treatments-xml/data/01/36/3A/01363A2314255B03FECCFA2BFE5EEB7F.xml

<document id="650E0A52CFEF81ADA58C6F5DE2B5DC02" ID-DOI="10.5281/zenodo.156033" ID-GBIF-Dataset="3f262a5f-c86b-481f-a65b-e92affdb3c9c" ID-ISSN="1175­5326" ID-Zenodo-Dep="156033" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438414948" checkinUser="plazi" docAuthor="Anderson, Robert S." docDate="2002" docId="01363A2314255B03FECCFA2BFE5EEB7F" docLanguage="en" docName="zt00080.pdf" docOrigin="Zootaxa 80" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Rhodobaenus patriciae Anderson, new species" docType="treatment" docVersion="12" lastPageNumber="93" masterDocId="FD0F425B147D5B5FFFC4FF8FFFB8E869" masterDocTitle="The Dryophthoridae of Costa Rica and Panama: Checklist with keys, new synonymy and descriptions of new species of Cactophagus, Mesocordylus, Metamasius and Rhodobaenus (Coleoptera; Curculionoidea)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="89" updateTime="1698188049892" updateUser="plazi">
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<mods:title id="C6D79E7F792A6D578D3666B2314EB5DA">The Dryophthoridae of Costa Rica and Panama: Checklist with keys, new synonymy and descriptions of new species of Cactophagus, Mesocordylus, Metamasius and Rhodobaenus (Coleoptera; Curculionoidea)</mods:title>
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<mods:date id="63F918FED34162494CBEAD7B814D4748">2002</mods:date>
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<treatment id="01363A2314255B03FECCFA2BFE5EEB7F" ID-DOI="http://doi.org/10.5281/zenodo.6277962" ID-GBIF-Taxon="119330211" ID-Zenodo-Dep="6277962" LSID="urn:lsid:zoobank.org:act:7D717B9A-EA93-418B-A83B-3B6974417301" httpUri="http://treatment.plazi.org/id/01363A2314255B03FECCFA2BFE5EEB7F" lastPageId="92" lastPageNumber="93" pageId="88" pageNumber="89">
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<paragraph id="89208B3514255B07FECCFA2BFCF8EDD4" blockId="88.[264,832,1443,1510]" box="[264,832,1443,1470]" pageId="88" pageNumber="89">
<heading id="D2683C5914255B07FECCFA2BFCF8EDD4" bold="true" box="[264,832,1443,1470]" fontSize="11" level="1" pageId="88" pageNumber="89" reason="1">
<emphasis id="BBEB572714255B07FECCFA2BFCF8EDD4" bold="true" box="[264,832,1443,1470]" pageId="88" pageNumber="89">
<taxonomicName id="4E9FF0B614255B07FECCFA2BFD1BEDD4" authority="Anderson" authorityName="Anderson" box="[264,675,1443,1470]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="patriciae" status="sp. nov.">
<emphasis id="BBEB572714255B07FECCFA2BFD98EDD7" bold="true" box="[264,544,1444,1470]" italics="true" pageId="88" pageNumber="89">Rhodobaenus patriciae</emphasis>
Anderson
</taxonomicName>
, 
<taxonomicNameLabel id="A0D8EA5C14255B07FD74FA2CFCF8EDD4" box="[688,832,1443,1469]" pageId="88" pageNumber="89" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="89208B3514255B07FECCFA43FD88ED8F" blockId="88.[264,832,1443,1510]" box="[264,560,1484,1510]" pageId="88" pageNumber="89">
(
<figureCitation id="11A497B014255B07FED5FA43FE00ED8F" box="[273,440,1484,1510]" captionStart="FIGURES 129 - 136" captionStartId="83.[264,382,1559,1582]" captionTargetBox="[276,1313,509,1512]" captionTargetId="figure@83.[276,1313,509,1518]" captionTargetPageId="83" captionText="FIGURES 129 - 136. Rhodobaenus species. 129 - 130. Rhodobaenus howelli Anderson. 129. Dorsal habitus. 130. Lateral habitus. 131 - 132. Rhodobaenus labrecheae Anderson. 131. Dorsal habitus. 132. Lateral habitus. 133 - 134. Rhodobaenus patriciae Anderson. 133. Dorsal habitus. 134. Lateral habitus. 135 - 136. Rhodobaenus tenorio Anderson. 135. Dorsal habitus. 136. Lateral habitus." httpUri="https://zenodo.org/record/168668/files/figure.png" pageId="88" pageNumber="89">Figs. 133­134</figureCitation>
, 
<figureCitation id="11A497B014255B07FE00FA43FD9FED8F" box="[452,551,1484,1510]" captionStart="FIGURES 147 ­ 150" captionStartId="89.[264,383,1106,1130]" captionTargetBox="[282,1293,586,1085]" captionTargetId="figure@89.[281,1293,585,1085]" captionTargetPageId="89" captionText="FIGURES 147 ­ 150. Rhodobaenus patriciae Anderson. 147. Lateral view of head, female. 148. Antenna. 149. Hind leg, female. 150. Pygidium, female." httpUri="https://zenodo.org/record/156064/files/figure.png" pageId="88" pageNumber="89">147­150</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C185D8BE14255B03FECCF993FE5EEB7F" lastPageId="92" lastPageNumber="93" pageId="88" pageNumber="89" type="description">
<paragraph id="89208B3514255B07FECCF993FB4AEF27" blockId="88.[264,1324,1564,1870]" pageId="88" pageNumber="89">
Identification.—This species is known only from three female specimens from the Osa Penninsula. 
<taxonomicName id="4E9FF0B614255B07FE5BF9CBFD08EE34" box="[415,688,1604,1629]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="BBEB572714255B07FE5BF9CBFD08EE34" box="[415,688,1604,1629]" italics="true" pageId="88" pageNumber="89">Rhodobaenus patriciae</emphasis>
</taxonomicName>
is similar to the southern South American 
<taxonomicName id="4E9FF0B614255B07FB63F9CBFECDEEEF" authority="Brethes" authorityName="Brethes" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="tornowii">
<emphasis id="BBEB572714255B07FB63F9CBFA92EE34" box="[1191,1322,1604,1629]" italics="true" pageId="88" pageNumber="89">R. tornowii</emphasis>
(Bréthes)
</taxonomicName>
in elytral color pattern (
<figureCitation id="11A497B014255B07FD6FF9E3FCACEEEF" box="[683,788,1644,1670]" captionStart="FIGURES 129 - 136" captionStartId="83.[264,382,1559,1582]" captionTargetBox="[276,1313,509,1512]" captionTargetId="figure@83.[276,1313,509,1518]" captionTargetPageId="83" captionText="FIGURES 129 - 136. Rhodobaenus species. 129 - 130. Rhodobaenus howelli Anderson. 129. Dorsal habitus. 130. Lateral habitus. 131 - 132. Rhodobaenus labrecheae Anderson. 131. Dorsal habitus. 132. Lateral habitus. 133 - 134. Rhodobaenus patriciae Anderson. 133. Dorsal habitus. 134. Lateral habitus. 135 - 136. Rhodobaenus tenorio Anderson. 135. Dorsal habitus. 136. Lateral habitus." httpUri="https://zenodo.org/record/168668/files/figure.png" pageId="88" pageNumber="89">Fig. 133</figureCitation>
) and in that the antennal scape is widened throughout most of its length (
<figureCitation id="11A497B014255B07FDB7F91BFD64EEC7" box="[627,732,1684,1710]" captionStart="FIGURES 147 ­ 150" captionStartId="89.[264,383,1106,1130]" captionTargetBox="[282,1293,586,1085]" captionTargetId="figure@89.[281,1293,585,1085]" captionTargetPageId="89" captionText="FIGURES 147 ­ 150. Rhodobaenus patriciae Anderson. 147. Lateral view of head, female. 148. Antenna. 149. Hind leg, female. 150. Pygidium, female." httpUri="https://zenodo.org/record/156064/files/figure.png" pageId="88" pageNumber="89">Fig. 148</figureCitation>
); however, in 
<taxonomicName id="4E9FF0B614255B07FC42F91BFBA9EEC4" box="[902,1041,1684,1709]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="BBEB572714255B07FC42F91BFBA9EEC4" box="[902,1041,1684,1709]" italics="true" pageId="88" pageNumber="89">R. patriciae</emphasis>
</taxonomicName>
, the pronotal color pattern is different, the punctures of the femora are large, deep and dense, and the vestiture of the third tarsal articles has less extensive pilosity. The species may also be confused with 
<taxonomicName id="4E9FF0B614255B07FECCF883FE2CEF4C" box="[264,404,1804,1829]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="bicinctus">
<emphasis id="BBEB572714255B07FECCF883FE2CEF4C" box="[264,404,1804,1829]" italics="true" pageId="88" pageNumber="89">R. bicinctus</emphasis>
</taxonomicName>
but that species has a different elytral pattern, the rostrum is usually red, and the antennal scape is not expanded throughout its length, but only clavate at the apex.
</paragraph>
<paragraph id="89208B3514245B06FEFCFE93FC4FE997" blockId="89.[264,1324,284,510]" pageId="89" pageNumber="90">
Description.— Male, not known. Female, length, 
<quantity id="4E6726D014245B06FC6CFE93FB80E95F" box="[936,1080,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="6.8" metricValueMin="6.7" pageId="89" pageNumber="90" unit="mm" value="6.75" valueMax="6.8" valueMin="6.7">6.7­6.8 mm</quantity>
; width 
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. Color black and red. Head and rostrum black. Pronotum red with median broad black line as well as lateral broad black line along midheight of flanks, apical margin also infuscate. Elytra red with apical one­third black, this black marking extended forward along sutural region to base of elytra in an inverted “V” shape; humeral region at interval 7 also black. Ventrally with prosternum red, otherwise black or deep piceous.
</paragraph>
<caption id="DDE0DBBD14245B06FECCFBDDFCD4ECF8" httpUri="https://zenodo.org/record/156064/files/figure.png" pageId="89" pageNumber="90" targetBox="[282,1293,586,1085]" targetPageId="89">
<paragraph id="89208B3514245B06FECCFBDDFCD4ECF8" blockId="89.[264,1323,1106,1169]" pageId="89" pageNumber="90">
<emphasis id="BBEB572714245B06FECCFBDDFE5FEC03" bold="true" box="[264,487,1106,1130]" pageId="89" pageNumber="90">FIGURES 147­150.</emphasis>
<taxonomicName id="4E9FF0B614245B06FE35FBDCFCDDEC00" authority="Anderson." authorityName="Anderson." box="[497,869,1106,1130]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="patriciae">
<emphasis id="BBEB572714245B06FE35FBDCFD54EC03" box="[497,748,1107,1130]" italics="true" pageId="89" pageNumber="90">Rhodobaenus patriciae</emphasis>
Anderson.
</taxonomicName>
147. Lateral view of head, female. 148. Antenna. 149. Hind leg, female. 150. Pygidium, female.
</paragraph>
</caption>
<paragraph id="89208B3514245B05FEFCFB47FC1DEA27" blockId="89.[264,1324,1224,1890]" lastBlockId="90.[264,1325,284,990]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">Rostrum slightly shorter than pronotum, strongly humped at base; elongate, narrow, cylindrical, almost straight, virtually impunctate except for moderately dense punctures dorsally on basal; base of rostrum abruptly expanded, basal expanded area very short, about one­sixth total rostral length; base of rostrum immediately in front of eyes subdentate ventrally. Peduncle flat basally, markedly and sharply keeled towards apex. Scrobe with posterior margin about width of base of scape from anterior margin of eyes. Antenna with scape long, about two­thirds length rostrum; scape expanded throughout length, subequal in width from basal one­third to apex, flattened, inner face with shallow irregular sulcus; club slightly laterally compressed, oval; apical pilose part subequal in length to basal glabrous part. Pronotum with lateral margins subparallel to slightly convergent in basal one­half, gradually convergent to apex; very sparsely and shallowly, indistinctly punctate except across base; moderately deeply transversely creased subbasally. Elytra with length one and four­fifths times length pronotum; with striae and intervals impunctate, flat. Scutellum elongate­narrow, lateral margins subparallel in basal one­half; length twice width at base, flat. Pygidium with subapex at middle slightly tumescent, punctate throughout; apical margin with row of elongate, fine setae; apex broadly acuminate. Ventrally with front coxae separated by width of base of scape; prosternum flat between and anterior to coxae. Meso­ and metasterna and ventrites sparsely, shallowly punctate laterally, punctures finer and sparsely medially; last ventrite flat, slightly deflexed at apex. Legs elongate, matte, coarsely, deeply punctate throughout; femora clavate, widest at extreme apex; long, hind femur almost reaching apex of ventrite 5; inner margins of all tibiae with only very short, sparse setae; all tibiae slightly inwardly arcuate. Tarsi each with third article moderately widely dilated, pilose ventrally except in narrow band along basolateral margins; tarsi with third articles symmetrical; apical margin of third articles truncate; claw­bearing segment bilamellate ventrally at apex.</paragraph>
<paragraph id="89208B3514275B05FEFCFDD3FABBEB7F" blockId="90.[264,1325,284,990]" pageId="90" pageNumber="91">
Material Examined.—Female 
<typeStatus id="5624359714275B05FD5FFDD3FC8EEA1F" box="[667,822,604,630]" pageId="90" pageNumber="91" type="holotype">HOLOTYPE</typeStatus>
labelled “Sirena, Corcovado N.P. / Puntarenas Province / 
<collectingCountry id="F188CBA514275B05FE0DFD0BFDF6EAF7" box="[457,590,644,670]" name="Costa Rica" pageId="90" pageNumber="91">Costa Rica</collectingCountry>
, 
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/ G. Fonseca. 
<date id="FD21ADF514275B05FCB6FD0BFC51EAF7" box="[882,1001,644,670]" pageId="90" pageNumber="91" value="1989-11">Nov 1989</date>
/ 
<date id="FD21ADF514275B05FBC0FD0BFBE7EAF7" box="[1028,1119,644,670]" pageId="90" pageNumber="91" value="00-05-27">270500</date>
, 508300”, INBio barcode 113978 (INBC). 
<typeStatus id="5624359714275B05FDFAFD23FD5EEAAF" box="[574,742,684,710]" pageId="90" pageNumber="91" type="paratype">PARATYPES</typeStatus>
(2). 
<collectingCountry id="F188CBA514275B05FC8FFD23FC45EAAF" box="[843,1021,684,710]" name="Costa Rica" pageId="90" pageNumber="91">COSTA RICA</collectingCountry>
. PUNTARENAS PROV­ INCE. Osa Penninsula. Rincon (
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. SW.). 
<geoCoordinate id="ECABEDF214275B05FCEEFD5BFC2AEA87" box="[810,914,724,750]" direction="north" orientation="latitude" pageId="90" pageNumber="91" precision="925" value="8.7">08°42’N</geoCoordinate>
, 
<geoCoordinate id="ECABEDF214275B05FC5BFD5BFBB6EA87" box="[927,1038,724,750]" direction="west" orientation="longitude" pageId="90" pageNumber="91" precision="925" value="-83.48333">83°29’W</geoCoordinate>
”, 
<date id="FD21ADF514275B05FBE2FD5BFB21EA87" box="[1062,1177,724,750]" pageId="90" pageNumber="91" value="1968-03-06">6.iii.1968</date>
, H. Hespenheide (1). Cerro Nora, 
<date id="FD21ADF514275B05FDF1FD73FD72EB7F" box="[565,714,764,790]" pageId="90" pageNumber="91" value="1972-02-10">10 Feb 1972</date>
, S. Medica (1). 
<typeStatus id="5624359714275B05FC63FD73FBA1EB7F" box="[935,1049,764,790]" pageId="90" pageNumber="91" type="paratype">Paratypes</typeStatus>
in CMNC, CWOB.
</paragraph>
<paragraph id="89208B3514275B05FEFCFCABFCB5EB57" blockId="90.[264,1325,284,990]" box="[312,781,804,830]" pageId="90" pageNumber="91">
Distribution.— 
<collectingCountry id="F188CBA514275B05FE2EFCABFDD2EB57" box="[490,618,804,830]" name="Costa Rica" pageId="90" pageNumber="91">Costa Rica</collectingCountry>
(Puntarenas).
</paragraph>
<paragraph id="89208B3514275B05FEFCFCC3FB49EB0F" blockId="90.[264,1325,284,990]" box="[312,1265,844,870]" pageId="90" pageNumber="91">Natural history.—Specimens were collected near sea level in the Osa Penninsula.</paragraph>
<paragraph id="89208B3514275B05FEFCFCFBFB25EBE7" blockId="90.[264,1325,284,990]" box="[312,1181,884,910]" pageId="90" pageNumber="91">Derivation of specific name.—This species is named after Patricia Vaurie.</paragraph>
<paragraph id="89208B3514275B05FEFCFC13FB47EBB7" blockId="90.[264,1325,284,990]" pageId="90" pageNumber="91">
Comments.—This species was first recognized as new by Patricia Vaurie. She placed a determination label on one of the 
<typeStatus id="5624359714275B05FD50FC4BFCBCEBB7" box="[660,772,964,990]" pageId="90" pageNumber="91" type="paratype">paratypes</typeStatus>
stating that it was near but not 
<taxonomicName id="4E9FF0B614275B05FBB0FC4BFB4FEBB4" box="[1140,1271,964,989]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="tornowii">
<emphasis id="BBEB572714275B05FBB0FC4BFB4FEBB4" box="[1140,1271,964,989]" italics="true" pageId="90" pageNumber="91">R. tornowii</emphasis>
</taxonomicName>
.
</paragraph>
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<paragraph id="89208B3514275B05FECCFA7DFCD4EE45" blockId="90.[264,1323,1522,1580]" pageId="90" pageNumber="91">
<emphasis id="BBEB572714275B05FECCFA7DFE51EE63" bold="true" box="[264,489,1522,1546]" pageId="90" pageNumber="91">FIGURES 151­154.</emphasis>
<taxonomicName id="4E9FF0B614275B05FE30FA7CFCE0EE60" authority="Anderson." authorityName="Anderson." box="[500,856,1522,1546]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="tenorio">
<emphasis id="BBEB572714275B05FE30FA7CFD66EE63" box="[500,734,1523,1546]" italics="true" pageId="90" pageNumber="91">Rhodobaenus tenorio</emphasis>
Anderson.
</taxonomicName>
151. Lateral view of head, female. 152. Antenna. 153. Hind leg, female. 154. Pygidium, female.
</paragraph>
</caption>
<paragraph id="89208B3514275B05FECCF908FD97EEA5" blockId="90.[264,836,1671,1740]" pageId="90" pageNumber="91">
<taxonomicName id="4E9FF0B614275B05FECCF908FD1FEECA" authority="Anderson" authorityName="Anderson" box="[264,679,1671,1699]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="tenorio" status="sp. nov.">
<emphasis id="BBEB572714275B05FECCF908FD9BEECA" bold="true" box="[264,547,1671,1699]" italics="true" pageId="90" pageNumber="91">Rhodobaenus tenorio</emphasis>
<emphasis id="BBEB572714275B05FDEFF906FD1FEECA" bold="true" box="[555,679,1673,1699]" pageId="90" pageNumber="91">Anderson</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A0D8EA5C14275B05FD70F906FCFCEECA" box="[692,836,1673,1699]" pageId="90" pageNumber="91" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="11A497B014275B05FED5F93DFE0FEEA5" box="[273,439,1714,1740]" captionStart="FIGURES 129 - 136" captionStartId="83.[264,382,1559,1582]" captionTargetBox="[276,1313,509,1512]" captionTargetId="figure@83.[276,1313,509,1518]" captionTargetPageId="83" captionText="FIGURES 129 - 136. Rhodobaenus species. 129 - 130. Rhodobaenus howelli Anderson. 129. Dorsal habitus. 130. Lateral habitus. 131 - 132. Rhodobaenus labrecheae Anderson. 131. Dorsal habitus. 132. Lateral habitus. 133 - 134. Rhodobaenus patriciae Anderson. 133. Dorsal habitus. 134. Lateral habitus. 135 - 136. Rhodobaenus tenorio Anderson. 135. Dorsal habitus. 136. Lateral habitus." httpUri="https://zenodo.org/record/168668/files/figure.png" pageId="90" pageNumber="91">Figs. 135­136</figureCitation>
, 
<figureCitation id="11A497B014275B05FE07F93DFD9EEEA5" box="[451,550,1714,1740]" captionStart="FIGURES 151 ­ 154" captionStartId="90.[264,383,1522,1546]" captionTargetBox="[278,1308,1089,1495]" captionTargetId="figure@90.[276,1311,1089,1495]" captionTargetPageId="90" captionText="FIGURES 151 ­ 154. Rhodobaenus tenorio Anderson. 151. Lateral view of head, female. 152. Antenna. 153. Hind leg, female. 154. Pygidium, female." httpUri="https://zenodo.org/record/156065/files/figure.png" pageId="90" pageNumber="91">151­154</figureCitation>
)
</paragraph>
<paragraph id="89208B3514275B04FECCF88DFB1CEA87" blockId="90.[264,1323,1794,1900]" lastBlockId="91.[264,1324,284,1870]" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
Identification.—The paired setose patches on the dorsal surface of the female rostrum divided medially by a distinct fine carina (
<figureCitation id="11A497B014275B05FCD7F8A5FCC4EF2D" box="[787,892,1834,1860]" captionStart="FIGURES 151 ­ 154" captionStartId="90.[264,383,1522,1546]" captionTargetBox="[278,1308,1089,1495]" captionTargetId="figure@90.[276,1311,1089,1495]" captionTargetPageId="90" captionText="FIGURES 151 ­ 154. Rhodobaenus tenorio Anderson. 151. Lateral view of head, female. 152. Antenna. 153. Hind leg, female. 154. Pygidium, female." httpUri="https://zenodo.org/record/156065/files/figure.png" pageId="90" pageNumber="91">Fig. 151</figureCitation>
), and the unique sculpturing of the pronotum and elytra (
<figureCitation id="11A497B014275B05FDCCF8DDFD16EF05" box="[520,686,1874,1900]" captionStart="FIGURES 129 - 136" captionStartId="83.[264,382,1559,1582]" captionTargetBox="[276,1313,509,1512]" captionTargetId="figure@83.[276,1313,509,1518]" captionTargetPageId="83" captionText="FIGURES 129 - 136. Rhodobaenus species. 129 - 130. Rhodobaenus howelli Anderson. 129. Dorsal habitus. 130. Lateral habitus. 131 - 132. Rhodobaenus labrecheae Anderson. 131. Dorsal habitus. 132. Lateral habitus. 133 - 134. Rhodobaenus patriciae Anderson. 133. Dorsal habitus. 134. Lateral habitus. 135 - 136. Rhodobaenus tenorio Anderson. 135. Dorsal habitus. 136. Lateral habitus." httpUri="https://zenodo.org/record/168668/files/figure.png" pageId="90" pageNumber="91">Figs. 135­136</figureCitation>
) immediately characterize this odd, perhaps flightless species. Whereas 
<taxonomicName id="4E9FF0B614265B04FE24FE93FDEFE95C" box="[480,599,284,309]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="howelli">
<emphasis id="BBEB572714265B04FE24FE93FDEFE95C" box="[480,599,284,309]" italics="true" pageId="91" pageNumber="92">R. howelli</emphasis>
</taxonomicName>
also has the dorsal surface of the female rostrum setose, in 
<taxonomicName id="4E9FF0B614265B04FAD5FE93FEE4E934" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="tenorio">
<emphasis id="BBEB572714265B04FAD5FE93FEE4E934" italics="true" pageId="91" pageNumber="92">R. tenorio</emphasis>
</taxonomicName>
, the setosity is shorter, with each hair broader and scale­like, and the extent of the pilosity is limited to a smaller patch extended from immediately above the point of antennal insertion to the midlength of the rostrum (
<figureCitation id="11A497B014265B04FCDEFE1BFC38E9C7" box="[794,896,404,430]" captionStart="FIGURES 151 ­ 154" captionStartId="90.[264,383,1522,1546]" captionTargetBox="[278,1308,1089,1495]" captionTargetId="figure@90.[276,1311,1089,1495]" captionTargetPageId="90" captionText="FIGURES 151 ­ 154. Rhodobaenus tenorio Anderson. 151. Lateral view of head, female. 152. Antenna. 153. Hind leg, female. 154. Pygidium, female." httpUri="https://zenodo.org/record/156065/files/figure.png" pageId="91" pageNumber="92">Fig. 151</figureCitation>
). The patch is also divided medially by a distinct, fine, sharp carina. The pronotum and elytra are scupltured unlike any other 
<taxonomicName id="4E9FF0B614265B04FECCFE6BFE1FE994" box="[264,423,484,509]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB572714265B04FECCFE6BFE1FE994" box="[264,423,484,509]" italics="true" pageId="91" pageNumber="92">Rhodobaenus</emphasis>
</taxonomicName>
. The pronotum has a subbasal transverse impression (as do many 
<taxonomicName id="4E9FF0B614265B04FECCFD83FE1FEA4C" box="[264,423,524,549]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB572714265B04FECCFD83FE1FEA4C" box="[264,423,524,549]" italics="true" pageId="91" pageNumber="92">Rhodobaenus</emphasis>
</taxonomicName>
) but this impression is extended anteriorly for a short distance on either side of the broadly elevated midline. The elytra has the scutellar region broadly impressed and also has a distinct oblique impression extended from the humerus to interval 2 at about the apical one­third. The pygidium is markedly, acutely tumescent apically (
<figureCitation id="11A497B014265B04FBA6FD0BFB7EEAF7" box="[1122,1222,644,670]" captionStart="FIGURES 151 ­ 154" captionStartId="90.[264,383,1522,1546]" captionTargetBox="[278,1308,1089,1495]" captionTargetId="figure@90.[276,1311,1089,1495]" captionTargetPageId="90" captionText="FIGURES 151 ­ 154. Rhodobaenus tenorio Anderson. 151. Lateral view of head, female. 152. Antenna. 153. Hind leg, female. 154. Pygidium, female." httpUri="https://zenodo.org/record/156065/files/figure.png" pageId="91" pageNumber="92">Fig. 154</figureCitation>
) and the tarsi have the venter of the claw bearing articles flat (as in 
<taxonomicName id="4E9FF0B614265B04FC14FD23FBF1EAAC" box="[976,1097,684,709]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="howelli">
<emphasis id="BBEB572714265B04FC14FD23FBF1EAAC" box="[976,1097,684,709]" italics="true" pageId="91" pageNumber="92">R. howelli</emphasis>
</taxonomicName>
). The short elytra with somewhat rounded humeri (
<figureCitation id="11A497B014265B04FD48FD5BFD48EA87" box="[652,752,724,750]" captionStart="FIGURES 129 - 136" captionStartId="83.[264,382,1559,1582]" captionTargetBox="[276,1313,509,1512]" captionTargetId="figure@83.[276,1313,509,1518]" captionTargetPageId="83" captionText="FIGURES 129 - 136. Rhodobaenus species. 129 - 130. Rhodobaenus howelli Anderson. 129. Dorsal habitus. 130. Lateral habitus. 131 - 132. Rhodobaenus labrecheae Anderson. 131. Dorsal habitus. 132. Lateral habitus. 133 - 134. Rhodobaenus patriciae Anderson. 133. Dorsal habitus. 134. Lateral habitus. 135 - 136. Rhodobaenus tenorio Anderson. 135. Dorsal habitus. 136. Lateral habitus." httpUri="https://zenodo.org/record/168668/files/figure.png" pageId="91" pageNumber="92">Fig. 135</figureCitation>
) indicates species is likely flightless.
</paragraph>
<paragraph id="89208B3514265B04FEFCFD73FC6AEB57" blockId="91.[264,1324,284,1870]" pageId="91" pageNumber="92">
Description.—Female, length, 
<quantity id="4E6726D014265B04FD60FD73FCB9EB7F" box="[676,769,764,790]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="91" pageNumber="92" unit="mm" value="9.5">9.5 mm</quantity>
; width 
<quantity id="4E6726D014265B04FC9CFD73FC0FEB7F" box="[856,951,764,790]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="91" pageNumber="92" unit="mm" value="3.2">3.2 mm</quantity>
. Male not known. Color black with dark red­brown head, rostrum, mesosternum and coxae.
</paragraph>
<paragraph id="89208B3514265B03FEFCFCC3FEF1E9C7" blockId="91.[264,1324,284,1870]" lastBlockId="92.[264,1324,284,790]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">Rostrum shorter than pronotum, slightly humped at base; elongate, tapered apically, laterally compressed basally in front of point of antennal insertion, slightly evenly arcuate; with deep, large punctures laterally in front of point of antennal insertion, dorsally with short, dense, broad, dark golden scale­like hairs arranged in paramedian patches extedned from above point of antennal insertion to about midlength of rostrum; base of rostrum slightly expanded, basal expanded area short, about one­fourth total rostral length. Rostrum glabrous ventrally; peduncle flat, not produced or toothed. Scrobe with posterior margin about twice width of base of scape from anterior margin of eyes. Antenna with scape slightly longer than one­half length rostrum; scape slightly expanded apically, not flattened; club cylindrical, elongate­oval; apical pilose part slightly less than one­half length of basal glabrous part. Pronotum with lateral margins evenly slightly arcuate from base to subapical constriction, apex distinctly constricted, tubulate; sparsely and shallowly, indistinctly punctate throughout; with subbasal transverse impression and longitudical paramedian impressions extended anteriorly of subbasal impression for short distance on either side of broadly elevated midline. Elytra with length subequal to length pronotum; with striae with indistinct, shallow, elongate punctures, intervals impunctate, flat; elytra with scutellar region broadly impressed, also each elytron with distinct oblique impression extended from the humerus to interval 2 at about the apical one­third. Scutellum elongate, lateral margins subparallel in basal one­half; length twice width at base, flat. Pygidium with subapex at middle markedly, acutely tumescent, regularly punctate with dense long, scale­like setae on tumosity; apex narrowly rounded. Ventrally with front coxae separated by less than width of base of scape; prosternum flat between and anterior to coxae. Meso­ and metasterna and ventrite 1­5 sparsely, shallowly punctate throughout; last ventrite flat at middle at apex. Legs (especially hind) elongate, matte, coarsely, shallowly punctate throughout; femora clavate, long, hind femur reaching apex of pygidium; inner margins of all tibiae with only very short, peg­like setae; all tibiae straight. Tarsi each with third article slightly dilated, pilose ventrally except in narrow median line; tarsi with third articles assymmetrical; apical margin of third articles very slightly emarginate; claw­bearing segment flat ventrally at apex; tarsi moderately long, first article less than twice as long as wide.</paragraph>
<paragraph id="89208B3514215B03FEFCFE33FC43EA4F" blockId="92.[264,1324,284,790]" pageId="92" pageNumber="93">
Material examined.—Female 
<typeStatus id="5624359714215B03FD60FE33FC87E9BF" box="[676,831,444,470]" pageId="92" pageNumber="93" type="holotype">HOLOTYPE</typeStatus>
labelled &quot; 
<collectingCountry id="F188CBA514215B03FC04FE33FBC0E9BF" box="[960,1144,444,470]" name="Costa Rica" pageId="92" pageNumber="93">COSTA RICA</collectingCountry>
, Prov. Limón. PILA / Bratsi, Refugio Valle del Silencio / 
<quantity id="4E6726D014215B03FCC1FE6BFCE2E997" box="[773,858,484,510]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="92" pageNumber="93" unit="m" value="2400.0">2400m</quantity>
. 
<date id="FD21ADF514215B03FCA1FE6BFBB5E997" box="[869,1037,484,510]" pageId="92" pageNumber="93" value="2001-04-17">17 ABR 2001</date>
. R. González. / Manual. L_S_341400_577250 #62561&quot; INBio barcode 3307215 (INBC).
</paragraph>
<paragraph id="89208B3514215B03FEFCFDBBFD6AEA27" blockId="92.[264,1324,284,790]" box="[312,722,564,590]" pageId="92" pageNumber="93">
Distribution.— 
<collectingCountry id="F188CBA514215B03FE2DFDBBFDD1EA27" box="[489,617,564,590]" name="Costa Rica" pageId="92" pageNumber="93">Costa Rica</collectingCountry>
(Limón)
</paragraph>
<paragraph id="89208B3514215B03FEFCFDD3FB91EA1F" blockId="92.[264,1324,284,790]" box="[312,1065,604,630]" pageId="92" pageNumber="93">Natural history.—No information is available on natural history.</paragraph>
<paragraph id="89208B3514215B03FEFCFD0BFA91EAAF" blockId="92.[264,1324,284,790]" pageId="92" pageNumber="93">
Derivation of specific name.—This species is named after INBio parataxonomist Roger Gonzalez 
<taxonomicName id="4E9FF0B614215B03FE09FD23FD90EAAF" box="[461,552,684,710]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="92" phylum="Arthropoda" rank="species" species="tenorio">Tenorio</taxonomicName>
for his diligence (and success) in finding new species of weevils.
</paragraph>
<paragraph id="89208B3514215B03FEFCFD5BFE5EEB7F" blockId="92.[264,1324,284,790]" pageId="92" pageNumber="93">
Comments.—This is the only known species of 
<taxonomicName id="4E9FF0B614215B03FCB7FD5BFBABEA84" box="[883,1043,724,749]" class="Insecta" family="Dryophthoridae" genus="Rhodobaenus" kingdom="Animalia" order="Coleoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="BBEB572714215B03FCB7FD5BFBABEA84" box="[883,1043,724,749]" italics="true" pageId="92" pageNumber="93">Rhodobaenus</emphasis>
</taxonomicName>
with such pronotal and elytral scuplturing.
</paragraph>
</subSubSection>
</treatment>
</document>