasis>
<bibRefCitation id="A27797D8FFBA8719FEFACC6FFDD1F112" author="Kugelann, J. G." box="[280,518,858,885]" pageId="65" pageNumber="66" pagination="513 - 582" refId="ref101871" refString="Kugelann, J. G. (1794) Verzeichniss der in einigen Gegenden Preussens bis jetzt entdeckten Kaefer-Arten nebst kurzen Nachrichten von denselben. Neuestes Magazin fur die Liebhaber der Entomologie, 1, 513 - 582." type="journal article" year="1794">Kugelann, 1794: 535</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(not preoccupied by
<taxonomicName id="01E691AAFFBA8719FD1BCC6FFBD0F112" authority="Linnaeus [1758]" authorityName="Linnaeus" authorityYear="1758" box="[761,1031,858,885]" class="Chlorophyceae" family="Volvocaceae" genus="Volvox" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="65" pageNumber="66" phylum="Zoophyta" rank="genus">
<emphasis id="F492363BFFBA8719FD1BCC6FFC93F112" box="[761,836,859,885]" italics="true" pageId="65" pageNumber="66">Volvox</emphasis>
Linnaeus [1758]
</taxonomicName>
, Chlorophyta).
<typeStatus id="195D548BFFBA8719FB5ACC6FFB27F112" box="[1208,1264,859,885]" pageId="65" pageNumber="66">Type</typeStatus>
species:
<taxonomicName id="01E691AAFFBA8719FABBCC68FE1AF1FE" authority="Herbst" authorityName="Herbst" authorityYear="1792" class="Insecta" family="Tetratomidae" genus="Tetratoma" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="F492363BFFBA8719FABBCC68FEACF1FE" italics="true" pageId="65" pageNumber="66">Tetratoma globosa</emphasis>
Herbst
</taxonomicName>
(=
<taxonomicName id="01E691AAFFBA8719FE17CC4BFCAAF1FE" authority="Linnaeus [1758]" authorityName="Linnaeus" authorityYear="1758" box="[501,893,894,921]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="seminulum">
<emphasis id="F492363BFFBA8719FE17CC4BFD6BF1FE" box="[501,700,895,921]" italics="true" pageId="65" pageNumber="66">Silpha seminulum</emphasis>
Linnaeus [1758]
</taxonomicName>
by subsequent designation of
<treatmentCitation id="4747CC38FFBA8719FB30CC4BFA4BF1FE" author="Newton, A. F. Jr." box="[1234,1436,894,921]" page="95" pageId="65" pageNumber="66" year="1998">
<bibRefCitation id="A27797D8FFBA8719FB30CC4BFA4BF1FE" author="Newton, A. F. Jr." box="[1234,1436,894,921]" pageId="65" pageNumber="66" pagination="41 - 177" refId="ref102584" refString="Newton, A. F. Jr. (1998) Phylogenetic problems, current classification and generic catalog of World Leiodidae (including Cholevidae). Museo Regionale di Scienze Naturali di Torino Atti, 8, 41 - 177." type="journal article" year="1998">Newton, 1998: 95</bibRefCitation>
</treatmentCitation>
(10 species in original description). Note: Proposed as nomen oblitum by provision of
<bibRefCitation id="A27797D8FFBA8719FB49CC97FAF8F1DA" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[1195,1327,931,957]" pageId="65" pageNumber="66" refId="ref101125" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Fourth edition. London. International Trust for Zoological Nomenclature. Available from: http: // www. iczn. org / iczn / index. jsp (accessed 24 September 2019)" type="url" year="1999">ICZN 1999</bibRefCitation>
: Art. 23.9 (
<treatmentCitation id="4747CC38FFBA8719FF08CCF3FDF2F186" author="Wheeler, Q. D. &amp; Miller, K. B." box="[234,549,966,993]" page="34" pageId="65" pageNumber="66" year="2005">
<bibRefCitation id="A27797D8FFBA8719FF08CCF3FDF2F186" author="Wheeler, Q. D. &amp; Miller, K. B." box="[234,549,966,993]" pageId="65" pageNumber="66" pagination="1 - 95" refId="ref110046" refString="Wheeler, Q. D. &amp; Miller, K. B. (2005) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part I. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 290, 1 - 95. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 290 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Wheeler &amp; Miller, 2005: 34</bibRefCitation>
</treatmentCitation>
) due to lack of use as a valid name after 1899.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C659EA29FFBA8719FF75CCD8FBCBF6DE" blockId="65.[151,1437,750,1785]" pageId="65" pageNumber="66">
<treatmentCitationGroup id="E6F6CD07FFBA8719FF75CCD8FBCBF6DE" pageId="65" pageNumber="66">
<emphasis id="F492363BFFBA8719FF75CCD8FF7DF662" box="[151,170,1004,1029]" italics="true" pageId="65" pageNumber="66">=</emphasis>
<treatmentCitation id="4747CC38FFBA8719FF53CCDFFE27F662" author="Illiger, J. K. W." box="[177,496,1003,1029]" page="81" pageId="65" pageNumber="66" year="1798">
<taxonomicName id="01E691AAFFBA8719FF53CCDFFE27F662" ID-CoL="8VRFW" authority="Illiger, 1798: 81" authorityName="Illiger" authorityPageNumber="81" authorityYear="1798" box="[177,496,1003,1029]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FF53CCDFFEE3F662" box="[177,308,1003,1029]" italics="true" pageId="65" pageNumber="66">Agathidium</emphasis>
<bibRefCitation id="A27797D8FFBA8719FED9CCDFFE27F662" author="Illiger, J. K. W." box="[315,496,1003,1029]" pageId="65" pageNumber="66" refId="ref101176" refString="Illiger, J. K. W. (1798) Verzeichniss der Kafer Preussens. J. J. Gebauer, Halle, xlii + 510 pp." type="book" year="1798">Illiger, 1798: 81</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(unnecessary replacement name for
<taxonomicName id="01E691AAFFBA8719FC6CCCDFFB65F662" ID-CoL="9LKHG" authority="Kugelann [1794]" authorityName="Kugelann" authorityYear="1794" box="[910,1202,1002,1029]" class="Insecta" family="Leiodidae" genus="Volvoxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FC6CCCDFFC3CF662" box="[910,1003,1003,1029]" italics="true" pageId="65" pageNumber="66">Volvoxis</emphasis>
<bibRefCitation id="A27797D8FFBA8719FC11CCDFFB65F662" author="Kugelann, J. G." box="[1011,1202,1002,1029]" pageId="65" pageNumber="66" pagination="513 - 582" refId="ref101871" refString="Kugelann, J. G. (1794) Verzeichniss der in einigen Gegenden Preussens bis jetzt entdeckten Kaefer-Arten nebst kurzen Nachrichten von denselben. Neuestes Magazin fur die Liebhaber der Entomologie, 1, 513 - 582." type="journal article" year="1794">Kugelann [1794]</bibRefCitation>
</taxonomicName>
(in part), incorrectly assumed to be preoccupied by
<taxonomicName id="01E691AAFFBA8719FDA2CB3BFC98F64E" authority="Linnaeus [1758]" authorityName="Linnaeus" authorityYear="1758" box="[576,847,1038,1065]" class="Chlorophyceae" family="Volvocaceae" genus="Volvox" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="65" pageNumber="66" phylum="Zoophyta" rank="genus">
<emphasis id="F492363BFFBA8719FDA2CB3BFD5CF64E" box="[576,651,1039,1065]" italics="true" pageId="65" pageNumber="66">Volvox</emphasis>
Linnaeus [1758]
</taxonomicName>
).
<typeStatus id="195D548BFFBA8719FC81CB3BFC4CF64E" box="[867,923,1039,1065]" pageId="65" pageNumber="66">Type</typeStatus>
species:
<taxonomicName id="01E691AAFFBA8719FBE1CB24FAACF64E" authority="Herbst [1792]" authorityName="Herbst" authorityYear="1792" box="[1027,1403,1038,1065]" class="Insecta" family="Tetratomidae" genus="Tetratoma" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="F492363BFFBA8719FBE1CB24FB01F64E" box="[1027,1238,1039,1065]" italics="true" pageId="65" pageNumber="66">Tetratoma globosa</emphasis>
Herbst [1792]
</taxonomicName>
(=
<taxonomicName id="01E691AAFFBA8719FF01CB07FDBDF62A" authority="Linnaeus [1785]" authorityName="Linnaeus" authorityYear="1785" box="[227,618,1074,1101]" class="Insecta" family="Curculionidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="seminulum">
<emphasis id="F492363BFFBA8719FF01CB07FE7EF62A" box="[227,425,1075,1101]" italics="true" pageId="65" pageNumber="66">Silpha seminulum</emphasis>
Linnaeus [1785]
</taxonomicName>
), fixed by autotypy as replacement name for
<taxonomicName id="01E691AAFFBA8719FBBFCB07FB6DF62A" ID-CoL="9LKHG" authorityName="Kugelann" authorityYear="1794" box="[1117,1210,1075,1101]" class="Insecta" family="Leiodidae" genus="Volvoxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FBBFCB07FB6DF62A" box="[1117,1210,1075,1101]" italics="true" pageId="65" pageNumber="66">Volvoxis</emphasis>
</taxonomicName>
. Note: It is not clear if Illiger intended to propose a new generic name here; he did not refer the generic name to
<bibRefCitation id="A27797D8FFBA8719FB14CB63FA40F616" author="Panzer, G. W." box="[1270,1431,1111,1137]" pageId="65" pageNumber="66" refId="ref103189" refString="Panzer, G. W. (1797) Fauna insectorum Germanicae initia oder deutschlands Insekten. Heft 37. Felssecker, Nurnberg." type="book" year="1797">Panzer (1797)</bibRefCitation>
, although he did refer to
<treatmentCitation id="4747CC38FFBA8719FE0CCB4FFD1EF6F2" author="Panzer, G. W." box="[494,713,1147,1173]" page="13" pageId="65" pageNumber="66" year="1797">
<bibRefCitation id="A27797D8FFBA8719FE0CCB4FFD1EF6F2" author="Panzer, G. W." box="[494,713,1147,1173]" pageId="65" pageNumber="66" refId="ref103189" refString="Panzer, G. W. (1797) Fauna insectorum Germanicae initia oder deutschlands Insekten. Heft 37. Felssecker, Nurnberg." type="book" year="1797">Panzers (1797: 13)</bibRefCitation>
</treatmentCitation>
species treatment of
<taxonomicName id="01E691AAFFBA8719FC56CB4FFB7DF6F2" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[948,1194,1147,1173]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="globosum">
<emphasis id="F492363BFFBA8719FC56CB4FFB7DF6F2" box="[948,1194,1147,1173]" italics="true" pageId="65" pageNumber="66">Agathidium globosum</emphasis>
</taxonomicName>
. The above treatment assumes it was an intentional proposal of a new name to replace
<taxonomicName id="01E691AAFFBA8719FC55CBABFBC3F6DE" ID-CoL="9LKHG" authorityName="Kugelann" authorityYear="1794" box="[951,1044,1183,1209]" class="Insecta" family="Leiodidae" genus="Volvoxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FC55CBABFBC3F6DE" box="[951,1044,1183,1209]" italics="true" pageId="65" pageNumber="66">Volvoxis</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C659EA29FFBA8719FF75CBF7FD3CF422" blockId="65.[151,1437,750,1785]" pageId="65" pageNumber="66">
Note 1:
<taxonomicName id="01E691AAFFBA8719FF12CBF7FE16F6BA" ID-CoL="9LKHG" authority="Kugelann" authorityName="Kugelann" authorityYear="1794" box="[240,449,1219,1245]" class="Insecta" family="Leiodidae" genus="Volvoxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FF12CBF7FE9AF6BA" box="[240,333,1219,1245]" italics="true" pageId="65" pageNumber="66">Volvoxis</emphasis>
Kugelann
</taxonomicName>
has priority for the name of this genus, but this name was rejected and replaced with
<taxonomicName id="01E691AAFFBA8719FA96CBF7FE90F766" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FA96CBF7FE90F766" italics="true" pageId="65" pageNumber="66">Agathidium</emphasis>
</taxonomicName>
by
<bibRefCitation id="A27797D8FFBA8719FE92CBD3FDD1F766" author="Illiger, J. K. W." box="[368,518,1255,1281]" pageId="65" pageNumber="66" refId="ref101176" refString="Illiger, J. K. W. (1798) Verzeichniss der Kafer Preussens. J. J. Gebauer, Halle, xlii + 510 pp." type="book" year="1798">Illiger (1798)</bibRefCitation>
in the mistaken belief that
<taxonomicName id="01E691AAFFBA8719FCD4CBD3FC44F766" ID-CoL="9LKHG" authorityName="Kugelann" authorityYear="1794" box="[822,915,1255,1281]" class="Insecta" family="Leiodidae" genus="Volvoxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FCD4CBD3FC44F766" box="[822,915,1255,1281]" italics="true" pageId="65" pageNumber="66">Volvoxis</emphasis>
</taxonomicName>
was preoccupied by
<taxonomicName id="01E691AAFFBA8719FB9DCBD3FAEDF766" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1758" box="[1151,1338,1255,1281]" class="Chlorophyceae" family="Volvocaceae" genus="Volvox" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="65" pageNumber="66" phylum="Zoophyta" rank="genus">
<emphasis id="F492363BFFBA8719FB9DCBD3FB1DF766" box="[1151,1226,1255,1281]" italics="true" pageId="65" pageNumber="66">Volvox</emphasis>
Linnaeus
</taxonomicName>
, and this replacement name became generally accepted. However,
<bibRefCitation id="A27797D8FFBA8719FC8ACA3FFBD1F742" author="Panzer, G. W." box="[872,1030,1291,1317]" pageId="65" pageNumber="66" refId="ref103189" refString="Panzer, G. W. (1797) Fauna insectorum Germanicae initia oder deutschlands Insekten. Heft 37. Felssecker, Nurnberg." type="book" year="1797">Panzer (1797)</bibRefCitation>
had already used
<taxonomicName id="01E691AAFFBA8719FB31CA3FFA81F742" authorityName="Illiger" authorityYear="1798" box="[1235,1366,1291,1317]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FB31CA3FFA81F742" box="[1235,1366,1291,1317]" italics="true" pageId="65" pageNumber="66">Agathidium</emphasis>
</taxonomicName>
in the same sense a year earlier, without explanation.
<bibRefCitation id="A27797D8FFBA8719FD12CA1BFC51F72E" author="Illiger, J. K. W." box="[752,902,1327,1353]" pageId="65" pageNumber="66" refId="ref101176" refString="Illiger, J. K. W. (1798) Verzeichniss der Kafer Preussens. J. J. Gebauer, Halle, xlii + 510 pp." type="book" year="1798">Illiger (1798)</bibRefCitation>
made no mention of
<bibRefCitation id="A27797D8FFBA8719FB90CA1BFAF4F72E" author="Panzer, G. W." box="[1138,1315,1327,1353]" pageId="65" pageNumber="66" refId="ref103189" refString="Panzer, G. W. (1797) Fauna insectorum Germanicae initia oder deutschlands Insekten. Heft 37. Felssecker, Nurnberg." type="book" year="1797">Panzers (1797)</bibRefCitation>
use of
<taxonomicName id="01E691AAFFBA8719FA96CA1BFE90F70A" authorityName="Illiger" authorityYear="1798" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FA96CA1BFE90F70A" italics="true" pageId="65" pageNumber="66">Agathidium</emphasis>
</taxonomicName>
so these have often been treated as independent names (e.g.,
<bibRefCitation id="A27797D8FFBA8719FC1ACA67FB38F709" author="Newton, A. F. Jr." box="[1016,1263,1362,1390]" pageId="65" pageNumber="66" pagination="41 - 177" refId="ref102584" refString="Newton, A. F. Jr. (1998) Phylogenetic problems, current classification and generic catalog of World Leiodidae (including Cholevidae). Museo Regionale di Scienze Naturali di Torino Atti, 8, 41 - 177." type="journal article" year="1998">
Newton, 1998: 95
<emphasis id="F492363BFFBA8719FB20CA60FB07F70A" box="[1218,1232,1364,1389]" italics="true" pageId="65" pageNumber="66"></emphasis>
96
</bibRefCitation>
), with Panzers name having priority and Illigers name being a junior homonym of it. All three names have the same
<typeStatus id="195D548BFFBA8719FA8ECA43FA4BF7F6" box="[1388,1436,1399,1425]" pageId="65" pageNumber="66">type</typeStatus>
species as indicated above, and thus are objective synonyms.
<bibRefCitation id="A27797D8FFBA8719FC72CAAFFB03F7D2" author="Wheeler, Q. D. &amp; Miller, K. B." box="[912,1236,1434,1461]" pageId="65" pageNumber="66" pagination="1 - 95" refId="ref110046" refString="Wheeler, Q. D. &amp; Miller, K. B. (2005) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part I. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 290, 1 - 95. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 290 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Wheeler &amp; Miller (2005: 34)</bibRefCitation>
cited Article 23.9 of the Code (
<bibRefCitation id="A27797D8FFBA8719FE91CA8BFE2AF7BE" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[371,509,1471,1497]" pageId="65" pageNumber="66" refId="ref101125" refString="ICZN (International Commission on Zoological Nomenclature) (1999) International Code of Zoological Nomenclature. Fourth edition. London. International Trust for Zoological Nomenclature. Available from: http: // www. iczn. org / iczn / index. jsp (accessed 24 September 2019)" type="url" year="1999">ICZN, 1999</bibRefCitation>
) to set aside priority and accept
<taxonomicName id="01E691AAFFBA8719FC84CA8BFBECF7BE" authority="Panzer" authorityName="Panzer" box="[870,1083,1471,1497]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FC84CA8BFC3EF7BE" box="[870,1001,1471,1497]" italics="true" pageId="65" pageNumber="66">Agathidium</emphasis>
Panzer
</taxonomicName>
as the valid name for the genus (nomen protectum) and treat
<taxonomicName id="01E691AAFFBA8719FDC8CAD7FD50F79A" ID-CoL="9LKHG" authorityName="Kugelann" authorityYear="1794" box="[554,647,1507,1533]" class="Insecta" family="Leiodidae" genus="Volvoxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FDC8CAD7FD50F79A" box="[554,647,1507,1533]" italics="true" pageId="65" pageNumber="66">Volvoxis</emphasis>
</taxonomicName>
as a forgotten name (nomen oblitum).
<taxonomicName id="01E691AAFFBA8719FBDCCAD7FB16F79A" box="[1086,1217,1507,1533]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="F492363BFFBA8719FBDCCAD7FB16F79A" box="[1086,1217,1507,1533]" italics="true" pageId="65" pageNumber="66">Agathidium</emphasis>
</taxonomicName>
has been attributed to both Panzer and Illiger at various times, sometimes (by accident) even in the same publication (see
<bibRefCitation id="A27797D8FFBA8719FABCC933FEAEF422" author="Newton, A. F. Jr." pageId="65" pageNumber="66" pagination="41 - 177" refId="ref102584" refString="Newton, A. F. Jr. (1998) Phylogenetic problems, current classification and generic catalog of World Leiodidae (including Cholevidae). Museo Regionale di Scienze Naturali di Torino Atti, 8, 41 - 177." type="journal article" year="1998">Newton, 1998: 94</bibRefCitation>
, 95), but
<bibRefCitation id="A27797D8FFBA8719FE01C91FFDA3F422" author="Panzer, G. W." box="[483,628,1579,1605]" pageId="65" pageNumber="66" refId="ref103189" refString="Panzer, G. W. (1797) Fauna insectorum Germanicae initia oder deutschlands Insekten. Heft 37. Felssecker, Nurnberg." type="book" year="1797">Panzer, 1797</bibRefCitation>
is correct.
</paragraph>
</subSubSection>
<subSubSection id="8EFCB9A2FFBA8719FF75C97BFCC3F5CA" pageId="65" pageNumber="66" type="description">
<paragraph id="C659EA29FFBA8719FF75C97BFACEF40E" blockId="65.[151,1437,750,1785]" box="[151,1305,1614,1642]" pageId="65" pageNumber="66">
Note 2: Revised in
<bibRefCitation id="A27797D8FFBA8719FE8DC97BFDB4F40D" author="Hlisnikovsky, J." box="[367,611,1615,1642]" pageId="65" pageNumber="66" pagination="3 - 239" refId="ref100735" refString="Hlisnikovsky, J. (1964 a) Monographische bearbeitung der gattung Agathidium Panzer (Coleoptera). Acta Entomologica Musei Nationalis Prague, Supplementum 5, 3 - 239." type="journal article" year="1964">Hlisnikovský (1964a)</bibRefCitation>
and
<bibRefCitation id="A27797D8FFBA8719FD7BC97BFC16F40E" author="Miller, K. B. &amp; Wheeler, Q. D." box="[665,961,1614,1641]" pageId="65" pageNumber="66" pagination="1 - 167" refId="ref102247" refString="Miller, K. B. &amp; Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Miller &amp; Wheeler (2005b)</bibRefCitation>
and
<bibRefCitation id="A27797D8FFBA8719FC15C97BFAC3F40E" author="Wheeler, Q. D. &amp; Miller, K. B." box="[1015,1300,1614,1641]" pageId="65" pageNumber="66" pagination="1 - 95" refId="ref110046" refString="Wheeler, Q. D. &amp; Miller, K. B. (2005) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part I. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 290, 1 - 95. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 290 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Wheeler &amp; Miller (2005)</bibRefCitation>
.
</paragraph>
<paragraph id="C659EA29FFBA8719FF75C947FD79F49E" blockId="65.[151,1437,750,1785]" pageId="65" pageNumber="66">
Note 3:
<bibRefCitation id="A27797D8FFBA8719FF12C947FDDEF4EA" author="Wheeler, Q. D. &amp; Miller, K. B." box="[240,521,1650,1677]" pageId="65" pageNumber="66" pagination="1 - 95" refId="ref110046" refString="Wheeler, Q. D. &amp; Miller, K. B. (2005) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part I. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 290, 1 - 95. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 290 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Wheeler &amp; Miller (2005)</bibRefCitation>
abandoned the use of subgenera and proposed a system of species groups, as used below, which may have some relationship to previously established subgenera. Most Mexican and Central American species are in the
<taxonomicName id="01E691AAFFBA8719FDD3C98FFD70F4B2" authorityName="Palisot de Beauvois" authorityYear="1805" box="[561,679,1723,1749]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="oniscoides">
<emphasis id="F492363BFFBA8719FDD3C98FFD70F4B2" box="[561,679,1723,1749]" italics="true" pageId="65" pageNumber="66">oniscoides</emphasis>
</taxonomicName>
species group. The rich species diversity of
<collectingCountry id="BEF1AAB9FFBA8719FB55C98FFADAF4B2" box="[1207,1293,1723,1749]" name="Mexico" pageId="65" pageNumber="66">Mexico</collectingCountry>
and Central America is obviously only partly known.
</paragraph>
<paragraph id="C659EA29FFBA8719FF75C813FD2FF526" blockId="65.[151,760,1830,1857]" box="[151,760,1830,1857]" pageId="65" pageNumber="66">
Key: to species groups in
<bibRefCitation id="A27797D8FFBA8719FE5BC813FD23F526" author="Wheeler, Q. D. &amp; Miller, K. B." box="[441,756,1830,1857]" pageId="65" pageNumber="66" pagination="1 - 95" refId="ref110046" refString="Wheeler, Q. D. &amp; Miller, K. B. (2005) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part I. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 290, 1 - 95. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 290 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Wheeler &amp; Miller, 2005: 37</bibRefCitation>
.
</paragraph>
<paragraph id="C659EA29FFBA8719FF75C85BFC19F5EE" blockId="65.[151,1436,1903,2038]" box="[151,974,1903,1929]" pageId="65" pageNumber="66">Distribution: 822 spp.—Holarctic, Africa, Oriental, northern Neotropical.</paragraph>
<paragraph id="C659EA29FFBA8719FF75C8A7FCC3F5CA" blockId="65.[151,1436,1903,2038]" box="[151,788,1938,1965]" pageId="65" pageNumber="66">
Note 4—
<emphasis id="F492363BFFBA8719FF1DC8A6FCC3F5CA" bold="true" box="[255,788,1938,1965]" pageId="65" pageNumber="66">Species wrongly reported for the Neotropics:</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="8EFCB9A2FFBA871AFF01C883FBDFF29C" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="reference_group">
<paragraph id="C659EA29FFBA8719FF01C883FA4BF592" blockId="65.[151,1436,1903,2038]" pageId="65" pageNumber="66">
<taxonomicName id="01E691AAFFBA8719FF01C883FDE0F5B6" authority="Melsheimer [1844]" authorityName="Melsheimer" authorityYear="1844" box="[227,567,1974,2001]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="exiguum">
<emphasis id="F492363BFFBA8719FF01C883FEB7F5B6" box="[227,352,1975,2001]" italics="true" pageId="65" pageNumber="66">A. exiguum</emphasis>
Melsheimer [1844]
</taxonomicName>
—The record of
<taxonomicName id="01E691AAFFBA8719FD17C883FBE4F5B6" authority="Horn [1880]" authorityName="Horn" authorityYear="1880" box="[757,1075,1974,2001]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="californicum">
<emphasis id="F492363BFFBA8719FD17C883FC75F5B6" box="[757,930,1975,2001]" italics="true" pageId="65" pageNumber="66">A. californicum</emphasis>
<bibRefCitation id="A27797D8FFBA8719FC4BC883FBE4F5B6" author="Horn, G. H." box="[937,1075,1974,2001]" pageId="65" pageNumber="66" pagination="219 - 319" refId="ref100993" refString="Horn, G. H. (1880) Synopsis of the Silphidae of the United States with reference to the genera of other countries. Transactions of the American Entomological Society, 8, 219 - 319, pls. 5 - 7. https: // doi. org / 10.2307 / 25076393" type="journal article" year="1880">Horn [1880]</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="01E691AAFFBA8719FBBAC883FAFFF5B6" authority=", Miller" authorityName="Miller" box="[1112,1320,1975,2001]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="exiguum">
<emphasis id="F492363BFFBA8719FBBAC883FB02F5B6" box="[1112,1237,1975,2001]" italics="true" pageId="65" pageNumber="66">A. exiguum</emphasis>
, Miller
</taxonomicName>
&amp; Wheel- er, 2005b: 156 syn] for
<collectingCountry id="BEF1AAB9FFBA8719FE05C8EFFDB7F592" box="[487,608,2011,2037]" name="Guatemala" pageId="65" pageNumber="66">Guatemala</collectingCountry>
(
<collectingRegion id="042224CBFFBA8719FD8CC8EFFD16F592" box="[622,705,2011,2037]" country="Guatemala" name="Quiche" pageId="65" pageNumber="66">Quiché</collectingRegion>
Mts.,
<quantity id="011E47CCFFBA8719FCE4C8EFFC73F591" box="[774,932,2011,2038]" metricMagnitude="3" metricUnit="m" metricValue="2.4384" metricValueMax="2.7432" metricValueMin="2.1336" pageId="65" pageNumber="66" unit="ft" value="8000.0" valueMax="9000.0" valueMin="7000.0">
7000
<emphasis id="F492363BFFBA8719FCDDC8E8FC9AF592" box="[831,845,2012,2037]" italics="true" pageId="65" pageNumber="66"></emphasis>
9000 ft.
</quantity>
) in
<bibRefCitation id="A27797D8FFBA8719FC2DC8EFFB7BF592" author="Matthews, A." box="[975,1196,2010,2037]" pageId="65" pageNumber="66" pagination="72 - 101" refId="ref102147" refString="Matthews, A. (1887 - 88) Fam. Silphidae. In: Biologia Centrali-Americana. Insects. Coleoptera. Vol. 2 (1). Taylor &amp; Francis, London, pp. 72 - 101, pl. 3." type="journal article" year="1887">Matthews, 1887: 75</bibRefCitation>
is a misidentification
</paragraph>
<paragraph id="C659EA29FFB9871AFF01CFADFC69F2D3" blockId="66.[151,1436,152,323]" box="[227,958,152,180]" pageId="66" pageNumber="67">
of an unknown different species (
<bibRefCitation id="A27797D8FFB9871AFDBACFADFC65F2D3" author="Miller, K. B. &amp; Wheeler, Q. D." box="[600,946,152,180]" pageId="66" pageNumber="67" pagination="1 - 167" refId="ref102247" refString="Miller, K. B. &amp; Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Miller &amp; Wheeler, 2005b: 160</bibRefCitation>
).
</paragraph>
<paragraph id="C659EA29FFB9871AFF01CF89FBDFF29C" blockId="66.[151,1436,152,323]" pageId="66" pageNumber="67">
<treatmentCitationGroup id="E6F6CD07FFB9871AFF01CF89FBDFF29C" pageId="66" pageNumber="67">
<taxonomicName id="01E691AAFFB9871AFF01CF89FD79F2B0" ID-CoL="6633Y" authority="Palisot de Beauvois [1805]" authorityName="Palisot de Beauvois" authorityYear="1805" box="[227,686,188,215]" class="Insecta" family="Leiodidae" genus="Agathidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="oniscoides">
<emphasis id="F492363BFFB9871AFF01CF89FEAEF2B0" box="[227,377,189,215]" italics="true" pageId="66" pageNumber="67">A. oniscoides</emphasis>
Palisot de Beauvois [1805]
</taxonomicName>
—The record of
<treatmentCitation id="4747CC38FFB9871AFC96CF89FB81F2B0" author="Matthews, A." box="[884,1110,189,215]" page="74" pageId="66" pageNumber="67" year="1887">
<bibRefCitation id="A27797D8FFB9871AFC96CF89FB81F2B0" author="Matthews, A." box="[884,1110,189,215]" pageId="66" pageNumber="67" pagination="72 - 101" refId="ref102147" refString="Matthews, A. (1887 - 88) Fam. Silphidae. In: Biologia Centrali-Americana. Insects. Coleoptera. Vol. 2 (1). Taylor &amp; Francis, London, pp. 72 - 101, pl. 3." type="journal article" year="1887">Matthews, 1887: 74</bibRefCitation>
</treatmentCitation>
for
<collectingCountry id="BEF1AAB9FFB9871AFB65CF89FB0AF2B0" box="[1159,1245,189,215]" name="Mexico" pageId="66" pageNumber="67">Mexico</collectingCountry>
(Jacala,
<collectingRegion id="042224CBFFB9871AFAA0CF89FECAF29C" country="Mexico" name="Hidalgo" pageId="66" pageNumber="67">Hidalgo State</collectingRegion>
) is in error (a misidentification, in
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<bibRefCitation id="A27797D8FFB9871AFD46CFD5FC2CF29C" author="Miller, K. B. &amp; Wheeler, Q. D." box="[676,1019,224,251]" pageId="66" pageNumber="67" pagination="1 - 167" refId="ref102247" refString="Miller, K. B. &amp; Wheeler, Q. D. (2005 b) Slime-mold beetles of the genus Agathidium Panzer in North and Central America, Part II. Coleoptera: Leiodidae. Bulletin of the American Museum of Natural History, 291, 1 - 167. https: // doi. org / 10.1206 / 0003 - 0090 (2005) 291 &lt;0001: SBOTGA&gt; 2.0. CO; 2" type="journal article" year="2005">Miller &amp; Wheeler, 2005b: 151</bibRefCitation>
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Biology: Usually in forested habitats and on slime molds (
<bibRefCitation id="A27797D8FFB9871AFCD9CE31FC1CF378" author="Hatch, M. H." box="[827,971,260,287]" pageId="66" pageNumber="67" pagination="1 - 384" refId="ref100508" refString="Hatch, M. H. (1957) The beetles of the Pacific Northwest. Part II. Staphyliniformia. University of Washington Publications in Biology, 16, i-ix + 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
;
<bibRefCitation id="A27797D8FFB9871AFC3BCE31FBA0F378" author="Russell, L. K." box="[985,1143,261,287]" pageId="66" pageNumber="67" pagination="1 - 9" refId="ref106044" refString="Russell, L. K. (1979) Beetles associated with slime molds (Mycetozoa) in Oregon and California (Coleoptera: Leiodidae, Sphindidae, Lathridiidae). Pan-Pacific Entomologist, 55, 1 - 9." type="journal article" year="1979">Russell, 1979</bibRefCitation>
;
<bibRefCitation id="A27797D8FFB9871AFB67CE31FAFDF378" author="Newton, A. F. Jr." box="[1157,1322,261,287]" pageId="66" pageNumber="67" pagination="302 - 353" refId="ref102466" refString="Newton, A. F. Jr. (1984) Mycophagy in Staphylinoidea (Coleoptera). In: Wheeler, Q. &amp; Blackwell, M. (Eds.), Fungus-insect relationships: perspectives in ecology and evolution. Columbia University Press, New York, pp. 302 - 353." type="book chapter" year="1984">Newton, 1984</bibRefCitation>
;
<bibRefCitation id="A27797D8FFB9871AFAD5CE31FEC9F324" author="Wheeler, Q. D." pageId="66" pageNumber="67" pagination="395 - 403" refId="ref109967" refString="Wheeler, Q. D. (1987) A new species of Agathidium associated with an &quot; epimycetic &quot; slime mold plasmodium on Pleurotus fungi (Coleoptera: Leiodidae-Myxomycetes: Physarales-Basidiomycetes: Tricholomataceae). Coleopterists Bulletin, 41, 395 - 403." type="journal article" year="1987">Wheeler, 1987</bibRefCitation>
,
<bibRefCitation id="A27797D8FFB9871AFECBCE1DFEB2F324" author="Wheeler, Q. D." box="[297,357,297,323]" pageId="66" pageNumber="67" pagination="1 - 46" refId="ref110012" refString="Wheeler, Q. D. (1990) Morphology and ontogeny of postembryonic larval Agathidium and Anisotoma (Coleoptera: Leiodidae). American Museum Novitates, 2986, 1 - 46." type="journal article" year="1990">1990</bibRefCitation>
).
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