457 lines
65 KiB
XML
457 lines
65 KiB
XML
<document id="3FDB6BF65DA5809880B2332663B94A6C" ID-CLB-Dataset="22056" ID-DOI="10.5852/ejt.2016.177" ID-GBIF-Dataset="33a17d32-75b4-423c-bc63-4bd22ae66a4f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832770" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589832188929" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2016" docId="0137952B7533CB176A17C54DFC2543F7" docLanguage="en" docName="ejt-2016-177.pdf.imf" docOrigin="European Journal of Taxonomy 177" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Geotypodon millemanus Enghoff, 2016, gen. et sp. nov." docType="treatment" docUuid="D65889F9-1063-4FBB-AEBB-A594052220B1" docUuidSource="ZooBank" docVersion="9" lastPageNumber="12" masterDocId="FD0EED537534CB1C6847C63BFFC9414A" masterDocTitle="A mountain of millipedes III: A new genus for three new species from the Udzungwa Mountains and surroundings, Tanzania, as well as several ‘ orphaned’ species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="8" updateTime="1698840199972" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6DE362C8279F7775BD4689D95E35FDA8">A mountain of millipedes III: A new genus for three new species from the Udzungwa Mountains and surroundings, Tanzania, as well as several ‘ orphaned’ species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:name id="BBF0DBC2D43A5435B53D70AE4C5004E5" type="personal">
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<mods:roleTerm id="1304FFE903CFFAD74516130B08600C31">Author</mods:roleTerm>
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<mods:namePart id="3EC5DA21C6D4FD26BF4CA8A4956C32D7">Enghoff, Henrik</mods:namePart>
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<mods:title id="AB614F5FD1E7BCB7BCFF51824AD99342">European Journal of Taxonomy</mods:title>
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<mods:part id="F02146FF34B3F1067116B144F3722F9A">
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<mods:date id="85E84B5243298B57B11E82349822EBA7">2016</mods:date>
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<mods:number id="7715D6474D2B54B6975B8A4FB70E1EE8">2016-02-26</mods:number>
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<mods:number id="2A195134234F41B036076A459F537BAC">177</mods:number>
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<treatment id="0137952B7533CB176A17C54DFC2543F7" ID-DOI="http://doi.org/10.5281/zenodo.3850247" ID-GBIF-Taxon="164167485" ID-Zenodo-Dep="3850247" LSID="urn:lsid:zoobank.org:act:D65889F9-1063-4FBB-AEBB-A594052220B1" httpUri="http://treatment.plazi.org/id/0137952B7533CB176A17C54DFC2543F7" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8">
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<subSubSection id="C18477B67533CB1B6A17C54DFB2042FE" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8921243D7533CB1B6A17C54DFBE742DB" blockId="7.[404,1257,886,983]" box="[592,1070,886,913]" pageId="7" pageNumber="8">
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<heading id="D26993517533CB1B6A17C54DFBE742DB" box="[592,1070,886,913]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
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<taxonomicName id="4E9E5FBE7533CB1B6A17C54DFCBF42DB" ID-CoL="3FTNP" authorityName="Enghoff" authorityYear="2016" box="[592,886,886,913]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="millemanus" status="gen. et sp. nov.">
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<emphasis id="BBEAF82F7533CB1B6A17C54DFCBF42DB" bold="true" box="[592,886,886,913]" italics="true" pageId="7" pageNumber="8">Geotypodon millemanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0D945547533CB1B6B39C54CFBE742DB" box="[894,1070,887,913]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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urn:lsid:zoobank.org:act:
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<uuid id="FD381EE87533CB1B6AF6C5A2FB2042FE" box="[689,1257,921,948]" pageId="7" pageNumber="8">D65889F9-1063-4FBB-AEBB-A594052220B1</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C18477B67533CB1B6B4BC586FCB8429D" box="[780,881,957,983]" pageId="7" pageNumber="8" type="description">
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<paragraph id="8921243D7533CB1B6B4BC586FCB8429D" blockId="7.[404,1257,886,983]" box="[780,881,957,983]" pageId="7" pageNumber="8">
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<figureCitation id="11A538B87533CB1B6B4BC586FCB8429D" box="[780,881,957,983]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig 4" captionStartId-0="1.[227,270,1894,1920]" captionStartId-1="7.[225,268,2002,2028]" captionStartId-2="8.[227,270,1966,1992]" captionStartId-3="9.[225,266,1790,1816]" captionTargetBox-0="[236,1429,925,1845]" captionTargetBox-1="[227,1435,1194,1972]" captionTargetBox-2="[329,1330,1159,1900]" captionTargetBox-3="[231,1427,302,1763]" captionTargetId-0="figure@1.[227,1436,918,1853]" captionTargetId-1="figure@7.[227,1436,1162,1972]" captionTargetId-2="figure@8.[301,1353,1155,1914]" captionTargetId-3="figure@9.[231,1428,302,1764]" captionTargetPageId-0="1" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="Fig. 1. Map of the Udzungwa Mountains, showing the collecting sites for the three new Geotypodon species, as well as names of the Forest Reserves in question and names of individual mountains in West Kilombero FR. Red diamonds = G. millemanus gen. et sp. nov., yellow dot: G. submontanus gen. et sp. nov., blue triangle: G. iringensis gen. et sp. nov. Based on fig. 1 in Marshall et al. (2010) and information in Doody et al. (2001)." captionText-1="Fig. 2. Geotypodon millemanus gen. et sp. nov., paratype from West Kilombero Scarp FR after nine years in alcohol. Photograph by N. Ioannou." captionText-2="Fig. 3. Body size of males of Geotypodon spp. Bold symbols indicate numbers of podous rings and midbody vertical diameter of the new species described here. Small circles and shaded areas indicate" captionText-3="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3832772" figureDoi-1="http://doi.org/10.5281/zenodo.3832774" figureDoi-2="http://doi.org/10.5281/zenodo.3832776" figureDoi-3="http://doi.org/10.5281/zenodo.3832778" httpUri-0="https://zenodo.org/record/3832772/files/figure.png" httpUri-1="https://zenodo.org/record/3832774/files/figure.png" httpUri-2="https://zenodo.org/record/3832776/files/figure.png" httpUri-3="https://zenodo.org/record/3832778/files/figure.png" pageId="7" pageNumber="8">Figs 1–4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C18477B67533CB1468A6C5C5FCB340FF" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="8921243D7533CB1B68A6C5C5FE934552" blockId="7.[225,346,1022,1048]" box="[225,346,1022,1048]" pageId="7" pageNumber="8">
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<heading id="D26993517533CB1B68A6C5C5FE934552" bold="true" box="[225,346,1022,1048]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
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<emphasis id="BBEAF82F7533CB1B68A6C5C5FE934552" bold="true" box="[225,346,1022,1048]" pageId="7" pageNumber="8">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8921243D7533CB1468A6C216FCB340FF" blockId="7.[225,1437,1069,1131]" lastBlockId="8.[227,1436,305,437]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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A species of
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<taxonomicName id="4E9E5FBE7533CB1B693CC216FDC3450D" authorityName="Enghoff" authorityYear="2016" box="[379,522,1069,1095]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="BBEAF82F7533CB1B693CC216FDC3450D" box="[379,522,1069,1095]" italics="true" pageId="7" pageNumber="8">Geotypodon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0D945547533CB1B6A54C216FDB6450D" box="[531,639,1069,1095]" pageId="7" pageNumber="8" rank="genus">gen. nov.</taxonomicNameLabel>
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in which the gonopod coxal metaplica is produced mesad in a thin lamella, a small longitudinal lamella, perpendicular to the other, is present on the anterior part of the metaplica, the telomere is apically divided into two lobes of approximately equal size, and the limbus lobes are multi-cusped, hand-like. It shares these characters with
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<taxonomicName id="4E9E5FBE753CCB146BACC76EFB624024" authorityName="Enghoff" authorityYear="2016" box="[1003,1195,340,367]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="submontanus" status="gen. et sp. nov.">
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<emphasis id="BBEAF82F753CCB146BACC76EFB624024" box="[1003,1195,340,367]" italics="true" pageId="8" pageNumber="9">G. submontanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0D94554753CCB146CF4C76EFAA34025" box="[1203,1386,341,367]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, but differs from that species in larger size, straight mesal margin of metaplical lamella and absence of a spine-like process on the posterior apical telomeral lobe.
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</paragraph>
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</subSubSection>
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<caption id="DDE174B57533CB1B68A6C1E9FD20495A" ID-DOI="http://doi.org/10.5281/zenodo.3832774" ID-Zenodo-Dep="3832774" httpUri="https://zenodo.org/record/3832774/files/figure.png" pageId="7" pageNumber="8" startId="7.[225,268,2002,2028]" targetBox="[227,1435,1194,1972]" targetPageId="7">
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<paragraph id="8921243D7533CB1B68A6C1E9FD20495A" blockId="7.[225,1437,2002,2064]" pageId="7" pageNumber="8">
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<emphasis id="BBEAF82F7533CB1B68A6C1E9FEF846A6" bold="true" box="[225,305,2002,2028]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
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<taxonomicName id="4E9E5FBE7533CB1B697BC1E9FD9546A6" authorityName="Enghoff" authorityYear="2016" box="[316,604,2002,2028]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="millemanus" status="gen. et sp. nov.">
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<emphasis id="BBEAF82F7533CB1B697BC1E9FD9546A6" box="[316,604,2002,2028]" italics="true" pageId="7" pageNumber="8">Geotypodon millemanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0D945547533CB1B6A21C1E8FCE846A7" box="[614,801,2003,2029]" pageId="7" pageNumber="8" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, paratype from West Kilombero Scarp FR after nine years in alcohol. Photograph by N. Ioannou.
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</paragraph>
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</caption>
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<subSubSection id="C18477B6753CCB1468A4C7E7FCA54945" pageId="8" pageNumber="9" type="etymology">
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<paragraph id="8921243D753CCB1468A4C7E7FEA340BC" blockId="8.[227,362,476,502]" box="[227,362,476,502]" pageId="8" pageNumber="9">
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<heading id="D2699351753CCB1468A4C7E7FEA340BC" bold="true" box="[227,362,476,502]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
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<emphasis id="BBEAF82F753CCB1468A4C7E7FEA340BC" bold="true" box="[227,362,476,502]" pageId="8" pageNumber="9">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8921243D753CCB1468A4C430FC5843FB" blockId="8.[227,1437,523,689]" pageId="8" pageNumber="9">
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The name is a Latin noun in apposition, meaning “a thousand hands” and referring to the hand-like limbus lobes. A specimen with a diameter of
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<quantity id="4E6689D8753CCB146AAAC415FCF94302" box="[749,816,558,585]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="mm" value="5.0">5 mm</quantity>
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will have a body perimeter of approximately
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<quantity id="4E6689D8753CCB146D04C415FA514302" box="[1347,1432,558,585]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="mm" value="15.0">15 mm</quantity>
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. Assuming that the limbus covers only
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<quantity id="4E6689D8753CCB146AE0C46AFD354326" box="[679,764,593,620]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="mm" value="10.0">10 mm</quantity>
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, and taking into account that each “hand” occupies about 10 microns of the perimeter, each body ring will carry about a thousand “hands”, and a specimen with 60 body rings will therefore have roughly 60,000 “hands”.
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</paragraph>
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<paragraph id="8921243D753CCB1468A4C4E2FD9A43B9" blockId="8.[227,595,727,755]" box="[227,595,727,755]" pageId="8" pageNumber="9">
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<emphasis id="BBEAF82F753CCB1468A4C4E2FE7B43B9" bold="true" box="[227,434,729,755]" pageId="8" pageNumber="9">Material studied</emphasis>
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(total: 4 ♂♂)
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</paragraph>
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<materialsCitation id="39F62E60753CCB146957C533FE3242C6" ID-GBIF-Occurrence="2620529301" collectingDate="2000-11-18" collectionCode="ZMUC" collectorName="West Kilombero Scarp FR & Plot Paradiso & Frontier Tanzania" country="Tanzania" elevation="1400" latitude="-7.8440003" location="West Kilombero Scarp" longLatPrecision="1" longitude="36.371555" municipality="Udzungwa Mts" pageId="8" pageNumber="9" specimenCode="ZMUC00040350" specimenCount="1" stateProvince="Iringa" typeStatus="holotype">
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<paragraph id="8921243D753CCB146957C533FE4B4268" blockId="8.[227,1436,776,908]" box="[272,386,776,802]" pageId="8" pageNumber="9">
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<emphasis id="BBEAF82F753CCB146957C533FE4B4268" bold="true" box="[272,386,776,802]" pageId="8" pageNumber="9">
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<typeStatus id="56259A9F753CCB146957C533FE4B4268" box="[272,386,776,802]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8921243D753CCB1468A4C510FE3242C6" blockId="8.[227,1436,776,908]" pageId="8" pageNumber="9">
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<collectingCountry id="F18964AD753CCB1468A4C510FEB4420F" box="[227,381,811,837]" name="Tanzania" pageId="8" pageNumber="9">TANZANIA</collectingCountry>
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: ♂,
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<collectingRegion id="4B5AEADF753CCB1469E0C510FD83420F" box="[423,586,811,837]" country="Tanzania" name="Iringa" pageId="8" pageNumber="9">Iringa Region</collectingRegion>
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||
,
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<collectingRegion id="4B5AEADF753CCB146A14C510FD30420F" box="[595,761,811,837]" country="Tanzania" name="Iringa" pageId="8" pageNumber="9">Iringa District</collectingRegion>
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,
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<collectingMunicipality id="6945BE47753CCB146B45C510FC7F420F" box="[770,950,811,837]" pageId="8" pageNumber="9">Udzungwa Mts</collectingMunicipality>
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,
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<collectorName id="246B41EB753CCB146BF8C510FB38420F" box="[959,1265,811,837]" pageId="8" pageNumber="9">
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<location id="8C4172E6753CCB146BF8C510FB0C420F" LSID="urn:lsid:plazi:treatment:0137952B7533CB176A17C54DFC2543F7:8C4172E6753CCB146BF8C510FB0C420F" box="[959,1221,811,837]" country="Tanzania" latitude="-7.8440003" longLatPrecision="1" longitude="36.371555" municipality="Udzungwa Mts" name="West Kilombero Scarp" pageId="8" pageNumber="9" stateProvince="Iringa">West Kilombero Scarp</location>
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FR
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</collectorName>
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,
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<geoCoordinate id="ECAA42FA753CCB146CBDC511FA5E420F" box="[1274,1431,810,837]" degrees="07" direction="south" minutes="50" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="38.4" value="-7.8440003">07°50'38.4"S</geoCoordinate>
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,
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<geoCoordinate id="ECAA42FA753CCB1468A4C575FE414222" box="[227,392,846,872]" degrees="36" direction="east" minutes="22" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="17.6" value="36.371555">36°22'17.6" E</geoCoordinate>
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, montane forest,
|
||
<elevation id="02B3C30E753CCB146A08C575FCD14222" box="[591,792,846,872]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.41" metricValueMin="1.39" pageId="8" pageNumber="9" unit="m" value="1400.0" valueMax="1410.0" valueMin="1390.0">
|
||
<quantity id="4E6689D8753CCB146A08C575FD384222" box="[591,753,846,872]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.41" metricValueMin="1.39" pageId="8" pageNumber="9" unit="m" value="1400.0" valueMax="1410.0" valueMin="1390.0">1390–1410 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<collectorName id="246B41EB753CCB146B62C575FC0A4222" box="[805,963,846,872]" pageId="8" pageNumber="9">Plot Paradiso</collectorName>
|
||
, casual,
|
||
<date id="FD2002FD753CCB146C63C575FB0D4222" box="[1060,1220,846,872]" pageId="8" pageNumber="9" value="2000-11-18">
|
||
<collectingDate id="ED64FB15753CCB146C63C575FB0D4222" box="[1060,1220,846,872]" pageId="8" pageNumber="9" value="2000-11-18">18 Nov. 2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="246B41EB753CCB146C89C575FA554222" box="[1230,1436,846,872]" pageId="8" pageNumber="9">Frontier Tanzania</collectorName>
|
||
leg. (
|
||
<specimenCode id="D9388C46753CCB14695AC54AFE2642C6" box="[285,495,881,908]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="8" pageNumber="9">ZMUC00040350</specimenCode>
|
||
).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8921243D753CCB146957C588FE444287" blockId="8.[227,1436,947,1079]" box="[272,397,947,973]" pageId="8" pageNumber="9">
|
||
<emphasis id="BBEAF82F753CCB146957C588FE444287" bold="true" box="[272,397,947,973]" pageId="8" pageNumber="9">
|
||
<typeStatus id="56259A9F753CCB146957C588FE444287" box="[272,397,947,973]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8921243D753CCB1468A4C5EDFC2F457D" blockId="8.[227,1436,947,1079]" pageId="8" pageNumber="9">
|
||
<collectingCountry id="F18964AD753CCB1468A4C5EDFEB442BA" box="[227,381,982,1008]" name="Tanzania" pageId="8" pageNumber="9">TANZANIA</collectingCountry>
|
||
: 2 ♂♂, same data as
|
||
<typeStatus id="56259A9F753CCB146ACCC5EDFD3B42BA" box="[651,754,982,1008]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
(ZMUC100969, ZMUC00040345); 1 ♂,
|
||
<collectingRegion id="4B5AEADF753CCB146CA8C5EDFA5142BA" box="[1263,1432,982,1008]" country="Tanzania" name="Iringa" pageId="8" pageNumber="9">Iringa Region</collectingRegion>
|
||
,
|
||
<collectingRegion id="4B5AEADF753CCB1468A4C5C2FE434559" box="[227,394,1017,1043]" country="Tanzania" name="Iringa" pageId="8" pageNumber="9">Iringa District</collectingRegion>
|
||
, Udzungwa Mts, Kiranzi-Kitungulu FR,
|
||
<geoCoordinate id="ECAA42FA753CCB146B21C5C2FC024559" box="[870,971,1017,1043]" degrees="08" direction="south" minutes="09" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-8.15">08°09' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="ECAA42FA753CCB146B90C5C2FBF44559" box="[983,1085,1017,1044]" degrees="35" direction="east" minutes="05" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="35.083332">35°05' E</geoCoordinate>
|
||
, forest,
|
||
<quantity id="4E6689D8753CCB146CDEC5C2FB384559" box="[1177,1265,1017,1044]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="m" value="1500.0">1500 m</quantity>
|
||
asl,
|
||
<date id="FD2002FD753CCB146D62C5C2FA5C4559" box="[1317,1429,1017,1043]" pageId="8" pageNumber="9" value="1996-01">Jan. 1996</date>
|
||
, M. Andersen, P. Gravlund & A. Jakobsen leg. (ZMUC00046991).
|
||
</paragraph>
|
||
<caption id="DDE174B5753CCB1468A4C195FA5546A1" ID-DOI="http://doi.org/10.5281/zenodo.3832776" ID-Zenodo-Dep="3832776" httpUri="https://zenodo.org/record/3832776/files/figure.png" pageId="8" pageNumber="9" startId="8.[227,270,1966,1992]" targetBox="[329,1330,1159,1900]" targetPageId="8">
|
||
<paragraph id="8921243D753CCB1468A4C195FA5546A1" blockId="8.[227,1436,1966,2028]" pageId="8" pageNumber="9">
|
||
<emphasis id="BBEAF82F753CCB1468A4C195FEFA4682" bold="true" box="[227,307,1966,1992]" pageId="8" pageNumber="9">Fig. 3.</emphasis>
|
||
Body size of males of
|
||
<taxonomicName id="4E9E5FBE753CCB146A15C195FD284682" authorityName="Enghoff" authorityYear="2016" box="[594,737,1966,1992]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BBEAF82F753CCB146A15C195FD284682" box="[594,737,1966,1992]" italics="true" pageId="8" pageNumber="9">Geotypodon</emphasis>
|
||
</taxonomicName>
|
||
spp. Bold symbols indicate numbers of podous rings and midbody vertical diameter of the new species described here. Small circles and shaded areas indicate
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8921243D753CCB1468A4C1CEFCA54945" blockId="8.[227,876,2037,2063]" box="[227,876,2037,2063]" pageId="8" pageNumber="9">
|
||
published measurements for other
|
||
<taxonomicName id="4E9E5FBE753CCB146A3FC1CEFCCE4945" authorityName="Enghoff" authorityYear="2016" box="[632,775,2037,2063]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BBEAF82F753CCB146A3FC1CEFCCE4945" box="[632,775,2037,2063]" italics="true" pageId="8" pageNumber="9">Geotypodon</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DDE174B5753DCB1568A6C0C5FB254945" ID-DOI="http://doi.org/10.5281/zenodo.3832778" ID-Zenodo-Dep="3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="9" pageNumber="10" startId="9.[225,266,1790,1816]" targetBox="[231,1427,302,1763]" targetPageId="9">
|
||
<paragraph id="8921243D753DCB1568A6C0C5FB254945" blockId="9.[225,1437,1790,2063]" pageId="9" pageNumber="10">
|
||
<emphasis id="BBEAF82F753DCB1568A6C0C5FEE54652" bold="true" box="[225,300,1790,1816]" pageId="9" pageNumber="10">Fig 4.</emphasis>
|
||
<taxonomicName id="4E9E5FBE753DCB15697EC0C5FD954652" authorityName="Enghoff" authorityYear="2016" box="[313,604,1790,1816]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="millemanus" status="gen. et sp. nov.">
|
||
<emphasis id="BBEAF82F753DCB15697EC0C5FD954652" box="[313,604,1790,1816]" italics="true" pageId="9" pageNumber="10">Geotypodon millemanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A0D94554753DCB156A2FC0C4FCE34653" box="[616,810,1791,1817]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, paratype from Kiranzi-Kitungulu FR.
|
||
<emphasis id="BBEAF82F753DCB156D48C0C5FA8F4652" bold="true" box="[1295,1350,1790,1816]" pageId="9" pageNumber="10">A–E</emphasis>
|
||
: Right gonopod.
|
||
<emphasis id="BBEAF82F753DCB156913C11AFEA04671" bold="true" box="[340,361,1825,1851]" pageId="9" pageNumber="10">A</emphasis>
|
||
. Posterior view.
|
||
<emphasis id="BBEAF82F753DCB156A60C119FDF24676" bold="true" box="[551,571,1826,1852]" pageId="9" pageNumber="10">B</emphasis>
|
||
.Anterior view, telomere in red oval.
|
||
<emphasis id="BBEAF82F753DCB156BA6C11AFC3F4671" bold="true" box="[993,1014,1825,1851]" pageId="9" pageNumber="10">C</emphasis>
|
||
. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured.
|
||
<emphasis id="BBEAF82F753DCB156B3DC17EFC464615" bold="true" box="[890,911,1861,1887]" pageId="9" pageNumber="10">D</emphasis>
|
||
. (Anterior-)mesal view.
|
||
<emphasis id="BBEAF82F753DCB156CE8C17EFB0A4615" bold="true" box="[1199,1219,1861,1887]" pageId="9" pageNumber="10">E</emphasis>
|
||
. Telomere (part of coxa at lower left), basal (dorsal) view.
|
||
<emphasis id="BBEAF82F753DCB156AF6C153FD0A46C8" bold="true" box="[689,707,1896,1922]" pageId="9" pageNumber="10">F</emphasis>
|
||
. Limbus. Abbreviations:
|
||
<emphasis id="BBEAF82F753DCB156BAAC153FBC546C8" box="[1005,1036,1896,1922]" italics="true" pageId="9" pageNumber="10">atl</emphasis>
|
||
= anterior distal lobe of telomere;
|
||
<emphasis id="BBEAF82F753DCB1568A6C1B0FF3146EF" box="[225,248,1931,1957]" italics="true" pageId="9" pageNumber="10">bl</emphasis>
|
||
= basal lamella of telomere;
|
||
<emphasis id="BBEAF82F753DCB156A1DC1B0FDBB46EF" box="[602,626,1931,1957]" italics="true" pageId="9" pageNumber="10">itl</emphasis>
|
||
= intermediate distal lamella of telomere;
|
||
<emphasis id="BBEAF82F753DCB156C34C1B0FB4A46EF" box="[1139,1155,1931,1957]" italics="true" pageId="9" pageNumber="10">ll</emphasis>
|
||
= longitudinal lamella;
|
||
<emphasis id="BBEAF82F753DCB1568A6C194FF374683" box="[225,254,1967,1993]" italics="true" pageId="9" pageNumber="10">mf</emphasis>
|
||
= anteriad metaplical flange;
|
||
<emphasis id="BBEAF82F753DCB156A19C195FD434682" box="[606,650,1966,1992]" italics="true" pageId="9" pageNumber="10">mla</emphasis>
|
||
= metaplical lamella;
|
||
<emphasis id="BBEAF82F753DCB156BD5C194FC7F4682" box="[914,950,1967,1992]" italics="true" pageId="9" pageNumber="10">mp</emphasis>
|
||
= metaplica;
|
||
<emphasis id="BBEAF82F753DCB156C1FC194FB4E4682" box="[1112,1159,1967,1992]" italics="true" pageId="9" pageNumber="10">msp</emphasis>
|
||
= metaplical spine-like process;
|
||
<emphasis id="BBEAF82F753DCB15690FC1E8FEAC46A6" box="[328,357,2003,2028]" italics="true" pageId="9" pageNumber="10">pn</emphasis>
|
||
= posttorsal narrowing;
|
||
<emphasis id="BBEAF82F753DCB156AC2C1E8FD6B46A6" box="[645,674,2003,2028]" italics="true" pageId="9" pageNumber="10">pp</emphasis>
|
||
= proplica;
|
||
<emphasis id="BBEAF82F753DCB156B77C1E9FC8646A6" box="[816,847,2002,2028]" italics="true" pageId="9" pageNumber="10">ptl</emphasis>
|
||
= posterior distal lobe of telomere;
|
||
<emphasis id="BBEAF82F753DCB156CB5C1E8FADD46A6" box="[1266,1300,2003,2028]" italics="true" pageId="9" pageNumber="10">pts</emphasis>
|
||
= proximal telomeral spine;
|
||
<emphasis id="BBEAF82F753DCB1569EBC1CEFE1C4945" box="[428,469,2037,2063]" italics="true" pageId="9" pageNumber="10">slm</emphasis>
|
||
= solenomere;
|
||
<emphasis id="BBEAF82F753DCB156ACCC1CDFD524945" box="[651,667,2038,2063]" italics="true" pageId="9" pageNumber="10">tt</emphasis>
|
||
= torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C18477B6753ECB1668A4C70AFB9040D7" pageId="10" pageNumber="11" type="materials_examined">
|
||
<paragraph id="8921243D753ECB1668A4C70AFE4B4000" blockId="10.[227,386,304,331]" box="[227,386,304,331]" pageId="10" pageNumber="11">
|
||
<heading id="D2699351753ECB1668A4C70AFE4B4000" bold="true" box="[227,386,304,331]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="BBEAF82F753ECB1668A4C70AFE4B4000" bold="true" box="[227,386,304,331]" pageId="10" pageNumber="11">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C764FB9040D7" blockId="10.[227,1437,351,413]" pageId="10" pageNumber="11">
|
||
<collectingCountry id="F18964AD753ECB1668A4C764FEB24033" box="[227,379,351,377]" name="Tanzania" pageId="10" pageNumber="11">TANZANIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="4B5AEADF753ECB1669C0C764FDE54033" box="[391,556,351,377]" country="Tanzania" name="Iringa" pageId="10" pageNumber="11">Iringa Region</collectingRegion>
|
||
,
|
||
<collectingRegion id="4B5AEADF753ECB166A70C764FD294033" box="[567,736,351,377]" country="Tanzania" name="Iringa" pageId="10" pageNumber="11">Iringa District</collectingRegion>
|
||
, Udzungwa Mts, West Kilombero Scarp FR,
|
||
<geoCoordinate id="ECAA42FA753ECB166CB5C764FA5E4033" box="[1266,1431,351,378]" degrees="07" direction="south" minutes="50" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="38.4" value="-7.8440003">07°50'38.4" S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="ECAA42FA753ECB1668A4C7B9FE4040D7" box="[227,393,386,413]" degrees="36" direction="east" minutes="22" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="17.6" value="36.371555">36°22'17.6" E</geoCoordinate>
|
||
, montane forest,
|
||
<quantity id="4E6689D8753ECB166A14C7B8FD3F40D7" box="[595,758,387,413]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.41" metricValueMin="1.39" pageId="10" pageNumber="11" unit="m" value="1400.0" valueMax="1410.0" valueMin="1390.0">1390–1410 m</quantity>
|
||
asl, cf.
|
||
<bibRefCitation id="ED0F59CC753ECB166B15C7B9FB9C40D7" author="Marshall A. R. & Brink H. & Topp-Jorgensen J. E." box="[850,1109,386,413]" pageId="10" pageNumber="11" pagination="124 - 132" refId="ref8586" refString="Marshall A. R., Brink H. & Topp-Jorgensen J. E. 2001. Millipede Diversity and Distribution - West Kilombero Scarp Forest Reserve. In: Doody K. Z., Howell K. M., & Fanning E. (eds) West Kilombero Scarp Forest Reserve - Zoological Report: 124 - 132. Report for the Udzungwa Mountains Forest Management and Biodiversity Conservation Project, MEMA, Iringa, Tanzania." type="book chapter" year="2001">
|
||
Marshall
|
||
<emphasis id="BBEAF82F753ECB166B86C7B8FC3C40D6" box="[961,1013,386,412]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
|
||
. (2001)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C18477B6753ECB1668A4C7FFFA9A46EC" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="8921243D753ECB1668A4C7FFFEBD4094" blockId="10.[227,372,452,478]" box="[227,372,452,478]" pageId="10" pageNumber="11">
|
||
<heading id="D2699351753ECB1668A4C7FFFEBD4094" bold="true" box="[227,372,452,478]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
|
||
<emphasis id="BBEAF82F753ECB1668A4C7FFFEBD4094" bold="true" box="[227,372,452,478]" pageId="10" pageNumber="11">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C7C9FA464346" blockId="10.[227,1423,498,525]" box="[227,1423,498,525]" pageId="10" pageNumber="11">
|
||
SIZE. Length ca.
|
||
<quantity id="4E6689D8753ECB1669E2C7C9FE2B4347" box="[421,482,498,525]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="cm" value="9.0">9 cm</quantity>
|
||
. Diameter
|
||
<quantity id="4E6689D8753ECB166A25C7C9FD3B4347" box="[610,754,498,525]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.1" metricValueMin="4.8" pageId="10" pageNumber="11" unit="mm" value="4.949999999999999" valueMax="5.1" valueMin="4.8">4.8–5.1 mm</quantity>
|
||
. 60–66 podous rings, no apodous rings in front of telson.
|
||
</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C40FFB474391" blockId="10.[227,1437,564,731]" pageId="10" pageNumber="11">
|
||
COLOUR. After 15 years in alcohol somewhat faded, but pattern still evident. Upper part of head blackish, lower part yellowish. Overall colour of body ventrally and laterally yellowish. Posterior ca. 40% of metazona amber, in front of amber zone a blackish zone; blackish zone narrow laterally, becoming wider dorsally where extending onto posterior
|
||
<emphasis id="BBEAF82F753ECB166A88C4A6FCC243FD" bold="true" box="[719,779,669,696]" pageId="10" pageNumber="11">1/4–1/3</emphasis>
|
||
of prozona; resulting mid-dorsal dark band becoming narrower towards head. Telson blackish with yellowish margins, legs yellowish.
|
||
</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C538FDE74256" blockId="10.[227,558,770,797]" box="[227,558,770,797]" pageId="10" pageNumber="11">HEAD. Without peculiarities.</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C57FFC8C4217" blockId="10.[227,837,835,862]" box="[227,837,835,862]" pageId="10" pageNumber="11">COLLUM. With a marginal and a submarginal furrow.</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C5BEFEFB4289" blockId="10.[227,1436,901,963]" pageId="10" pageNumber="11">BODY RINGS. Almost perfect cylinders, not vaulted; suture straight; ozopores ca. three diameters behind suture.</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C5D1FCCD456D" blockId="10.[227,1436,1001,1063]" pageId="10" pageNumber="11">
|
||
LIMBUS (
|
||
<figureCitation id="11A538B8753ECB166901C5D2FE44454E" box="[326,397,1001,1028]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
|
||
). Consisting of isolated hand-like lobes of ca. 10 microns’ length, each with 3–6 ‘fingers’ and each with a ridge running along its length.
|
||
</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C275FD1145C1" blockId="10.[227,1437,1102,1164]" pageId="10" pageNumber="11">ANAL VALVES. Each with a well-developed dorsal spine and a smaller, yet distinctive ventral one; margin raised, with 3 setae not borne on tubercles.</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C288FD1A45BA" blockId="10.[227,1436,1201,1264]" pageId="10" pageNumber="11">
|
||
MALE LEGS. From 5
|
||
<superScript id="7EEB8975753ECB166982C28AFE1B45F5" attach="left" box="[453,466,1201,1215]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
|
||
pair with postfemoral and tibial pads on all legs, except last four pairs; tibial pads absent from several pairs in front of these.
|
||
</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C323FA8644AB" blockId="10.[227,1437,1303,1506]" pageId="10" pageNumber="11">
|
||
GONOPOD COXA (
|
||
<figureCitation id="11A538B8753ECB1669E4C32CFE254478" box="[419,492,1303,1330]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
|
||
). Slender, slightly sigmoid. Proplica (
|
||
<emphasis id="BBEAF82F753ECB166BE8C323FC05447B" box="[943,972,1304,1329]" italics="true" pageId="10" pageNumber="11">pp</emphasis>
|
||
) ending in small proplical lobe hidden behind anterior edge of metaplical lamella (
|
||
<emphasis id="BBEAF82F753ECB166AB3C300FCE9441F" box="[756,800,1339,1365]" italics="true" pageId="10" pageNumber="11">mla</emphasis>
|
||
), on
|
||
<figureCitation id="11A538B8753ECB166B25C300FC09441F" box="[866,960,1339,1365]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4B</figureCitation>
|
||
. Metaplica (
|
||
<emphasis id="BBEAF82F753ECB166C10C307FBB2441F" box="[1111,1147,1340,1365]" italics="true" pageId="10" pageNumber="11">mp</emphasis>
|
||
) with poorly developed basal anteriad flange (
|
||
<emphasis id="BBEAF82F753ECB1669A3C365FDC84432" box="[484,513,1374,1400]" italics="true" pageId="10" pageNumber="11">mf</emphasis>
|
||
), disto-mesally expanded into thin lamella (
|
||
<emphasis id="BBEAF82F753ECB166C42C365FBF84432" box="[1029,1073,1374,1400]" italics="true" pageId="10" pageNumber="11">mla</emphasis>
|
||
), with a straight mesal margin, tip of metaplica formed by blunt-triangular extension of
|
||
<emphasis id="BBEAF82F753ECB166B3AC3BAFC6044D1" box="[893,937,1409,1435]" italics="true" pageId="10" pageNumber="11">mla</emphasis>
|
||
; a second smaller longitudinal lamella (
|
||
<emphasis id="BBEAF82F753ECB166DC5C3BAFA5B44D1" box="[1410,1426,1409,1435]" italics="true" pageId="10" pageNumber="11">ll</emphasis>
|
||
) on anterior surface of metaplica partly covering proplical lobe; metaplica at level of proplical lobe, with a long, slightly curved spine-like process (
|
||
<emphasis id="BBEAF82F753ECB166A93C3F3FCCA44AB" box="[724,771,1480,1505]" italics="true" pageId="10" pageNumber="11">msp</emphasis>
|
||
) directed toward base of coxa on its anterior side.
|
||
</paragraph>
|
||
<paragraph id="8921243D753ECB1668A4C032FA9A46EC" blockId="10.[227,1437,1545,1959]" pageId="10" pageNumber="11">
|
||
GONOPOD TELOPODITE (
|
||
<figureCitation id="11A538B8753ECB1669A4C032FDE24769" box="[483,555,1545,1571]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
|
||
). Arculus 90°. Torsotope (
|
||
<emphasis id="BBEAF82F753ECB166B27C031FCB94769" box="[864,880,1546,1571]" italics="true" pageId="10" pageNumber="11">tt</emphasis>
|
||
) simple, compact, without processes (
|
||
<figureCitation id="11A538B8753ECB166D75C032FA464769" box="[1330,1423,1545,1571]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4B</figureCitation>
|
||
). Posttorsal narrowing (
|
||
<emphasis id="BBEAF82F753ECB1669ADC016FDCE470C" box="[490,519,1581,1606]" italics="true" pageId="10" pageNumber="11">pn</emphasis>
|
||
) pronounced, very slender, without processes or spines (
|
||
<figureCitation id="11A538B8753ECB166CEBC017FAC3470C" box="[1196,1290,1580,1606]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4B</figureCitation>
|
||
). Telopodite just distal to posttorsal narrowing dividing into slender, whip-like solenomere and complicated telomere. Solenomere (
|
||
<emphasis id="BBEAF82F753ECB1669C6C049FE6347C6" box="[385,426,1650,1676]" italics="true" pageId="10" pageNumber="11">slm</emphasis>
|
||
) curved in 3 dimensions (at least on preserved specimens), apically pointed, without any outgrowths (
|
||
<figureCitation id="11A538B8753ECB1669EFC0ADFD8047FA" box="[424,585,1686,1712]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4A, C, E</figureCitation>
|
||
). Efferent groove continuing from posttorsal narrowing onto solenomere and running all the way to its tip. Telomere with long, stout basal spine (
|
||
<emphasis id="BBEAF82F753ECB166C5DC081FBF54799" box="[1050,1084,1722,1747]" italics="true" pageId="10" pageNumber="11">pts</emphasis>
|
||
), spine first curving in parallel with solenomere, but then becoming straight and directed mesad (
|
||
<figureCitation id="11A538B8753ECB166B9BC0E7FB9047BC" box="[988,1113,1756,1782]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4A, C</figureCitation>
|
||
). Telomere close to
|
||
<emphasis id="BBEAF82F753ECB166D07C0E6FAAB47BC" box="[1344,1378,1757,1782]" italics="true" pageId="10" pageNumber="11">pts</emphasis>
|
||
with a basal lamella (
|
||
<emphasis id="BBEAF82F753ECB1669EEC0C4FE094653" box="[425,448,1791,1817]" italics="true" pageId="10" pageNumber="11">bl</emphasis>
|
||
) (
|
||
<figureCitation id="11A538B8753ECB16699BC0C4FDF44653" box="[476,573,1791,1817]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4A</figureCitation>
|
||
), followed by several complicated lamellar parts; basal part of telomere partly sheathing base of solenomere, distal part divided into two equally-sized thin lobes (
|
||
<emphasis id="BBEAF82F753ECB166D55C119FAF84676" box="[1298,1329,1826,1852]" italics="true" pageId="10" pageNumber="11">atl</emphasis>
|
||
and
|
||
<emphasis id="BBEAF82F753ECB166D2BC119FA424676" box="[1388,1419,1826,1852]" italics="true" pageId="10" pageNumber="11">ptl</emphasis>
|
||
), which initially diverge at ca. 90° but then curve towards each other (
|
||
<figureCitation id="11A538B8753ECB166C66C17DFBB7462A" box="[1057,1150,1862,1888]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4B</figureCitation>
|
||
, D–E). An intermediate lamella (
|
||
<emphasis id="BBEAF82F753ECB16690DC152FEAB46C9" box="[330,354,1897,1923]" italics="true" pageId="10" pageNumber="11">itl</emphasis>
|
||
) lodged in the space between
|
||
<emphasis id="BBEAF82F753ECB166A85C152FD2846C9" box="[706,737,1897,1923]" italics="true" pageId="10" pageNumber="11">atl</emphasis>
|
||
and
|
||
<emphasis id="BBEAF82F753ECB166B5FC152FC8D46C9" box="[792,836,1897,1923]" italics="true" pageId="10" pageNumber="11">plm</emphasis>
|
||
(
|
||
<figureCitation id="11A538B8753ECB166B13C152FC6646C9" box="[852,943,1897,1923]" captionStart="Fig 4" captionStartId="9.[225,266,1790,1816]" captionTargetBox="[231,1427,302,1763]" captionTargetId="figure@9.[231,1428,302,1764]" captionTargetPageId="9" captionText="Fig 4. Geotypodon millemanus gen. et sp. nov., paratype from Kiranzi-Kitungulu FR. A–E: Right gonopod. A. Posterior view. B.Anterior view, telomere in red oval. C. (Posterior-)mesal view, solenomere (yellow) and proximal telomeral spine (green) coloured. D. (Anterior-)mesal view. E. Telomere (part of coxa at lower left), basal (dorsal) view. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; bl = basal lamella of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mf = anteriad metaplical flange; mla = metaplical lamella; mp = metaplica; msp = metaplical spine-like process; pn = posttorsal narrowing; pp = proplica; ptl = posterior distal lobe of telomere; pts = proximal telomeral spine; slm = solenomere; tt = torsotope. Scales: A–E = 0.1 mm, F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.3832778" httpUri="https://zenodo.org/record/3832778/files/figure.png" pageId="10" pageNumber="11">Fig. 4D</figureCitation>
|
||
). Surfaces of
|
||
<emphasis id="BBEAF82F753ECB166C16C152FBB946C9" box="[1105,1136,1897,1923]" italics="true" pageId="10" pageNumber="11">atl</emphasis>
|
||
and
|
||
<emphasis id="BBEAF82F753ECB166CEFC152FB0E46C9" box="[1192,1223,1897,1923]" italics="true" pageId="10" pageNumber="11">ptl</emphasis>
|
||
facing each other, concave, margins of each lobe subparallel, smooth; telomere entirely without denticles or spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C18477B6753FCB1768A6C70BFC2543F7" pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph id="8921243D753FCB1768A6C70BFDDB4000" blockId="11.[225,530,304,331]" box="[225,530,304,331]" pageId="11" pageNumber="12">
|
||
<heading id="D2699351753FCB1768A6C70BFDDB4000" bold="true" box="[225,530,304,331]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="BBEAF82F753FCB1768A6C70BFDDB4000" bold="true" box="[225,530,304,331]" pageId="11" pageNumber="12">Distribution and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8921243D753FCB1768A6C75BFDDB40D6" blockId="11.[225,1436,351,413]" pageId="11" pageNumber="12">
|
||
Known from West Kilombero FR and Kiranza-Kitungulu FR. Altitudinal range:
|
||
<quantity id="4E6689D8753FCB176C85C764FAA14030" box="[1218,1384,351,378]" metricMagnitude="3" metricUnit="m" metricValue="1.3225" metricValueMax="1.5" metricValueMin="1.145" pageId="11" pageNumber="12" unit="m" value="1322.5" valueMax="1500.0" valueMin="1145.0">1145–1500 m</quantity>
|
||
asl. Haibtat: (montane) forest.
|
||
</paragraph>
|
||
<paragraph id="8921243D753FCB1768A6C7FFFE0A4094" blockId="11.[225,451,452,478]" box="[225,451,452,478]" pageId="11" pageNumber="12">
|
||
<heading id="D2699351753FCB1768A6C7FFFE0A4094" bold="true" box="[225,451,452,478]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="BBEAF82F753FCB1768A6C7FFFE0A4094" bold="true" box="[225,451,452,478]" pageId="11" pageNumber="12">Coexisting species</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8921243D753FCB1768A6C7C8FC2543F7" blockId="11.[225,1437,498,701]" pageId="11" pageNumber="12">
|
||
In Kiranza-Kitungulu FR
|
||
<taxonomicName id="4E9E5FBE753FCB176A56C7C8FD754346" authorityName="Enghoff" authorityYear="2016" box="[529,700,498,525]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="millemanus" status="gen. et sp. nov.">
|
||
<emphasis id="BBEAF82F753FCB176A56C7C8FD754346" box="[529,700,498,525]" italics="true" pageId="11" pageNumber="12">G. millemanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A0D94554753FCB176A83C7C8FCBD4347" box="[708,884,499,525]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
was found in the same sample as
|
||
<taxonomicName id="4E9E5FBE753FCB176D44C7C9FDC0437A" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="dabagaensis">
|
||
<emphasis id="BBEAF82F753FCB176D44C7C9FEBD437A" italics="true" pageId="11" pageNumber="12">Chaleponcus dabagaensis</emphasis>
|
||
Kraus, 1958
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4E9E5FBE753FCB176A07C42DFCB5437A" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[576,892,533,560]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gracilior">
|
||
<emphasis id="BBEAF82F753FCB176A07C42DFD03437A" box="[576,714,534,560]" italics="true" pageId="11" pageNumber="12">C. gracilior</emphasis>
|
||
<bibRefCitation id="ED0F59CC753FCB176A96C42EFCB5437A" author="Enghoff H." box="[721,892,533,560]" pageId="11" pageNumber="12" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
. In West Kilombero FR no other odontopygids were found in the same sample as
|
||
<taxonomicName id="4E9E5FBE753FCB176A3EC402FCEC4319" authorityName="Enghoff" authorityYear="2016" box="[633,805,569,595]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="millemanus" status="gen. et sp. nov.">
|
||
<emphasis id="BBEAF82F753FCB176A3EC402FCEC4319" box="[633,805,569,595]" italics="true" pageId="11" pageNumber="12">G. millemanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A0D94554753FCB176B6AC402FC2B4319" box="[813,994,569,595]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, but
|
||
<taxonomicName id="4E9E5FBE753FCB176C5BC402FB124319" authorityName="Enghoff" authorityYear="2016" box="[1052,1243,569,595]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="submontanus" status="gen. et sp. nov.">
|
||
<emphasis id="BBEAF82F753FCB176C5BC402FB124319" box="[1052,1243,569,595]" italics="true" pageId="11" pageNumber="12">G. submontanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A0D94554753FCB176CA4C402FA514319" box="[1251,1432,569,595]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4E9E5FBE753FCB1768A6C467FD63433C" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[225,682,604,630]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basiliscus">
|
||
<emphasis id="BBEAF82F753FCB1768A6C467FE3F433C" box="[225,502,604,630]" italics="true" pageId="11" pageNumber="12">Chaleponcus basiliscus</emphasis>
|
||
<bibRefCitation id="ED0F59CC753FCB1769B9C467FD63433C" author="Enghoff H." box="[510,682,604,630]" pageId="11" pageNumber="12" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4E9E5FBE753FCB176AF1C467FBF2433C" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[694,1083,604,630]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="circumvallatus">
|
||
<emphasis id="BBEAF82F753FCB176AF1C467FC4F433C" box="[694,902,604,630]" italics="true" pageId="11" pageNumber="12">C. circumvallatus</emphasis>
|
||
<bibRefCitation id="ED0F59CC753FCB176BC9C467FBF2433C" author="Enghoff H." box="[910,1083,604,630]" pageId="11" pageNumber="12" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4E9E5FBE753FCB176C01C467FB18433C" authorityName="Enghoff" authorityYear="2014" box="[1094,1233,604,630]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gracilior">
|
||
<emphasis id="BBEAF82F753FCB176C01C467FB18433C" box="[1094,1233,604,630]" italics="true" pageId="11" pageNumber="12">C. gracilior</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4E9E5FBE753FCB176CA7C467FED643D3" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ibis">
|
||
<emphasis id="BBEAF82F753FCB176CA7C467FAE4433C" box="[1248,1325,604,630]" italics="true" pageId="11" pageNumber="12">C. ibis</emphasis>
|
||
<bibRefCitation id="ED0F59CC753FCB176D72C467FED643D3" author="Enghoff H." pageId="11" pageNumber="12" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4E9E5FBE753FCB176974C4BBFD9643D3" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[307,607,639,666]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="netus">
|
||
<emphasis id="BBEAF82F753FCB176974C4BBFE5243D3" box="[307,411,640,666]" italics="true" pageId="11" pageNumber="12">C. netus</emphasis>
|
||
<bibRefCitation id="ED0F59CC753FCB1769EDC444FD9643D3" author="Enghoff H." box="[426,607,639,665]" pageId="11" pageNumber="12" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4E9E5FBE753FCB176A35C4BBFC5643D0" authority="Enghoff, 2015" authorityName="Enghoff" authorityYear="2015" box="[626,927,639,666]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tintin">
|
||
<emphasis id="BBEAF82F753FCB176A35C4BBFD1243D3" box="[626,731,640,666]" italics="true" pageId="11" pageNumber="12">C. tintin</emphasis>
|
||
<bibRefCitation id="ED0F59CC753FCB176AADC444FC5643D0" author="Enghoff H. & Frederiksen S. B." box="[746,927,639,666]" pageId="11" pageNumber="12" pagination="1 - 25" refId="ref8050" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4E9E5FBE753FCB176BF5C4BBFAAA43D0" authority="Enghoff 2015" authorityName="Enghoff" authorityYear="2015" box="[946,1379,639,666]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="longipala">
|
||
<emphasis id="BBEAF82F753FCB176BF5C4BBFB6043D3" box="[946,1193,639,666]" italics="true" pageId="11" pageNumber="12">Aquattuor longipala</emphasis>
|
||
<bibRefCitation id="ED0F59CC753FCB176CFEC444FAAA43D0" author="Enghoff H. & Frederiksen S. B." box="[1209,1379,639,666]" pageId="11" pageNumber="12" pagination="1 - 25" refId="ref8050" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4E9E5FBE753FCB1768A6C498FD9C43F7" authority="Enghoff, 2015" authorityName="Enghoff" authorityYear="2015" box="[225,597,674,701]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="udzungwensis">
|
||
<emphasis id="BBEAF82F753FCB1768A6C498FE6C43F6" box="[225,421,674,701]" italics="true" pageId="11" pageNumber="12">A. udzungwensis</emphasis>
|
||
<bibRefCitation id="ED0F59CC753FCB1769EBC499FD9C43F7" author="Enghoff H. & Frederiksen S. B." box="[428,597,674,701]" pageId="11" pageNumber="12" pagination="1 - 25" refId="ref8050" refString="Enghoff H. & Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
also occur in West Kilombero FR.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |