164 lines
20 KiB
XML
164 lines
20 KiB
XML
<document id="DDA032E9B73CCEA6E7E0E2F842E88C84" ID-DOI="10.5281/zenodo.156033" ID-GBIF-Dataset="3f262a5f-c86b-481f-a65b-e92affdb3c9c" ID-ISSN="11755326" ID-Zenodo-Dep="156033" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438414948" checkinUser="plazi" docAuthor="Anderson, Robert S." docDate="2002" docId="01363A2314545B74FECCFA43FD83EC47" docLanguage="en" docName="zt00080.pdf" docOrigin="Zootaxa 80" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cactophagus sunatoriorum Anderson, new species" docType="treatment" docVersion="12" lastPageNumber="44" masterDocId="FD0F425B147D5B5FFFC4FF8FFFB8E869" masterDocTitle="The Dryophthoridae of Costa Rica and Panama: Checklist with keys, new synonymy and descriptions of new species of Cactophagus, Mesocordylus, Metamasius and Rhodobaenus (Coleoptera; Curculionoidea)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="42" updateTime="1698188049892" updateUser="plazi">
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<mods:titleInfo id="7E3CEF7805C12EC4E98A81808C929956">
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<mods:title id="9E16D5C21BF4759CDC9FFE514FE6BD0B">The Dryophthoridae of Costa Rica and Panama: Checklist with keys, new synonymy and descriptions of new species of Cactophagus, Mesocordylus, Metamasius and Rhodobaenus (Coleoptera; Curculionoidea)</mods:title>
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<mods:namePart id="5AB74E54526AE32F46ED3470CD9A075E">Anderson, Robert S.</mods:namePart>
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<mods:title id="33C162DAA7B37FD9795A13B1C46AC3E8">Zootaxa</mods:title>
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<mods:part id="F22CFC9D169F60A2187F9C45FAF96A6F">
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<mods:date id="D4AB1A992DD9C5C79BA603CA46082FB7">2002</mods:date>
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<mods:number id="3C295AA714902503F9AB9842E256214E">80</mods:number>
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<mods:identifier id="73F76168125E767B7221CDC1C802A45D" type="DOI">10.5281/zenodo.156033</mods:identifier>
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<treatment id="01363A2314545B74FECCFA43FD83EC47" ID-DOI="http://doi.org/10.5281/zenodo.6277922" ID-GBIF-Taxon="119330191" ID-Zenodo-Dep="6277922" LSID="urn:lsid:zoobank.org:act:97043022-1C47-4BA8-AE68-962EC24C4610" httpUri="http://treatment.plazi.org/id/01363A2314545B74FECCFA43FD83EC47" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
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<subSubSection id="C185D8BE14545B76FECCFA43FE4CEE67" pageId="41" pageNumber="42" type="nomenclature">
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<paragraph id="89208B3514545B76FECCFA43FCCCED8C" blockId="41.[264,884,1483,1550]" box="[264,884,1483,1510]" pageId="41" pageNumber="42">
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<heading id="D2683C5914545B76FECCFA43FCCCED8C" bold="true" box="[264,884,1483,1510]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
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<emphasis id="BBEB572714545B76FECCFA43FCCCED8C" bold="true" box="[264,884,1483,1510]" pageId="41" pageNumber="42">
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<taxonomicName id="4E9FF0B614545B76FECCFA43FD6FED8C" authority="Anderson" authorityName="Anderson" box="[264,727,1483,1510]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sunatoriorum" status="sp. nov.">
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<emphasis id="BBEB572714545B76FECCFA43FDEBED8F" bold="true" box="[264,595,1484,1510]" italics="true" pageId="41" pageNumber="42">Cactophagus sunatoriorum</emphasis>
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Anderson
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</taxonomicName>
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,
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<taxonomicNameLabel id="A0D8EA5C14545B76FD20FA44FCCCED8C" box="[740,884,1483,1509]" pageId="41" pageNumber="42" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="89208B3514545B76FECCFA7BFE4CEE67" blockId="41.[264,884,1483,1550]" box="[264,500,1524,1550]" pageId="41" pageNumber="42">
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(
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<figureCitation id="11A497B014545B76FED5FA7BFE22EE67" box="[273,410,1524,1550]" captionStart="FIGURES 33 - 38" captionStartId="38.[264,382,1334,1357]" captionTargetBox="[279,1310,489,1310]" captionTargetId="figure@38.[279,1315,489,1310]" captionTargetPageId="38" captionText="FIGURES 33 - 38. Cactophagus species. 33 - 34. Cactophagus riesenorum Anderson. 33. Dorsal habitus. 34. Lateral habitus. 35 - 36. Cactophagus silron Anderson. 35. Dorsal habitus. 36. Lateral habitus. 37 - 38. Cactophagus sunatoriorum Anderson. 37. Dorsal habitus. 38. Lateral habitus." httpUri="https://zenodo.org/record/168663/files/figure.png" pageId="41" pageNumber="42">Figs. 3738</figureCitation>
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,
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<figureCitation id="11A497B014545B76FE62FA7BFE54EE67" box="[422,492,1524,1550]" captionStart="FIGURES 46 50" captionStartId="42.[264,383,975,999]" captionTargetBox="[278,1311,510,957]" captionTargetId="figure@42.[275,1311,508,957]" captionTargetPageId="42" captionText="FIGURES 46 50. Cactophagus sunatoriorum Anderson. 46. Lateral view of head, male. 47. Lateral view of head, female. 48. Antenna. 49. Hind leg, male. 50. Pygidium, male." httpUri="https://zenodo.org/record/156045/files/figure.png" pageId="41" pageNumber="42">4650</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C185D8BE14545B74FECCF9CBFD83EC47" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42" type="description">
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<paragraph id="89208B3514545B76FECCF9CBFC35EF27" blockId="41.[264,1323,1604,1870]" pageId="41" pageNumber="42">
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Identification.—
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<taxonomicName id="4E9FF0B614545B76FE0FF9CBFD49EE34" box="[459,753,1604,1629]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sunatorium">
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<emphasis id="BBEB572714545B76FE0FF9CBFD49EE34" box="[459,753,1604,1629]" italics="true" pageId="41" pageNumber="42">Cactophagus sunatorium</emphasis>
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</taxonomicName>
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bears a great resemblance to
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<taxonomicName id="4E9FF0B614545B76FB8CF9CBFB52EE34" box="[1096,1258,1604,1629]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="personatus">
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<emphasis id="BBEB572714545B76FB8CF9CBFB52EE34" box="[1096,1258,1604,1629]" italics="true" pageId="41" pageNumber="42">C. personatus</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4E9FF0B614545B76FAD4F9CBFE2CEEEC" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="rectistriatus">
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<emphasis id="BBEB572714545B76FAD4F9CBFE2CEEEC" italics="true" pageId="41" pageNumber="42">C. rectistriatus</emphasis>
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</taxonomicName>
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but is clearly distinct based upon the lack of ventral pilosity on the rostrum (as in
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<taxonomicName id="4E9FF0B614545B76FEE3F91BFE73EEC4" box="[295,459,1684,1709]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="personatus">
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<emphasis id="BBEB572714545B76FEE3F91BFE73EEC4" box="[295,459,1684,1709]" italics="true" pageId="41" pageNumber="42">C. personatus</emphasis>
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</taxonomicName>
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) by the more widely separated front coxae, and by the flat prosternum in males (tuberculate in males of
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<taxonomicName id="4E9FF0B614545B76FDB0F933FC9BEEBC" box="[628,803,1724,1749]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="rectistriatus">
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<emphasis id="BBEB572714545B76FDB0F933FC9BEEBC" box="[628,803,1724,1749]" italics="true" pageId="41" pageNumber="42">C. rectistriatus</emphasis>
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</taxonomicName>
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). Additionally, the humeri are orange in
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<taxonomicName id="4E9FF0B614545B76FAD4F933FE10EE94" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="sunatoriorum">
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<emphasis id="BBEB572714545B76FAD4F933FE10EE94" italics="true" pageId="41" pageNumber="42">C. sunatoriorum</emphasis>
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</taxonomicName>
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whereas they are black in
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<taxonomicName id="4E9FF0B614545B76FD22F96BFC32EE94" box="[742,906,1764,1789]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="personatus">
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<emphasis id="BBEB572714545B76FD22F96BFC32EE94" box="[742,906,1764,1789]" italics="true" pageId="41" pageNumber="42">C. personatus</emphasis>
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</taxonomicName>
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and infuscate subbasally in
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<taxonomicName id="4E9FF0B614545B76FB1DF96BFEC8EF4C" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="rectistriatus">
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<emphasis id="BBEB572714545B76FB1DF96BFEC8EF4C" italics="true" pageId="41" pageNumber="42">C. rectistriatus</emphasis>
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</taxonomicName>
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. The elytral maculations do not distort the striae as in
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<taxonomicName id="4E9FF0B614545B76FC33F883FBCEEF4C" box="[1015,1142,1804,1829]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dragoni">
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<emphasis id="BBEB572714545B76FC33F883FBCEEF4C" box="[1015,1142,1804,1829]" italics="true" pageId="41" pageNumber="42">C. dragoni</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4E9FF0B614545B76FB6BF883FA93EF4C" box="[1199,1323,1804,1829]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="ornatus">
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<emphasis id="BBEB572714545B76FB6BF883FA93EF4C" box="[1199,1323,1804,1829]" italics="true" pageId="41" pageNumber="42">C. ornatus</emphasis>
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</taxonomicName>
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and form a broader, more continuous arc (
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<figureCitation id="11A497B014545B76FD31F8BBFC38EF27" box="[757,896,1844,1870]" captionStart="FIGURES 33 - 38" captionStartId="38.[264,382,1334,1357]" captionTargetBox="[279,1310,489,1310]" captionTargetId="figure@38.[279,1315,489,1310]" captionTargetPageId="38" captionText="FIGURES 33 - 38. Cactophagus species. 33 - 34. Cactophagus riesenorum Anderson. 33. Dorsal habitus. 34. Lateral habitus. 35 - 36. Cactophagus silron Anderson. 35. Dorsal habitus. 36. Lateral habitus. 37 - 38. Cactophagus sunatoriorum Anderson. 37. Dorsal habitus. 38. Lateral habitus." httpUri="https://zenodo.org/record/168663/files/figure.png" pageId="41" pageNumber="42">Figs. 3738</figureCitation>
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).
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</paragraph>
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<paragraph id="89208B3514575B75FEFCFE93FCC7E9C7" blockId="42.[264,1323,284,430]" pageId="42" pageNumber="43">
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Description.— Male, length,
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<quantity id="4E6726D014575B75FD54FE93FD43E95F" box="[656,763,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="42" pageNumber="43" unit="mm" value="12.2">12.2 mm</quantity>
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; width,
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<quantity id="4E6726D014575B75FC9DFE93FC0EE95F" box="[857,950,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="42" pageNumber="43" unit="mm" value="5.6">5.6 mm</quantity>
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. Female, length, 11.212.0 mm; width,
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<quantity id="4E6726D014575B75FE99FECBFE55E937" box="[349,493,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="42" pageNumber="43" unit="mm" value="5.35" valueMax="5.5" valueMin="5.2">5.25.5 mm</quantity>
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. Color black with humeri and basal portion of interval 10 yelloworange, with broad orange band extending from interval 10 at basal onethird in a slightly oblique manner to sutural interval at about midlength.
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</paragraph>
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<caption id="DDE0DBBD14575B75FECCFC40FBD4EC61" httpUri="https://zenodo.org/record/156045/files/figure.png" pageId="42" pageNumber="43" targetBox="[278,1311,510,957]" targetPageId="42">
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<paragraph id="89208B3514575B75FECCFC40FBD4EC61" blockId="42.[264,1323,975,1032]" pageId="42" pageNumber="43">
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<emphasis id="BBEB572714575B75FECCFC40FE71EB8E" bold="true" box="[264,457,975,999]" pageId="42" pageNumber="43">FIGURES 4650.</emphasis>
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<taxonomicName id="4E9FF0B614575B75FE12FC40FCD7EB8F" authority="Anderson." authorityName="Anderson." box="[470,879,975,998]" class="Insecta" family="Dryophthoridae" genus="Cactophagus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="sunatoriorum">
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<emphasis id="BBEB572714575B75FE12FC40FD42EB8F" box="[470,762,975,998]" italics="true" pageId="42" pageNumber="43">Cactophagus sunatoriorum</emphasis>
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Anderson.
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</taxonomicName>
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46. Lateral view of head, male. 47. Lateral view of head, female. 48. Antenna. 49. Hind leg, male. 50. Pygidium, male.
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</paragraph>
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</caption>
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<paragraph id="89208B3514575B74FEFCFBB3FED0EA4F" blockId="42.[264,1324,1084,1870]" lastBlockId="43.[264,1324,284,1070]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">Rostrum subequal to length pronotum; elongate, cylindrical, moderately robust, evenly rather strongly curved, somewhat coarsely, irregularly punctate laterally at base, punctures much finer, sparser apically in male and throughout length in female; base of rostrum expanded in dorsal view at point of antennal insertion only; in lateral view more or less subequal in width throughout length, basal expanded area about onethird total rostral length. Rostrum glabrous ventrally; peduncle bilamellate, projected anteriorly and ventrally. Scrobe with posterior margin about 4 or 5 times width of base of scape from anterior margin of eyes. Antenna with scape slightly longer than length rostrum; club laterally compressed, elongateoval; apical pilose part very slightly shorter than length basal glabrous part. Pronotum with lateral margins slightly convergent in basal twothirds, convergent subapically, tubulate to apex; disc and flanks uniformly shallowly punctate throughout; with broad transverse subbasal impression at middle. Pronotum with length subequal to width. Elytra one and onehalf times length of pronotum; intervals impunctate, flat; striae with moderately deep, widely spaced, small punctures. Scutellum large, “V” shaped, length slightly longer than width at base, concave at base. Pygidium rounded medially, not tumescent; coarsely deeply regularly punctate throughout; apex broadly subtruncate, with transverse row of longer setae. Ventrally with front coxae widely separated by slightly more than onehalf width of coxa; prosternum flat between and anterior to front coxae. Meso, metasternum and ventrite 1 irregularly sparsely punctate; ventrites 2 to 5 sparsely, finely punctate; last ventrite flat, glabrous, with shallow apical depression. Legs moderately long, moderately densely regularly punctate, especially on outer face of femora; femora slightly clavate, moderately long, hind femur reaching apex of pygidium; inner margins of middle and hind tibiae straight; all tibiae with very fine, uniformly short, stout, peglike setae throughout length arranged in pectinate manner. Tarsi with third article very widely dilated, pilose ventrally in apical twothirds except for “vshaped” basal area; all tarsi with third articles symmetrical; apical margin of third articles truncate, straight.</paragraph>
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<paragraph id="89208B3514565B74FEFCFDBBFE4FEB57" blockId="43.[264,1324,284,1070]" pageId="43" pageNumber="44">
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Material Examined.—Male
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<typeStatus id="5624359714565B74FD45FDBBFCA3EA27" box="[641,795,564,590]" pageId="43" pageNumber="44" type="holotype">HOLOTYPE</typeStatus>
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labelled “Pan: Chiriqui Prov. / Cont'l Divide Trl. /
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<date id="FD21ADF514565B74FE83FDD3FE5DEA1F" box="[327,485,604,630]" pageId="43" pageNumber="44" value="1999-05-11" valueMax="1999-05-15" valueMin="1999-05-11">1115.V.1999</date>
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/ Morris & Wappes" (CMNC). Female ALLOTYPE labelled "
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<collectingCountry id="F188CBA514565B74FB04FDD3FA9CEA1F" box="[1216,1316,604,630]" name="Panama" pageId="43" pageNumber="44">Panama</collectingCountry>
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: Chir. Res. / For. La
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<collectingCountry id="F188CBA514565B74FDD5FD0BFDD7EAF7" box="[529,623,644,670]" name="Wallis and Futuna" pageId="43" pageNumber="44">Fortuna</collectingCountry>
|
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, Cont. / Divide Tr.,
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<date id="FD21ADF514565B74FCBEFD0BFC57EAF7" box="[890,1007,644,670]" pageId="43" pageNumber="44" value="1995-07-20">7.20.1995</date>
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/ C.W. & L.B. O'Brien" (CWOB).
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<typeStatus id="5624359714565B74FE54FD23FD9FEAAF" box="[400,551,684,710]" pageId="43" pageNumber="44" type="paratype">PARATYPE</typeStatus>
|
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(1).
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<collectingCountry id="F188CBA514565B74FD40FD23FCBFEAAF" box="[644,775,684,710]" name="Panama" pageId="43" pageNumber="44">PANAMA</collectingCountry>
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. CHIRIQUI PROVINCE. Continental Divide Trail,
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<date id="FD21ADF514565B74FE8AFD5BFE55EA87" box="[334,493,724,750]" pageId="43" pageNumber="44" value="1999-05-12" valueMax="1999-05-14" valueMin="1999-05-12">1214.V.1999</date>
|
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(1).
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<typeStatus id="5624359714565B74FD8CFD5BFD16EA87" box="[584,686,724,750]" pageId="43" pageNumber="44" type="paratype">Paratype</typeStatus>
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in CMNC. There is also an additional specimen from
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<collectingCountry id="F188CBA514565B74FECCFD73FEDDEB7F" box="[264,357,764,790]" name="Panama" pageId="43" pageNumber="44">Panama</collectingCountry>
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Province, Cerro Campana,
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<date id="FD21ADF514565B74FD75FD73FC8EEB7F" box="[689,822,764,790]" pageId="43" pageNumber="44" value="1970-04-15">15.IV.1970</date>
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, H. Hespenheide, from which the rostrum has been broken off.
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</paragraph>
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<paragraph id="89208B3514565B74FEFCFCC3FC8FEB0F" blockId="43.[264,1324,284,1070]" box="[312,823,844,870]" pageId="43" pageNumber="44">
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Distribution.—
|
||
<collectingCountry id="F188CBA514565B74FE2DFCC3FDFFEB0F" box="[489,583,844,870]" name="Panama" pageId="43" pageNumber="44">Panama</collectingCountry>
|
||
(Chiriqui,
|
||
<collectingCountry id="F188CBA514565B74FD0CFCC3FC92EB0F" box="[712,810,844,870]" name="Panama" pageId="43" pageNumber="44">Panama</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="89208B3514565B74FEFCFCFBFCD6EBDF" blockId="43.[264,1324,284,1070]" pageId="43" pageNumber="44">
|
||
Natural history.—Three of the four known specimens were collected in cloud forest in the La
|
||
<collectingCountry id="F188CBA514565B74FE9EFC13FE0DEBDF" box="[346,437,924,950]" name="Wallis and Futuna" pageId="43" pageNumber="44">Fortuna</collectingCountry>
|
||
area of Chiriqui province in
|
||
<collectingCountry id="F188CBA514565B74FCCEFC13FCD3EBDF" box="[778,875,924,950]" name="Panama" pageId="43" pageNumber="44">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="89208B3514565B74FEFCFC4BFD83EC47" blockId="43.[264,1324,284,1070]" pageId="43" pageNumber="44">Derivation of specific name.—Through their support of the Nature Discovery Fund at the Canadian Museum of Nature, this species is named after the Sunatori family as a gift from Michèle Provencher.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |