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<mods:title id="77B609F969C23F06F4A9FBEA2F50D13E">The Dryophthoridae of Costa Rica and Panama: Checklist with keys, new synonymy and descriptions of new species of Cactophagus, Mesocordylus, Metamasius and Rhodobaenus (Coleoptera; Curculionoidea)</mods:title>
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<paragraph id="89208B3514675B45FECCFA2BFD35EDD4" blockId="26.[264,653,1443,1510]" box="[264,653,1443,1470]" pageId="26" pageNumber="27">
<heading id="D2683C5914675B45FECCFA2BFD35EDD4" bold="true" box="[264,653,1443,1470]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<taxonomicName id="4E9FF0B614675B45FECCFA2BFD35EDD4" authority="Anderson" authorityName="Anderson" box="[264,653,1443,1470]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="dragoni">
<emphasis id="BBEB572714675B45FECCFA2BFD35EDD4" bold="true" box="[264,653,1443,1470]" pageId="26" pageNumber="27">
<emphasis id="BBEB572714675B45FECCFA2BFDB4EDD7" bold="true" box="[264,524,1444,1470]" italics="true" pageId="26" pageNumber="27">Cactophagus dragoni</emphasis>
Anderson
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="89208B3514675B45FECCFA43FE5DED8F" blockId="26.[264,653,1443,1510]" box="[264,485,1484,1510]" pageId="26" pageNumber="27">
(
<figureCitation id="11A497B014675B45FED5FA43FE33ED8F" box="[273,395,1484,1510]" captionStart="FIGURES 7 - 14" captionStartId="27.[264,382,1596,1619]" captionTargetBox="[272,1315,489,1569]" captionTargetId="figure@27.[272,1315,489,1569]" captionTargetPageId="27" captionText="FIGURES 7 - 14. Cactophagus species. 7 - 10. Cactophagus dragoni Anderson. 7. Dorsal habitus, immaculate form. 8. Lateral habitus, immaculate form. 9. Dorsal habitus, maculate form. 10. Lateral habitus, maculate form. 11 - 12. Cactophagus gasbarrinorum Anderson. 11. Dorsal habitus. 12. Lateral habitus. 13 - 14. Cactophagus lineatus Anderson. 13. Dorsal habitus. 14. Lateral habitus." httpUri="https://zenodo.org/record/168661/files/figure.png" pageId="26" pageNumber="27">Figs. 7­10</figureCitation>
, 
<figureCitation id="11A497B014675B45FE53FA43FE64ED8F" box="[407,476,1484,1510]" captionStart="FIGURES 15 ­ 18" captionStartId="28.[264,383,917,941]" captionTargetBox="[288,1300,473,890]" captionTargetId="figure@28.[287,1300,473,890]" captionTargetPageId="28" captionText="FIGURES 15 ­ 18. Cactophagus dragoni Anderson. 15. Lateral view of head, female. 16. Antenna. 17. Hind leg, male. 18. Pygidium, male." httpUri="https://zenodo.org/record/156037/files/figure.png" pageId="26" pageNumber="27">15­18</figureCitation>
)
</paragraph>
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<subSubSection id="C185D8BE14675B42FECCF993FBA6EE37" lastPageId="29" lastPageNumber="30" pageId="26" pageNumber="27" type="description">
<paragraph id="89208B3514675B44FECCF993FE15E9C4" blockId="26.[264,1324,1564,1870]" lastBlockId="27.[264,1323,284,429]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Identification.— 
<taxonomicName id="4E9FF0B614675B45FE0FF993FD74EE5C" box="[459,716,1564,1589]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="dragoni">
<emphasis id="BBEB572714675B45FE0FF993FD74EE5C" box="[459,716,1564,1589]" italics="true" pageId="26" pageNumber="27">Cactophagus dragoni</emphasis>
</taxonomicName>
appears to be closely related to 
<taxonomicName id="4E9FF0B614675B45FB90F993FB6BEE5C" box="[1108,1235,1564,1589]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="BBEB572714675B45FB90F993FB6BEE5C" box="[1108,1235,1564,1589]" italics="true" pageId="26" pageNumber="27">C. ornatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4E9FF0B614675B45FAD4F993FED2EE34" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="BBEB572714675B45FAD4F993FA9BEE5C" box="[1296,1315,1564,1589]" italics="true" pageId="26" pageNumber="27">C</emphasis>
. 
<emphasis id="BBEB572714675B45FECCF9CBFED2EE34" box="[264,362,1604,1629]" italics="true" pageId="26" pageNumber="27">sinuatus</emphasis>
</taxonomicName>
based on similarities in the placement of the antennae on the rostrum far from the eye (
<figureCitation id="11A497B014675B45FE87F9E3FE23EEEF" box="[323,411,1644,1670]" captionStart="FIGURES 15 ­ 18" captionStartId="28.[264,383,917,941]" captionTargetBox="[288,1300,473,890]" captionTargetId="figure@28.[287,1300,473,890]" captionTargetPageId="28" captionText="FIGURES 15 ­ 18. Cactophagus dragoni Anderson. 15. Lateral view of head, female. 16. Antenna. 17. Hind leg, male. 18. Pygidium, male." httpUri="https://zenodo.org/record/156037/files/figure.png" pageId="26" pageNumber="27">Fig. 15</figureCitation>
), the bilamellate apical portion of the peduncle of the postmentum which is somewhat projected anteriorly and ventrally, and the elytral maculations which (if present) distort the form of the elytal striae (
<figureCitation id="11A497B014675B45FD60F933FD54EEBF" box="[676,748,1724,1750]" captionStart="FIGURES 7 - 14" captionStartId="27.[264,382,1596,1619]" captionTargetBox="[272,1315,489,1569]" captionTargetId="figure@27.[272,1315,489,1569]" captionTargetPageId="27" captionText="FIGURES 7 - 14. Cactophagus species. 7 - 10. Cactophagus dragoni Anderson. 7. Dorsal habitus, immaculate form. 8. Lateral habitus, immaculate form. 9. Dorsal habitus, maculate form. 10. Lateral habitus, maculate form. 11 - 12. Cactophagus gasbarrinorum Anderson. 11. Dorsal habitus. 12. Lateral habitus. 13 - 14. Cactophagus lineatus Anderson. 13. Dorsal habitus. 14. Lateral habitus." httpUri="https://zenodo.org/record/168661/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
). Most specimens of 
<taxonomicName id="4E9FF0B614675B45FC2EF933FBD1EEBC" box="[1002,1129,1724,1749]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="dragoni">
<emphasis id="BBEB572714675B45FC2EF933FBD1EEBC" box="[1002,1129,1724,1749]" italics="true" pageId="26" pageNumber="27">C. dragoni</emphasis>
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have elytra with orange humeri and a series of disjunct orange maculations extended in an oblique arc from the anterior one­third of the lateral elytal margin to the posterior one­third of the elytral length at interval 3 (
<figureCitation id="11A497B014675B45FE33F8BBFDF8EF27" box="[503,576,1844,1870]" captionStart="FIGURES 7 - 14" captionStartId="27.[264,382,1596,1619]" captionTargetBox="[272,1315,489,1569]" captionTargetId="figure@27.[272,1315,489,1569]" captionTargetPageId="27" captionText="FIGURES 7 - 14. Cactophagus species. 7 - 10. Cactophagus dragoni Anderson. 7. Dorsal habitus, immaculate form. 8. Lateral habitus, immaculate form. 9. Dorsal habitus, maculate form. 10. Lateral habitus, maculate form. 11 - 12. Cactophagus gasbarrinorum Anderson. 11. Dorsal habitus. 12. Lateral habitus. 13 - 14. Cactophagus lineatus Anderson. 13. Dorsal habitus. 14. Lateral habitus." httpUri="https://zenodo.org/record/168661/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
). These maculations are tumescent and slightly distort the elytral striae. One immaculate conspecific specimen (near Boquete) is known (
<figureCitation id="11A497B014665B44FB6AFE93FAA1E95C" box="[1198,1305,284,309]" captionStart="FIGURES 7 - 14" captionStartId="27.[264,382,1596,1619]" captionTargetBox="[272,1315,489,1569]" captionTargetId="figure@27.[272,1315,489,1569]" captionTargetPageId="27" captionText="FIGURES 7 - 14. Cactophagus species. 7 - 10. Cactophagus dragoni Anderson. 7. Dorsal habitus, immaculate form. 8. Lateral habitus, immaculate form. 9. Dorsal habitus, maculate form. 10. Lateral habitus, maculate form. 11 - 12. Cactophagus gasbarrinorum Anderson. 11. Dorsal habitus. 12. Lateral habitus. 13 - 14. Cactophagus lineatus Anderson. 13. Dorsal habitus. 14. Lateral habitus." httpUri="https://zenodo.org/record/168661/files/figure.png" pageId="27" pageNumber="28">Figs. 7-8</figureCitation>
). Whereas similar, 
<taxonomicName id="4E9FF0B614665B44FE1EFECBFDEEE934" box="[474,598,324,349]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="BBEB572714665B44FE1EFECBFDEEE934" box="[474,598,324,349]" italics="true" pageId="27" pageNumber="28">C. ornatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4E9FF0B614665B44FD57FECBFCA1E934" box="[659,793,324,349]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="BBEB572714665B44FD57FECBFCA1E934" box="[659,793,324,349]" italics="true" pageId="27" pageNumber="28">C. sinuatus</emphasis>
</taxonomicName>
are not as markedly sculptured, have only a transverse subbasal pronotal impression, and the peduncle is not as sharply bilamellate as in 
<taxonomicName id="4E9FF0B614665B44FEE2FE1BFE1EE9C4" box="[294,422,404,429]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="dragoni">
<emphasis id="BBEB572714665B44FEE2FE1BFE1EE9C4" box="[294,422,404,429]" italics="true" pageId="27" pageNumber="28">C. dragoni</emphasis>
</taxonomicName>
.
</paragraph>
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<paragraph id="89208B3514665B44FECCF9B3FE97EEB7" blockId="27.[264,1323,1596,1758]" pageId="27" pageNumber="28">
<emphasis id="BBEB572714665B44FECCF9B3FE06EE3A" bold="true" box="[264,446,1596,1619]" pageId="27" pageNumber="28">FIGURES 7-14.</emphasis>
<taxonomicName id="4E9FF0B614665B44FE0AF9B2FDE3EE3D" box="[462,603,1597,1620]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB572714665B44FE0AF9B2FDE3EE3D" box="[462,603,1597,1620]" italics="true" pageId="27" pageNumber="28">Cactophagus</emphasis>
</taxonomicName>
species. 7-10. 
<taxonomicName id="4E9FF0B614665B44FCC1F9B2FBDCEE3A" authority="Anderson." authorityName="Anderson." box="[773,1124,1596,1620]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="dragoni">
<emphasis id="BBEB572714665B44FCC1F9B2FC56EE3D" box="[773,1006,1597,1620]" italics="true" pageId="27" pageNumber="28">Cactophagus dragoni</emphasis>
Anderson.
</taxonomicName>
7. Dorsal habitus, immaculate form. 8. Lateral habitus, immaculate form. 9. Dorsal habitus, maculate form. 10. Lateral habitus, maculate form. 11-12. 
<taxonomicName id="4E9FF0B614665B44FD5FF90DFBF2EEF0" authority="Anderson." authorityName="Anderson." box="[667,1098,1666,1689]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gasbarrinorum">
<emphasis id="BBEB572714665B44FD5FF90DFC6BEEF0" box="[667,979,1666,1689]" italics="true" pageId="27" pageNumber="28">Cactophagus gasbarrinorum</emphasis>
Anderson.
</taxonomicName>
11. Dorsal habitus. 12. Lateral habitus. 13-14. 
<taxonomicName id="4E9FF0B614665B44FDF7F92AFC29EED5" authority="Anderson." authorityName="Anderson." box="[563,913,1701,1724]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="BBEB572714665B44FDF7F92AFCA2EED5" box="[563,794,1701,1724]" italics="true" pageId="27" pageNumber="28">Cactophagus lineatus</emphasis>
Anderson.
</taxonomicName>
13. Dorsal habitus. 14. Lateral habitus.
</paragraph>
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Description. Male, length, 
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<date id="FD21ADF514665B44FDB0F89DFD76EF42" box="[628,718,1810,1835]" pageId="27" pageNumber="28" value="1915-08-13">13.8-15</date>
.3 mm
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; width, 
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. Female, length, 13.4-
</paragraph>
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14.4 mm; width, 6. 
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. Color black with elytral orange maculations at humeri, on Rostrum subequal to length pronotum; elongate, cylindrical, robust, evenly rather strongly curved, very coarsely, irregularly punctate basally, punctures somewhat finer, sparser apically; base of rostrum expanded slightly in dorsal view at point of antennal insertion only; in lateral view slightly wider at base but not obviously expanded, basal area long, about two­fifths total rostral length. Rostrum glabrous ventrally; peduncle markedly bilamellate apically, projected anteriorly and ventrally. Scrobe with posterior margin about 6 or 7 times width of base of scape from anterior margin of eyes. Antenna with scape more or less one­half length rostrum; club slightly laterally compressed, elongateoval; apical pilose part subequal to length basal glabrous part. Pronotum with lateral margins irregular, subparallel in basal one­half, convergent subapically, tubulate to apex; disc and flanks moderately deeply irregularly punctate throughout, some punctures coalescent forming irregular shallow rugae; surface irregular, with shallow impressions laterally at midlength and deep transverse impression subbasally at middle; anterolateral margin immediately behind eyes with surface somewhat crenulate. Pronotum slightly longer than wide. Elytra with length one and one­half times length pronotum; intervals finely punctate, irregularly creased; intervals 3­5 swollen and irregularly scupltured at base; striae with moderately deep, widely spaced, small punctures. Scutellum large, “V” shaped, length slightly longer than width at base, flat. Pygidium tumescent medially; shallowly regularly punctate throughout; apex broadly rounded, with transverse row of longer setae. Ventrally with front coxae very narrowly separated by no more than width of base of scape; prosternum markedly sloping between and anterior to front coxae, with low tubercle between coxae. Meso­, metasternum and ventrite 1 irregularly sparsely punctate; ventrites 2 to 5 sparsely, finely punctate; last ventrite flat, glabrous. Legs moderately long, moderately densely regularly punctate, especially on outer face of femora; front coxae of male with inwardly projected low rounded swelling, lacking in female; femora clavate, moderately long, hind femur reaching apex of pygidium; inner margins of middle and hind tibiae very slightly inwardly expanded subbasally; all tibiae with very fine, uniformly short, stout, peglike setae throughout length arranged in pectinate manner. Tarsi with article 1 of front leg pilose on both lobes; third article very widely dilated, pilose ventrally except for narrow median line in basal one­half; all tarsi with third articles symmetrical; apical margin of third articles very shallowly emarginate.
</paragraph>
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<th id="747EE00B1461A4A0FECCFE93FDDBE95F" box="[264,611,284,310]" gridcol="0" gridrow="0" pageId="28" pageNumber="29">intervals 10 and 11 at basal</th>
<th id="747EE00B1461A4A0FDAEFE93FD5FE95F" box="[618,743,284,310]" gridcol="1" gridrow="0" pageId="28" pageNumber="29">one­third,</th>
<th id="747EE00B1461A4A0FD34FE93FCDFE95F" box="[752,871,284,310]" gridcol="2" gridrow="0" pageId="28" pageNumber="29">and in an</th>
<th id="747EE00B1461A4A0FCAAFE93FC44E95F" box="[878,1020,284,310]" gridcol="3" gridrow="0" pageId="28" pageNumber="29">oblique arc</th>
<th id="747EE00B1461A4A0FBC1FE93FA94E95F" box="[1029,1324,284,310]" gridcol="4" gridrow="0" pageId="28" pageNumber="29">through intervals 5­7 at</th>
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<th id="747EE00B1461A4A0FECCFECBFDDBE937" box="[264,611,324,350]" gridcol="0" gridrow="1" pageId="28" pageNumber="29">midlength, to intervals 3 and</th>
<td id="747EE00B1461A4A0FDAEFECBFD5FE937" box="[618,743,324,350]" gridcol="1" gridrow="1" pageId="28" pageNumber="29">4 at apical</td>
<td id="747EE00B1461A4A0FD34FECBFCDFE937" box="[752,871,324,350]" gridcol="2" gridrow="1" pageId="28" pageNumber="29">one­third;</td>
<td id="747EE00B1461A4A0FCAAFECBFC44E937" box="[878,1020,324,350]" gridcol="3" gridrow="1" pageId="28" pageNumber="29">maculations</td>
<td id="747EE00B1461A4A0FBC1FECBFA94E937" box="[1029,1324,324,350]" gridcol="4" gridrow="1" pageId="28" pageNumber="29">
slightly swollen and 
<tr id="37AF89771461A4A0FECCFEE3FA94E9EF" box="[264,1324,364,390]" gridrow="2" pageId="28" pageNumber="29" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="747EE00B1461A4A0FECCFEE3FDDBE9EF" box="[264,611,364,390]" gridcol="0" gridrow="2" pageId="28" pageNumber="29">distorting elytral striae.</th>
</tr>
</td>
</tr>
</table>
</paragraph>
<caption id="DDE0DBBD14615B43FECCFC1AFD02EBA6" httpUri="https://zenodo.org/record/156037/files/figure.png" pageId="28" pageNumber="29" targetBox="[288,1300,473,890]" targetPageId="28">
<paragraph id="89208B3514615B43FECCFC1AFD02EBA6" blockId="28.[264,1324,917,975]" pageId="28" pageNumber="29">
<emphasis id="BBEB572714615B43FECCFC1AFE71EBC4" bold="true" box="[264,457,917,941]" pageId="28" pageNumber="29">FIGURES 15­18.</emphasis>
<taxonomicName id="4E9FF0B614615B43FE14FC19FC94EBC5" authority="Anderson." authorityName="Anderson." box="[464,812,917,941]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dragoni">
<emphasis id="BBEB572714615B43FE14FC19FD00EBC4" box="[464,696,918,941]" italics="true" pageId="28" pageNumber="29">Cactophagus dragoni</emphasis>
Anderson.
</taxonomicName>
15. Lateral view of head, female. 16. Antenna. 17. Hind leg, male. 18. Pygidium, male.
</paragraph>
</caption>
<paragraph id="89208B3514605B42FEFCFD0BFD61EA87" blockId="29.[264,1325,284,1630]" pageId="29" pageNumber="30">
Variation.—Some variation exists in the extent of the elytral orange maculations which are lacking on interval 4 and intervals 6 and 
<quantity id="4E6726D014605B42FC9AFD23FC33EAAF" box="[862,907,684,710]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="29" pageNumber="30" unit="in" value="7.0">7 in</quantity>
some specimens, and one immaculate specimen (near Boquete) is known.
</paragraph>
<paragraph id="89208B3514605B42FEFCFD73FEECEB0F" blockId="29.[264,1325,284,1630]" pageId="29" pageNumber="30">Sexual dimorphism.—Sexes are difficult to separate in this species. Males have a slightly more concave metasternum and ventrite 1, and possess small tubercles on the front coxae.</paragraph>
<paragraph id="89208B3514605B42FEFCFCFBFBE4ECA7" blockId="29.[264,1325,284,1630]" pageId="29" pageNumber="30">
Material Examined.—Male 
<typeStatus id="5624359714605B42FD47FCFBFCA6EBE7" box="[643,798,884,910]" pageId="29" pageNumber="30" type="holotype">HOLOTYPE</typeStatus>
labelled “Pan. Chiriqui Prov. / Cont'l Divide Trl. / 
<date id="FD21ADF514605B42FE8CFC13FE5EEBDF" box="[328,486,924,950]" pageId="29" pageNumber="30" value="1995-05-11" valueMax="1995-05-15" valueMin="1995-05-11">11­15.V.1995</date>
/ Morris &amp; Wappes&quot; (CMNC). Female ALLOTYPE labelled &quot; 
<collectingCountry id="F188CBA514605B42FB01FC13FA91EBDF" box="[1221,1321,924,950]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
. Chiriqui / 
<collectingCountry id="F188CBA514605B42FE43FC4BFE5DEBB7" box="[391,485,964,990]" name="Wallis and Futuna" pageId="29" pageNumber="30">Fortuna</collectingCountry>
, 
<geoCoordinate id="ECABEDF214605B42FE31FC4BFDE5EBB7" box="[501,605,964,990]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="-82.25">82°15'W</geoCoordinate>
/ 
<geoCoordinate id="ECABEDF214605B42FDBEFC4BFD77EBB7" box="[634,719,964,990]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="8.733334">8°44'N</geoCoordinate>
, 
<date id="FD21ADF514605B42FD1BFC4BFC32EBB7" box="[735,906,964,990]" pageId="29" pageNumber="30" value="1978-05-20">May 20, 1978</date>
/ O'Briens &amp; 
<collectingCountry id="F188CBA514605B42FBFFFC4BFB1FEBB7" box="[1083,1191,964,990]" name="Marshall Islands" pageId="29" pageNumber="30">Marshall</collectingCountry>
&quot; (CWOB). 
<typeStatus id="5624359714605B42FECCFC63FE08EC6F" box="[264,432,1004,1030]" pageId="29" pageNumber="30" type="paratype">PARATYPES</typeStatus>
(5, 3). 
<collectingCountry id="F188CBA514605B42FD82FC63FD7FEC6F" box="[582,711,1004,1030]" name="Panama" pageId="29" pageNumber="30">PANAMA</collectingCountry>
. CHIRIQUI PROVINCE. Boquete (
<quantity id="4E6726D014605B42FBB7FC63FB71EC6F" box="[1139,1225,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="29" pageNumber="30" unit="km" value="5.6">5.6 km</quantity>
. W.), La Culebra Trail, 
<quantity id="4E6726D014605B42FE76FB9BFDBDEC47" box="[434,517,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="29" pageNumber="30" unit="m" value="1450.0">1450m</quantity>
, 
<date id="FD21ADF514605B42FDD5FB9BFD29EC47" box="[529,657,1044,1070]" pageId="29" pageNumber="30" value="1996-06-19">19.vi.1996</date>
, J.Ashe &amp; R. Brooks, 
<taxonomicName id="4E9FF0B614605B42FC58FB9BFC65EC44" box="[924,989,1044,1069]" class="Liliopsida" family="Araceae" genus="Arum" kingdom="Plantae" order="Alismatales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="BBEB572714605B42FC58FB9BFC65EC44" box="[924,989,1044,1069]" italics="true" pageId="29" pageNumber="30">Arum</emphasis>
</taxonomicName>
leaf axils, #178 (1). Continental Divide Trail, 
<date id="FD21ADF514605B42FDC2FBB3FD1FEC3F" box="[518,679,1084,1110]" pageId="29" pageNumber="30" value="1995-05-11" valueMax="1995-05-15" valueMin="1995-05-11">11­15.V.1995</date>
, Morris and Wappes (1). 
<collectingCountry id="F188CBA514605B42FC33FBB3FBEDEC3F" box="[1015,1109,1084,1110]" name="Wallis and Futuna" pageId="29" pageNumber="30">Fortuna</collectingCountry>
, 
<geoCoordinate id="ECABEDF214605B42FBA6FBB3FB71EC3F" box="[1122,1225,1084,1110]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="-82.25">82°15'W</geoCoordinate>
<geoCoordinate id="ECABEDF214605B42FB15FBB3FA9DEC3F" box="[1233,1317,1084,1110]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="8.733334">8°44'N</geoCoordinate>
, 
<date id="FD21ADF514605B42FECCFBEBFE17EC17" box="[264,431,1124,1150]" pageId="29" pageNumber="30" value="1978-05-20">May 20, 1978</date>
, O'Briens &amp; 
<collectingCountry id="F188CBA514605B42FD8DFBEBFD09EC17" box="[585,689,1124,1150]" name="Marshall Islands" pageId="29" pageNumber="30">Marshall</collectingCountry>
(1). 
<date id="FD21ADF514605B42FCC1FBEBFC12EC17" box="[773,938,1124,1150]" pageId="29" pageNumber="30" value="1978-05-16">May 16, 1978</date>
(2; 1). Reserva Forestal La 
<collectingCountry id="F188CBA514605B42FECCFB03FEDEECCF" box="[264,358,1164,1190]" name="Wallis and Futuna" pageId="29" pageNumber="30">Fortuna</collectingCountry>
, Quebrada Aleman, 
<date id="FD21ADF514605B42FD9EFB03FD54ECCF" box="[602,748,1164,1190]" pageId="29" pageNumber="30" value="1995-07-21">21.VII.1995</date>
, C.W. and L.B. O'Brien (1). Cuenca 
<collectingCountry id="F188CBA514605B42FB0CFB03FA9EECCF" box="[1224,1318,1164,1190]" name="Wallis and Futuna" pageId="29" pageNumber="30">Fortuna</collectingCountry>
, Aoki Camp, 
<date id="FD21ADF514605B42FE5BFB3BFDA2ECA7" box="[415,538,1204,1230]" pageId="29" pageNumber="30" value="1985-05-23">23.V.1985</date>
, R. Flowers (1). 
<typeStatus id="5624359714605B42FCC4FB3BFCCAECA7" box="[768,882,1204,1230]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
in CMNC, CWOB.
</paragraph>
<paragraph id="89208B3514605B42FEFCFB53FD73EC9F" blockId="29.[264,1325,284,1630]" box="[312,715,1244,1270]" pageId="29" pageNumber="30">
Distribution.— 
<collectingCountry id="F188CBA514605B42FE2DFB53FDFFEC9F" box="[489,583,1244,1270]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
(Chiriqui).
</paragraph>
<paragraph id="89208B3514605B42FEFCFA8BFCFFED07" blockId="29.[264,1325,284,1630]" pageId="29" pageNumber="30">
Natural history.—Adults have been collected in montane cloud forest in the La 
<collectingCountry id="F188CBA514605B42FB3DFA8BFE82ED2F" name="Wallis and Futuna" pageId="29" pageNumber="30">Fortuna</collectingCountry>
area of Chiriqui province, 
<collectingCountry id="F188CBA514605B42FDBCFAA3FD61ED2F" box="[632,729,1324,1350]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
. The specimen from 
<quantity id="4E6726D014605B42FC13FAA3FB94ED2F" box="[983,1068,1324,1350]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="29" pageNumber="30" unit="km" value="5.6">5.6 km</quantity>
. W. Boquete was collected in the axil of a large 
<taxonomicName id="4E9FF0B614605B42FD83FADBFD3BED07" box="[583,643,1364,1390]" class="Liliopsida" family="Araceae" genus="Arum" kingdom="Plantae" order="Alismatales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">arum</taxonomicName>
plant (
<taxonomicName id="4E9FF0B614605B42FD10FADBFC83ED07" box="[724,827,1364,1390]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
).
</paragraph>
<paragraph id="89208B3514605B42FEFCFAF3FDC9ED8F" blockId="29.[264,1325,284,1630]" pageId="29" pageNumber="30">
Derivation of specific name.— Through support for the Nature Discovery Fund at the Canadian Museum of Nature, this species is named after Jane Dragon of Fort Smith, Northwest Territories, 
<collectingCountry id="F188CBA514605B42FDD6FA43FDD6ED8F" box="[530,622,1484,1510]" name="Canada" pageId="29" pageNumber="30">Canada</collectingCountry>
.
</paragraph>
<paragraph id="89208B3514605B42FEFCFA7BFBA6EE37" blockId="29.[264,1325,284,1630]" pageId="29" pageNumber="30">
Comments.—This species approaches species previously placed in 
<taxonomicName id="4E9FF0B614605B42FB8AFA7BFAB4EE64" box="[1102,1292,1524,1549]" class="Insecta" family="Dryophthoridae" genus="Cactophagoides" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB572714605B42FB8AFA7BFAB4EE64" box="[1102,1292,1524,1549]" italics="true" pageId="29" pageNumber="30">Cactophagoides</emphasis>
</taxonomicName>
in the extent of pronotal and elytral sculpture. Such intermediate forms have led to the consideration of 
<taxonomicName id="4E9FF0B614605B42FE67F9CBFDDAEE34" box="[419,610,1604,1629]" class="Insecta" family="Dryophthoridae" genus="Cactophagoides" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB572714605B42FE67F9CBFDDAEE34" box="[419,610,1604,1629]" italics="true" pageId="29" pageNumber="30">Cactophagoides</emphasis>
</taxonomicName>
as a junior synonym of 
<taxonomicName id="4E9FF0B614605B42FCB9F9CBFBA0EE34" box="[893,1048,1604,1629]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB572714605B42FCB9F9CBFBA0EE34" box="[893,1048,1604,1629]" italics="true" pageId="29" pageNumber="30">Cactophagus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>