treatments-xml/data/01/36/3A/01363A2314585B77FECCFAF3FD0BE9BF.xml

<document id="E793F60D7AD5168D99AA05441CA3C287" ID-DOI="10.5281/zenodo.156033" ID-GBIF-Dataset="3f262a5f-c86b-481f-a65b-e92affdb3c9c" ID-ISSN="1175­5326" ID-Zenodo-Dep="156033" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438414948" checkinUser="plazi" docAuthor="Anderson, Robert S." docDate="2002" docId="01363A2314585B77FECCFAF3FD0BE9BF" docLanguage="en" docName="zt00080.pdf" docOrigin="Zootaxa 80" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cactophagus riesenorum Anderson, new species" docType="treatment" docVersion="11" lastPageNumber="41" masterDocId="FD0F425B147D5B5FFFC4FF8FFFB8E869" masterDocTitle="The Dryophthoridae of Costa Rica and Panama: Checklist with keys, new synonymy and descriptions of new species of Cactophagus, Mesocordylus, Metamasius and Rhodobaenus (Coleoptera; Curculionoidea)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="38" updateTime="1698188049892" updateUser="plazi">
<mods:mods id="5C742A68BC4CC3B76F9D1E500403957B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="31E3E88CF994523AFBFA2BDE9AFE8C02">
<mods:title id="7893D28086680CE59726CF64129B94D4">The Dryophthoridae of Costa Rica and Panama: Checklist with keys, new synonymy and descriptions of new species of Cactophagus, Mesocordylus, Metamasius and Rhodobaenus (Coleoptera; Curculionoidea)</mods:title>
</mods:titleInfo>
<mods:name id="DA4F56ACB903393EDFE0359DA1228144" type="personal">
<mods:role id="AA23A4745667F1FD5B688452A05F07D9">
<mods:roleTerm id="67310869057D3EFBB564A0A51156DA5E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="640CF4C11BA34389219176190CA5B818">Anderson, Robert S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="FC83A408B1134235CCBE77434ABFCB5A">text</mods:typeOfResource>
<mods:relatedItem id="64D76BFC8ED53768BFC57D54862D87F7" type="host">
<mods:titleInfo id="A6D9C705531DD6827485DFF7B973AE61">
<mods:title id="D26ED2C31C46C75AA2159B29215A0D68">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C066829025CD1C2C729A9D087BAA1AE1">
<mods:date id="07B30D0305FD1A7F6A98E1C031A081C0">2002</mods:date>
<mods:detail id="EE602BCBA6309B7AC69DD818FCAA5253" type="volume">
<mods:number id="30E79D70A2D77AAEF98DA53C1B2B97B6">80</mods:number>
</mods:detail>
<mods:extent id="73B0F8CC7BA6C45E430642C75A63B3AD" unit="page">
<mods:start id="13E84A4FDF9D96B40A971E3911AF087F">1</mods:start>
<mods:end id="D5F923636BCDF384C9C3D54BAABD2673">94</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A142BC7B67081278D408E6CE7974E916">journal article</mods:classification>
<mods:identifier id="11F28AA75FE31DCD396FB417759EAE6B" type="DOI">10.5281/zenodo.156033</mods:identifier>
<mods:identifier id="2B1131B615B9A6035546FB12DF9918A1" type="GBIF-Dataset">3f262a5f-c86b-481f-a65b-e92affdb3c9c</mods:identifier>
<mods:identifier id="DC190C42C093832058220584F8A298FB" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="5EBD9B339F264C67413E2624BE7E58FF" type="Zenodo-Dep">156033</mods:identifier>
</mods:mods>
<treatment id="01363A2314585B77FECCFAF3FD0BE9BF" ID-DOI="http://doi.org/10.5281/zenodo.6277918" ID-GBIF-Taxon="119330192" ID-Zenodo-Dep="6277918" LSID="urn:lsid:plazi:treatment:01363A2314585B77FECCFAF3FD0BE9BF" httpUri="http://treatment.plazi.org/id/01363A2314585B77FECCFAF3FD0BE9BF" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38">
<subSubSection id="C185D8BE14585B7AFECCFAF3FE4DEDD7" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="89208B3514585B7AFECCFAF3FCE0EDFC" blockId="37.[264,856,1403,1470]" box="[264,856,1403,1430]" pageId="37" pageNumber="38">
<heading id="D2683C5914585B7AFECCFAF3FCE0EDFC" bold="true" box="[264,856,1403,1430]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
<emphasis id="BBEB572714585B7AFECCFAF3FCE0EDFC" bold="true" box="[264,856,1403,1430]" pageId="37" pageNumber="38">
<taxonomicName id="4E9FF0B614585B7AFECCFAF3FD03EDFC" authority="Anderson" authorityName="Anderson" box="[264,699,1403,1430]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="riesenorum" status="sp. nov.">
<emphasis id="BBEB572714585B7AFECCFAF3FD8FEDFF" bold="true" box="[264,567,1404,1430]" italics="true" pageId="37" pageNumber="38">Cactophagus riesenorum</emphasis>
Anderson
</taxonomicName>
, 
<taxonomicNameLabel id="A0D8EA5C14585B7AFD0CFAF4FCE0EDFC" box="[712,856,1403,1429]" pageId="37" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="89208B3514585B7AFECCFA2BFE4DEDD7" blockId="37.[264,856,1403,1470]" box="[264,501,1444,1470]" pageId="37" pageNumber="38">
(
<figureCitation id="11A497B014585B7AFED5FA2BFE22EDD7" box="[273,410,1444,1470]" captionStart="FIGURES 33 - 38" captionStartId="38.[264,382,1334,1357]" captionTargetBox="[279,1310,489,1310]" captionTargetId="figure@38.[279,1315,489,1310]" captionTargetPageId="38" captionText="FIGURES 33 - 38. Cactophagus species. 33 - 34. Cactophagus riesenorum Anderson. 33. Dorsal habitus. 34. Lateral habitus. 35 - 36. Cactophagus silron Anderson. 35. Dorsal habitus. 36. Lateral habitus. 37 - 38. Cactophagus sunatoriorum Anderson. 37. Dorsal habitus. 38. Lateral habitus." httpUri="https://zenodo.org/record/168663/files/figure.png" pageId="37" pageNumber="38">Figs. 33­34</figureCitation>
, 
<figureCitation id="11A497B014585B7AFE62FA2BFE54EDD7" box="[422,492,1444,1470]" captionStart="FIGURES 39 ­ 41" captionStartId="39.[264,383,1642,1666]" captionTargetBox="[553,1035,1091,1615]" captionTargetId="figure@39.[551,1037,1089,1615]" captionTargetPageId="39" captionText="FIGURES 39 ­ 41. Cactophagus riesenorum Anderson. 39. Lateral view of head, male. 40. Antenna. 41. Hind leg, male." httpUri="https://zenodo.org/record/156043/files/figure.png" pageId="37" pageNumber="38">39­41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C185D8BE14585B77FECCFA7BFD0BE9BF" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38" type="description">
<paragraph id="89208B3514585B79FECCFA7BFD92E934" blockId="37.[264,1324,1524,1870]" lastBlockId="38.[264,1323,284,429]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
Identification.—The only known specimen of 
<taxonomicName id="4E9FF0B614585B7AFCEBFA7BFC61EE64" box="[815,985,1524,1549]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="riesenorum">
<emphasis id="BBEB572714585B7AFCEBFA7BFC61EE64" box="[815,985,1524,1549]" italics="true" pageId="37" pageNumber="38">C. riesenorum</emphasis>
</taxonomicName>
can be easily recognized by the pattern of elytral coloration (single transverse basal red band) (
<figureCitation id="11A497B014585B7AFBD1F993FB26EE5F" box="[1045,1182,1564,1590]" captionStart="FIGURES 33 - 38" captionStartId="38.[264,382,1334,1357]" captionTargetBox="[279,1310,489,1310]" captionTargetId="figure@38.[279,1315,489,1310]" captionTargetPageId="38" captionText="FIGURES 33 - 38. Cactophagus species. 33 - 34. Cactophagus riesenorum Anderson. 33. Dorsal habitus. 34. Lateral habitus. 35 - 36. Cactophagus silron Anderson. 35. Dorsal habitus. 36. Lateral habitus. 37 - 38. Cactophagus sunatoriorum Anderson. 37. Dorsal habitus. 38. Lateral habitus." httpUri="https://zenodo.org/record/168663/files/figure.png" pageId="37" pageNumber="38">Figs. 33­34</figureCitation>
), tumescent front of metasternum, slightly sinuate basal margin of pronotum (
<figureCitation id="11A497B014585B7AFBC0F9CBFBE0EE37" box="[1028,1112,1604,1630]" captionStart="FIGURES 33 - 38" captionStartId="38.[264,382,1334,1357]" captionTargetBox="[279,1310,489,1310]" captionTargetId="figure@38.[279,1315,489,1310]" captionTargetPageId="38" captionText="FIGURES 33 - 38. Cactophagus species. 33 - 34. Cactophagus riesenorum Anderson. 33. Dorsal habitus. 34. Lateral habitus. 35 - 36. Cactophagus silron Anderson. 35. Dorsal habitus. 36. Lateral habitus. 37 - 38. Cactophagus sunatoriorum Anderson. 37. Dorsal habitus. 38. Lateral habitus." httpUri="https://zenodo.org/record/168663/files/figure.png" pageId="37" pageNumber="38">Fig. 33</figureCitation>
) and apically carinate pygidium. This species is most similar in color pattern to 
<taxonomicName id="4E9FF0B614585B7AFBC1F9E3FB7FEEEC" box="[1029,1223,1644,1669]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="aurofasciatus">
<emphasis id="BBEB572714585B7AFBC1F9E3FB7FEEEC" box="[1029,1223,1644,1669]" italics="true" pageId="37" pageNumber="38">C. aurofasciatus</emphasis>
</taxonomicName>
but that species is much broader in dorsal aspect, has a distinctly sinuate basal pronotal margin, has a virtually impunctate, short rostrum, has a straight upper edge of the antennal scrobe, and has a wider basal red band on the elytra. In rostral form, the species resembles 
<taxonomicName id="4E9FF0B614585B7AFAD4F96BFE26EF4C" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="duplocinctus">
<emphasis id="BBEB572714585B7AFAD4F96BFE26EF4C" italics="true" pageId="37" pageNumber="38">C. duplocinctus</emphasis>
</taxonomicName>
but that species has a distinctly sinuate basal pronotal margin and an anteriorly medially excised scutellum. Other characters distinguish 
<taxonomicName id="4E9FF0B614585B7AFBD7F8BBFB06EF24" box="[1043,1214,1844,1869]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="riesenorum">
<emphasis id="BBEB572714585B7AFBD7F8BBFB06EF24" box="[1043,1214,1844,1869]" italics="true" pageId="37" pageNumber="38">C. riesenorum</emphasis>
</taxonomicName>
from all members of 
<taxonomicName id="4E9FF0B6145B5B79FE61FE93FDF8E95C" box="[421,576,284,309]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB5727145B5B79FE61FE93FDF8E95C" box="[421,576,284,309]" italics="true" pageId="38" pageNumber="39">Cactophagus</emphasis>
</taxonomicName>
with an anteriorly tumescent metasternum (
<taxonomicName id="4E9FF0B6145B5B79FBA0FE93FA93E95C" box="[1124,1323,284,309]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="aurofasciatus">
<emphasis id="BBEB5727145B5B79FBA0FE93FA93E95C" box="[1124,1323,284,309]" italics="true" pageId="38" pageNumber="39">C. aurofasciatus</emphasis>
</taxonomicName>
group of Vaurie [1967]).
</paragraph>
<paragraph id="89208B35145B5B79FEFCFEE3FB25E9C4" blockId="38.[264,1323,284,429]" pageId="38" pageNumber="39">
Description. Male, length, 
<quantity id="4E6726D0145B5B79FD46FEE3FD57E9EC" box="[642,751,364,389]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="38" pageNumber="39" unit="mm" value="14.6">14.6 mm</quantity>
; width 
<quantity id="4E6726D0145B5B79FC90FEE3FC0AE9EC" box="[852,946,364,389]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="38" pageNumber="39" unit="mm" value="5.1">5.1 mm</quantity>
. Female not known. Color black, with elytra with distinct transverse basal red band in apical one-quarter.
</paragraph>
<caption id="DDE0DBBD145B5B79FECCFAB9FE97EDDC" httpUri="https://zenodo.org/record/168663/files/figure.png" pageId="38" pageNumber="39" targetBox="[279,1310,489,1310]" targetPageId="38">
<paragraph id="89208B35145B5B79FECCFAB9FE97EDDC" blockId="38.[264,1323,1334,1461]" pageId="38" pageNumber="39">
<emphasis id="BBEB5727145B5B79FECCFAB9FE75ED24" bold="true" box="[264,461,1334,1357]" pageId="38" pageNumber="39">FIGURES 33-38.</emphasis>
<taxonomicName id="4E9FF0B6145B5B79FE1BFAB9FDD4ED24" box="[479,620,1334,1357]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB5727145B5B79FE1BFAB9FDD4ED24" box="[479,620,1334,1357]" italics="true" pageId="38" pageNumber="39">Cactophagus</emphasis>
</taxonomicName>
species. 33-34. 
<taxonomicName id="4E9FF0B6145B5B79FCEDFAB9FB08ED24" authority="Anderson." authorityName="Anderson." box="[809,1200,1334,1357]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="riesenorum">
<emphasis id="BBEB5727145B5B79FCEDFAB9FB80ED24" box="[809,1080,1334,1357]" italics="true" pageId="38" pageNumber="39">Cactophagus riesenorum</emphasis>
Anderson.
</taxonomicName>
33. Dorsal habitus. 34. Lateral habitus. 35-36. 
<taxonomicName id="4E9FF0B6145B5B79FD59FAD6FC5DED19" authority="Anderson." authorityName="Anderson." box="[669,997,1369,1392]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="silron">
<emphasis id="BBEB5727145B5B79FD59FAD6FCD7ED19" box="[669,879,1369,1392]" italics="true" pageId="38" pageNumber="39">Cactophagus silron</emphasis>
Anderson.
</taxonomicName>
35. Dorsal habitus. 36. Lateral habitus. 37-38. 
<taxonomicName id="4E9FF0B6145B5B79FE2AFAF3FC31EDFA" authority="Anderson." authorityName="Anderson." box="[494,905,1404,1427]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sunatoriorum">
<emphasis id="BBEB5727145B5B79FE2AFAF3FCABEDFA" box="[494,787,1404,1427]" italics="true" pageId="38" pageNumber="39">Cactophagus sunatoriorum</emphasis>
Anderson.
</taxonomicName>
37. Dorsal habitus. 38. Lateral habitus.
</paragraph>
</caption>
<paragraph id="89208B35145B5B78FEFCF989FC79EC6F" blockId="38.[264,1323,1542,1887]" lastBlockId="39.[264,1325,284,1030]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">Rostrum slightly longer than pronotum; elongate, cylindrical, evenly curved, densely, shallowly punctate dorsally and laterally in basal one-third; apical two-thirds glabrous and virtually impunctate, shining; base of rostrum slightly expanded in dorsal view, basal expanded area short, about one-fifth total rostral length; in dorsal view with shallow median sulcus in basal expanded; in lateral view with deep impressed semi-circular area behind point of antennal insertion. Peduncle slightly convex, bilamellate. Scrobe with posterior margin about two times width of base of scape from anterior margin of eyes. Antenna with scape moderately long, about one-half length rostrum; club elongate-oval; apical pilose part subequal in length to basal glabrous part. Pronotum with lateral margins subparallel to slightly convergent in basal one­half, more markedly convergent at apical one­third, subapically constricted, tubulate to apex; moderately densely, moderately deeply, regularly punctate on disc and flanks; disc moderately deeply impressed subbasally at middle; hind margin of pronotum very slightly sinuate at middle. Elytra one and one­third times length of pronotum; intervals very finely, shallowly punctate, convex; striae deeply impressed, sparsely punctate; bases on intervals 2­5 not produced anteriorly. Scutellum small, “V” shaped, length slightly greater than width at base, concave. Pygidium deeply regularly punctate throughout; apex broadly acuminate; with distinct, apical median finely setose carina. Ventrally with front coxae separated by width of scape; prosternum moderately densely, irregularly punctate, flat but sloping anteriorly. Metasternum impunctate medially, with large irregular punctures laterally; anterior process of metasternum markedly prominent and tumescent at middle; venter otherwise with moderately deep, sparse punctures laterally on ventrites 1­5, on mesepisternum, mesepimeron, less so on metepisternum; last ventrite flat, deflexed ventrally at apex, apex narrowly impressed at middle. Legs moderate in length, moderately densely, shallowly punctate; femora clavate, hind femur longer, almost reaching middle of ventrite 5; front coxae with inner faces rounded; inner margins of tibiae straight, with row of short, peglike setae. Tarsi each with third article widely dilated, pilose ventrally except in narrow median line at base; all tarsi with third articles symmetrical and apical margins truncate.</paragraph>
<caption id="DDE0DBBD145A5B78FECCF9E5FDF2EECD" httpUri="https://zenodo.org/record/156043/files/figure.png" pageId="39" pageNumber="40" targetBox="[553,1035,1091,1615]" targetPageId="39">
<paragraph id="89208B35145A5B78FECCF9E5FDF2EECD" blockId="39.[264,1323,1642,1700]" pageId="39" pageNumber="40">
<emphasis id="BBEB5727145A5B78FECCF9E5FE77EEEB" bold="true" box="[264,463,1642,1666]" pageId="39" pageNumber="40">FIGURES 39­41.</emphasis>
<taxonomicName id="4E9FF0B6145A5B78FE1FF9E4FCD1EEE8" authority="Anderson." authorityName="Anderson." box="[475,873,1642,1666]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="riesenorum">
<emphasis id="BBEB5727145A5B78FE1FF9E4FD56EEEB" box="[475,750,1643,1666]" italics="true" pageId="39" pageNumber="40">Cactophagus riesenorum</emphasis>
Anderson.
</taxonomicName>
39. Lateral view of head, male. 40. Antenna. 41. Hind leg, male.
</paragraph>
</caption>
<paragraph id="89208B35145A5B78FEFCF957FCB4EF2B" blockId="39.[264,1324,1752,1898]" pageId="39" pageNumber="40">
Material examined.—Male 
<typeStatus id="56243597145A5B78FD46F957FCA5EE9B" box="[642,797,1752,1778]" pageId="39" pageNumber="40" type="holotype">HOLOTYPE</typeStatus>
labelled &quot;S. Irazuzta 2/16 / CR, Puntarenas Prov. / Monteverde 
<quantity id="4E6726D0145A5B78FE34F88FFDD0EF73" box="[496,616,1792,1818]" metricMagnitude="2" metricUnit="m" metricValue="9.07" metricValueMax="18.0" metricValueMin="0.13999999999999999" pageId="39" pageNumber="40" unit="m" value="907.0" valueMax="1800.0" valueMin="14.0">14­1800m</quantity>
/ 
<geoCoordinate id="ECABEDF2145A5B78FD44F88FFD5BEF73" box="[640,739,1792,1818]" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="925" value="10.3">10°18'N</geoCoordinate>
, 
<geoCoordinate id="ECABEDF2145A5B78FD2BF88FFCEEEF73" box="[751,854,1792,1818]" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="925" value="-84.8">84°48'W</geoCoordinate>
/ 1986&quot; (CMNC). Distribution.— 
<collectingCountry id="F188CBA5145A5B78FE2DF8A7FDD1EF2B" box="[489,617,1832,1858]" name="Costa Rica" pageId="39" pageNumber="40">Costa Rica</collectingCountry>
(Puntarenas).
</paragraph>
<paragraph id="89208B35145A5B78FEFCF8DFFBD7EF03" blockId="39.[264,1324,1752,1898]" box="[312,1135,1872,1898]" pageId="39" pageNumber="40">Natural history.—There is no information available on natural history.</paragraph>
<paragraph id="89208B3514555B77FEFCFE93FE76E9EF" blockId="40.[264,1323,284,470]" pageId="40" pageNumber="41">
Derivation of specific name.­ Through their support of the Nature Discovery Fund at the Canadian Museum of Nature, this species is named after the Riesen family of Calgary, Alberta, 
<collectingCountry id="F188CBA514555B77FEABFEE3FE73E9EF" box="[367,459,364,390]" name="Canada" pageId="40" pageNumber="41">Canada</collectingCountry>
.
</paragraph>
<paragraph id="89208B3514555B77FEFCFE1BFD0BE9BF" blockId="40.[264,1323,284,470]" pageId="40" pageNumber="41">
Comments.—The 
<typeStatus id="5624359714555B77FDD6FE1BFDC1E9C7" box="[530,633,404,430]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
was discovered in the student collections of the Estacion Biologica Monteverde in 
<date id="FD21ADF514555B77FDF0FE33FD17E9BF" box="[564,687,444,470]" pageId="40" pageNumber="41" value="2001-06">June 2001</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>