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<document ID-CLB-Dataset="28237" ID-DOI="http://dx.doi.org/10.3897/jhr.53.9890" ID-GBIF-Dataset="e6b7983e-dc70-447a-bb89-c93b74d6f249" ID-Pensoft-Pub="1314-2607-53-77" ID-Pensoft-UUID="CB6EB5480F1F2F28FFF91521801EFF8C" ID-ZBK="989B7FE41F5048A784148D0AF4F4C064" ID-Zenodo-Dep="575132" ID-ZooBank="989B7FE41F5048A784148D0AF4F4C064" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2607-53-77" ModsDocOrigin="Journal of Hymenoptera Research " ModsDocTitle="New Dryocosmus Giraud species associated with Cyclobalanopsis and non-Quercus host plants from the Eastern Palaearctic (Hymenoptera, Cynipidae, Cynipini)" checkinTime="1553125490764" checkinUser="pensoft" docAuthor="Tang, Chang-Ti, Miko, Istvan, Nicholls, James A., Schweger, Szabina, Yang, Man-Miao, Stone, Graham N., Sinclair, Frazer, Bozso, Miklos, Melika, George &amp; Penzes, Zsolt" docDate="2016" docId="F0579F8A85D1D2A9BEA14E0FF4CF9307" docLanguage="en" docName="JourHymenoptRes 53: 77-162" docOrigin="Journal of Hymenoptera Research 53" docPubDate="2016-12-19" docSource="http://dx.doi.org/10.3897/jhr.53.9890" docTitle="Dryocosmus harrisonae Melika &amp; Tang 2016, sp. n." docType="treatment" docUuid="796EC89B-9267-464E-B2B3-C888AEC76CFC" docUuidSource="ZooBank" docVersion="6" id="CB6EB5480F1F2F28FFF91521801EFF8C" lastPageNumber="107" masterDocId="CB6EB5480F1F2F28FFF91521801EFF8C" masterDocTitle="New Dryocosmus Giraud species associated with Cyclobalanopsis and non- Quercus host plants from the Eastern Palaearctic (Hymenoptera, Cynipidae, Cynipini)" masterLastPageNumber="162" masterPageNumber="77" pageNumber="100" updateTime="1732741174302" updateUser="ExternalLinkService">
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<mods:title id="889FD6A635BC21DA4DF79B8E7CC1CD70">New Dryocosmus Giraud species associated with Cyclobalanopsis and non- Quercus host plants from the Eastern Palaearctic (Hymenoptera, Cynipidae, Cynipini)</mods:title>
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<mods:namePart id="9B0E1B869C14271015D42AAFA8777BFB">Tang, Chang-Ti</mods:namePart>
<mods:affiliation id="A5747F3CF15B10A3652D009446C00967">Department of Entomology, National Chung Hsing University, 250 Kuo Kuang Rd., Taichung 402, Taiwan</mods:affiliation>
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<mods:namePart id="1E07C5655E41C1C1F708795D3C55A510">Miko, Istvan</mods:namePart>
<mods:affiliation id="02E0C220FF0A3DE40A0AF855AFF9C19A">Pennsylvania State University, Department of Entomology, Frost Entomological Museum, PA, USA</mods:affiliation>
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<mods:namePart id="D921061A7898D0D91ABC3571D2984CC8">Nicholls, James A.</mods:namePart>
<mods:affiliation id="C0027800ABF95AC964CD07F2622523DD">Institute of Evolutionary Biology, University of Edinburgh, King's Buildings, West Mains Road, Edinburgh EH 9 3 JT, Scotland, U. K.</mods:affiliation>
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<mods:namePart id="4CDCD91734BED786DA8A04D51850FC26">Schweger, Szabina</mods:namePart>
<mods:affiliation id="5FF0AFA9E4F4C1BFF1B75B86574B753A">University of Szeged, Department of Ecology, Koezep fasor 52 Szeged, Hungary</mods:affiliation>
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<mods:namePart id="5F3688C0571B0C0D110522470DD66743">Yang, Man-Miao</mods:namePart>
<mods:affiliation id="35760B847F2176B28AC08D232EE88EAB">Department of Entomology, National Chung Hsing University, 250 Kuo Kuang Rd., Taichung 402, Taiwan</mods:affiliation>
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<mods:namePart id="50AF1C2B3FDBD824778B08ECAD2D2216">Stone, Graham N.</mods:namePart>
<mods:affiliation id="5B4D99429551CA90084FDF2AEA769F45">Institute of Evolutionary Biology, University of Edinburgh, King's Buildings, West Mains Road, Edinburgh EH 9 3 JT, Scotland, U. K.</mods:affiliation>
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<mods:namePart id="BA4F74FC844B28BE58143F97D0096141">Sinclair, Frazer</mods:namePart>
<mods:affiliation id="440D6E237F524559AEF01A8452517A39">Institute of Evolutionary Biology, University of Edinburgh, King's Buildings, West Mains Road, Edinburgh EH 9 3 JT, Scotland, U. K.</mods:affiliation>
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<mods:namePart id="0CE1D0CD3AFE419106116D629B742E74">Bozso, Miklos</mods:namePart>
<mods:affiliation id="5969E9BACED0035290230CB64E22946D">Plant Health and Molecular Biology Laboratory, National Food Chain Safety Office, Directorate of Plant Protection, Soil Conservation and Agri-environment, Budaoersi str. 141 - 145, Budapest 1118, Hungary</mods:affiliation>
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<mods:namePart id="960F98E43B48F1E2A1694C82E151F4FF">Melika, George</mods:namePart>
<mods:affiliation id="F67660104B9AC333DCBA34DFAE60BA97">Plant Health and Molecular Biology Laboratory, National Food Chain Safety Office, Directorate of Plant Protection, Soil Conservation and Agri-environment, Budaoersi str. 141 - 145, Budapest 1118, Hungary</mods:affiliation>
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<mods:namePart id="0405EA4B3EEAB9759F0743753A30D092">Penzes, Zsolt</mods:namePart>
<mods:affiliation id="4514137D1F767C59E7668A103E967608">University of Szeged, Department of Ecology, Koezep fasor 52 Szeged, Hungary</mods:affiliation>
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<mods:title id="BCC607732B0EA5A772AA67ED14369557">Journal of Hymenoptera Research</mods:title>
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<mods:date id="C8C3AB6003DC57CD238BC29C909C93EF">2016</mods:date>
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<mods:number id="7D055EB1A9D57C51D3739F28E984FDAE">53</mods:number>
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<mods:start id="4E7F232FD9BB115E07CC3C555EAC6965">77</mods:start>
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<mods:classification id="7EDA4D91DCF2335AB846A03C8E4A2412">journal article</mods:classification>
<mods:identifier id="DCE39F8028EDA7D53F724E532C3BF354" type="DOI">http://dx.doi.org/10.3897/jhr.53.9890</mods:identifier>
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<treatment id="F0579F8A85D1D2A9BEA14E0FF4CF9307" ID-GBIF-Taxon="154473345" LSID="urn:lsid:zoobank.org:act:796EC89B-9267-464E-B2B3-C888AEC76CFC" httpUri="http://treatment.plazi.org/id/F0579F8A85D1D2A9BEA14E0FF4CF9307" lastPageId="30" lastPageNumber="107" pageId="23" pageNumber="100" scope_family="Cynipidae" scope_order="Hymenoptera">
<subSubSection id="02802159CDF764BB9B3F50DE7B27D8DC" pageId="23" pageNumber="100" type="nomenclature">
<paragraph id="C47B800686A33E9CCD1E175B26EBB2E0" pageId="23" pageNumber="100">
<taxonomicName id="B0B4656C8F502D1A3495D52164AEAD75" LSID="http://zoobank.org/796EC89B-9267-464E-B2B3-C888AEC76CFC" authority="Melika &amp; Tang" authorityName="Melika &amp; Tang" authorityYear="2016" class="Insecta" family="Cynipidae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus harrisonae" order="Hymenoptera" pageId="23" pageNumber="100" phylum="Arthropoda" rank="species" species="harrisonae" status="sp. n.">Dryocosmus harrisonae Melika &amp; Tang</taxonomicName>
<taxonomicNameLabel id="AC9C694AF05D10C32EE1612DCA9E3179" pageId="23" pageNumber="100">sp. n.</taxonomicNameLabel>
<figureCitation id="6EBEC6E77E00C216FAFAA4AD57F1126A" captionStart="Figures 4855" captionStartId="F9" captionText="Figures 48 - 55. Dryocosmus harrisonae sp. n. 48 - 50 head, female: 48 frontal view 49 dorsal view 50 posterior view 51 - 52 head, male: 51 frontal view 52 dorsal view 53 - 54 antenna: 53 female 54 male 55 mesosoma, female, lateral view (gen = gena, spe = speculum)." figureDoi="10.3897/jhr.53.9890.figures48-55" httpUri="https://binary.pensoft.net/fig/115548" pageId="23" pageNumber="100">Figures 48-55</figureCitation>
<figureCitation id="59E40C57B3F2FC942EBDE15A91317E96" captionStart="Figures 5660" captionStartId="F10" captionText="Figures 56 - 60. Dryocosmus harrisonae, female sp. n. 56 mesoscutum, dorsal view 57 mesoscutellum, dorsal view 58 pronotum and propleuron, frontal view 59 metascutellum and propodeum, posterodorsal view 60 fore wing, part." figureDoi="10.3897/jhr.53.9890.figures56-60" httpUri="https://binary.pensoft.net/fig/115549" pageId="23" pageNumber="100">, 56-60</figureCitation>
<figureCitation id="4DD3DD4F024D0B95149C2E0542639DF0" captionStart="Figures 6163" captionStartId="F11" captionText="Figures 61 - 63. Dryocosmus harrisonae sp. n. 61 female, metasoma, lateral view 62 - 63 galls (photos by C. - T. Tang)." figureDoi="10.3897/jhr.53.9890.figures61-63" httpUri="https://binary.pensoft.net/fig/115550" pageId="23" pageNumber="100">, 61-63</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A83D05A4D46906F57FB7DD75AAEC5D6E" pageId="24" pageNumber="101" type="type material">
<paragraph id="12F16C71D89B9FB72E884DAE8383FC71" pageId="24" pageNumber="101">
<pageBreakToken id="3BCBD304FC4E81DBC9BEB4561766B754" pageId="24" pageNumber="101" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph id="387B7C332F0652BD5744AACE0A1C676C" pageId="24" pageNumber="101">
HOLOTYPE female: CHINA: Yunnan Prov., Xishuangbanna, tea plantation near Meng Hai, ex
<taxonomicName id="B6EEFB6EA94AD35A4B73D3C42875BEA2" class="Hexapoda" family="Fagaceae" genus="Castanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castanopsis echinocarpa" order="Hymenoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="echinocarpa">
<emphasis id="69450B6BE0BA9DECFED354FFE3EE7C39" italics="true" pageId="24" pageNumber="101">Castanopsis echinocarpa</emphasis>
</taxonomicName>
, 7.IV.2011 (CHI11),
<geoCoordinate id="3D4CE9E3CBB65C08D8B99E84E9223B69" degrees="21.968800" direction="north" orientation="latitude" precision="1" value="21.9688">21.968800°N</geoCoordinate>
,
<geoCoordinate id="13F8A523A012B61E9848D33FCC2FFE7C" degrees="100.600883" direction="east" orientation="longitude" precision="1" value="100.60088">100.600883°E</geoCoordinate>
, 1342m, ex oval chamber gall on stalk extending from petiole or midrib of very young leaf (AGWP-Morpho57), em. 13.IV.2011, leg. C. T. Tang, F. Sinclair, J. Hearn. Four female and two male PARATYPES: 2 female paratypes: CHINA: Yunnan Prov., Xishuangbanna, tea plantation near Meng Hai, ex
<taxonomicName id="1222BB588B3C0D2581A79695803E1C7D" class="Hexapoda" family="Fagaceae" genus="Castanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castanopsis echinocarpa" order="Hymenoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="echinocarpa">
<emphasis id="33C740EDCFECE7705410BCC1C5CB5101" italics="true" pageId="24" pageNumber="101">Castanopsis echinocarpa</emphasis>
</taxonomicName>
, 7.IV.2011 (CHI11),
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,
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, 1342m, ex oval chamber gall on stalk extending from petiole or midrib of very young leaf (AGWP-Morpho57), em. 17.IV.2011, leg. C. T. Tang, F. Sinclair, J. Hearn; 2 female paratypes: CHINA: Yunnan Prov., Xishuangbanna, tea plantation near Meng Hai, ex
<taxonomicName id="BF7ECF52DF8E8C4B3D1896641E076958" class="Hexapoda" family="Fagaceae" genus="Castanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castanopsis echinocarpa" order="Hymenoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="echinocarpa">
<emphasis id="B996474DD53EE4DB52EB19FA262A4D72" italics="true" pageId="24" pageNumber="101">Castanopsis echinocarpa</emphasis>
</taxonomicName>
, 7.IV.2011 (CHI11),
<geoCoordinate id="3490DC7BDCBDBD2ED41796EDC753116E" degrees="21.968800" direction="north" orientation="latitude" precision="1" value="21.9688">21.968800°N</geoCoordinate>
,
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, 1342m, ex oval chamber gall on stalk extending from petiole or midrib of very young leaf (AGWP-Morpho57), em. 19.IV.2011, leg. C. T. Tang, F. Sinclair, J. Hearn; 1 male paratype: CHINA: Yunnan Prov., Xishuangbanna, tea plantation near Meng Hai, ex
<taxonomicName id="BD13C0F6994BAFE13CDED57E7DB6AB50" class="Hexapoda" family="Fagaceae" genus="Castanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castanopsis echinocarpa" order="Hymenoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="echinocarpa">
<emphasis id="95F524C47071F22B6F228A7AA34A3904" italics="true" pageId="24" pageNumber="101">Castanopsis echinocarpa</emphasis>
</taxonomicName>
, 7.IV.2011 (CHI11),
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,
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, 1342m, ex oval chamber gall on stalk extending from petiole or midrib of very young leaf (AGWP-Morpho57), em. 15.IV.2011, leg. C. T. Tang, F. Sinclair, J. Hearn; 1 male paratype: CHINA: Yunnan Prov., Xishuangbanna, tea plantation near Meng Hai, ex
<taxonomicName id="AC99D6CA587CAF5AB7A20D47F296358B" class="Hexapoda" family="Fagaceae" genus="Castanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castanopsis echinocarpa" order="Hymenoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="echinocarpa">
<emphasis id="98D6E9FE39403D0C66C853939B66CA83" italics="true" pageId="24" pageNumber="101">Castanopsis echinocarpa</emphasis>
</taxonomicName>
, 7.IV.2011 (CHI11),
<geoCoordinate id="8424BD17791989524B43A49E771DEAD0" degrees="21.968800" direction="north" orientation="latitude" precision="1" value="21.9688">21.968800°N</geoCoordinate>
,
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, 1342m, ex oval chamber gall on stalk extending from petiole or midrib of very young leaf (AGWP-Morpho57), em. 18.IV.2011, leg. C. T. Tang, F. Sinclair, J. Hearn.
</paragraph>
<paragraph id="BEB3117FC9692BC76EDD1372AE992176" pageId="24" pageNumber="101">The female holotype and 1 male paratype are deposited in NMNS, 2 females and 1 male paratypes in PHMB, and 2 female paratypes in NCHU.</paragraph>
</subSubSection>
<subSubSection id="CEC6DE0C815B9EE48E4ED3EBB61B03EC" pageId="24" pageNumber="101" type="etymology">
<paragraph id="2F1335D0DC55C9809F7FD00EBBBC74BE" pageId="24" pageNumber="101">Etymology.</paragraph>
<paragraph id="8B1D6335FB24A1D2AC769CEEC92C2035" pageId="24" pageNumber="101">Named in honour of Ms Warin Harrison (Xishuangbanna Tropical Botanical Garden, Yunnan, China), who was our guide during the collecting trip in China in 2011.</paragraph>
</subSubSection>
<subSubSection id="DE2C693243D5011C021B7F4809930806" pageId="25" pageNumber="102" type="diagnosis">
<paragraph id="169C512C738524D9FB54EF5075CD7DF6" pageId="25" pageNumber="102">
<pageBreakToken id="780805DD084C1820117BD3E34736593D" pageId="25" pageNumber="102" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph id="1946E01C30327A392F243A32FE117B93" pageId="25" pageNumber="102">
See the diagnosis for
<taxonomicName id="1B394AA9AA40576882378EE50881FAF4" authorityName="Schweger &amp; Tang" authorityYear="2016" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus cannoni" order="Hymenoptera" pageId="25" pageNumber="102" phylum="Arthropoda" rank="species" species="cannoni">
<emphasis id="88422EBBB793A979285B367775BA161A" italics="true" pageId="25" pageNumber="102">Dryocosmus cannoni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4F3589FF7EF129317641C5EDB11B3808" lastPageId="29" lastPageNumber="106" pageId="25" pageNumber="102" type="description">
<paragraph id="56B854998929F41C44A04C7724C2BE05" pageId="25" pageNumber="102">Description.</paragraph>
<paragraph id="C89BB2723C060E17307EA37DA4FF3411" pageId="25" pageNumber="102">Sexual female: Head color: dark brown, except for lighter clypeus; mandibles, maxillary and labial palps yellowish; scape and pedicel yellow, flagellomeres progressively darker. Mesosoma and metasoma color: dark brown, except for lighter tegula and propodeum, legs yellowish.</paragraph>
<paragraph id="358BA43FCD10EB47184FEB536BDC343C" lastPageId="26" lastPageNumber="103" pageId="25" pageNumber="102">
Head sculpture: coriaceous. Head shape anterior view: rounded. Head width / head length: 1.86. Head width / head height: 1.10. Head width / maximum meso
<pageBreakToken id="F327DDB3E8603C054DC7804E863718EE" pageId="26" pageNumber="103" start="start">soma</pageBreakToken>
width: &lt;1. Gena sculpture: coriaceous. Gena reflectivity: NOT CODED. Gena length / eye width: 0.40. Gena shape: broadened posterior to eye, visible in frontal view. Malar striae dorsal limit: torulus-eye line on lower face and lower eye margin on malar area. Malar striae count: present. Malar area: alutaceous. Malar sulcus: absent. Eye height / malar distance: 3.33. Inner margins of eyes: converging ventrally. Median ocellus shape: NOT CODED. Lateral ocellus shape: NOT CODED. POL / OOL: 1.10. OOL / diameter of lateral ocellus: 2.30. OOL / LOL: 1.70. Diameter of lateral ocellus / diameter of median ocellus: 1. Transfacial distance / eye height: 1.16. Diameter of torulus / intertorular distance: 1.20. Intertorular distance / eye torulus distance: NOT CODED. Eye-torulus distance / diameter of torulus: 1.25. Lower face sculpture: alutaceous. Lower face pilosity color: white. Lower face pilosity density: rare. Clypeus convexity: flat. Clypeus sculpture: smooth. Clypeus ventral margin shape: straight. Clypeus reflectivity: NOT CODED. Clypeus shape anterior view: rectangular. Clypeus pilosity: NOT CODED. Clypeus pilosity density: NOT CODED. Clypeus pilosity color: NOT CODED. Anterior tentorial pit: large, distinct. Epistomal sulcus: indistinct. Clypeo-pleurostomal line: indistinct. Frons sculpture: coriaceous. Frons pilosity density: rare. Frons reflectivity: matt. Frons pilosity color: white. Impression around central ocellus: present. Interocellar area sculpture: coriaceous. Interocellar area pilosity density: rare. Interocellar area reflectivity: matt. Interocellar area pilosity color: white. Vertex sculpture: coriaceous. Vertex pilosity density: rare. Vertex reflectivity: matt. Vertex pilosity color: white. Occiput sculpture: coriaceous. Occiput pilosity density: rare. Occiput reflectivity: matt. Occiput pilosity color: white. Postocciput sculpture: coriaceous. Postocciput reflectivity: matt. Postocciput pilosity: present. Postocciput pilosity density: rare. Postocciput pilosity color: white. Median impression of postocciput dorsal to occipital foramen: present. Posterior tentorial pit: distinct, ovate, deep. Impression abjacent ventrally to posterior tentorial pit: present. Postgena sculpture: alutaceous in outer area, smooth in inner area. Postgena reflectivity: glossy medially. Postgena pilosity color: white. Postgena pilosity density: rare. Postgena pilosity count: absent medially, present laterally. Postgenal bridge / height of occipital foramen: 1. Postgenal bridge / length of oral foramen: &lt;1. Postgenal bridge sculpture: NOT CODED. Number of flagellomeres (female): 12. Antenna length / body length: &lt;1. Pedicel length / pedicel width: 1.70. Pedicel length / length of broadened part of scape: NOT CODED. Combined length of scape and pedicel / first flagellomere length: NOT CODED. F1 length / F2 length: 1.16. F1 length / pedicel length: 1.9. F1 length / F3 length: NOT CODED. Flagellomeres relative length: F3&gt;F4&gt;F5&gt;F6&gt;F7&gt;F8&gt;F9&gt;F10&gt;F11. F4-F7 relative length: NOT CODED. F3 length/F2 length: NOT CODED. F3 length / F4 length: NOT CODED. F8 length / F7 length: NOT CODED. F9 length /F8 length: NOT CODED. F10 length / F9 length: NOT CODED. F9-F11 relative length: NOT CODED. F10 length / F11 length: NOT CODED. F12 length / F11 length: 1.5. F13 length / F12 length: NOT CODED. Placoid sensilla present on: F2-F12.
</paragraph>
<paragraph id="2D7054E0764BCB48F694F381535C4043" lastPageId="28" lastPageNumber="105" pageId="26" pageNumber="103">
Mesosoma length lateral view / mesosoma height lateral view: 1.03. Pronotum sculpture: alutaceous with some rugae laterally. Pronotal dorsal row of setae count:
<pageBreakToken id="7612B59368280C44516DF1F35A2C3126" pageId="27" pageNumber="104" start="start">present</pageBreakToken>
. Transverse pronotal sulcus depth: deep. Transverse pronotal sulcus sculpture: NOT CODED. Mesoscutum sculpture: smooth. Mesoscutum reflectivity: glossy. Adnotaular setae: present. Mesoscutum length / transscutal line: NOT CODED. Notaulus limits: well-impressed, posterior end adjacent to posterior margin of mesoscutum, anterior end adjacent to anterior margin of mesoscutum. Notaulus sculpture: NOT CODED. Notaulus posterior region width / anterior region width: NOT CODED. Mesoscutal suprahumeral sulcus anterior end vs notaulus anterior end: adjacent. Mesoscutal suprahumeral sulcus sculpture: NOT CODED. Median mesoscutal line: absent. Median mesoscutal line shape: NOT CODED. Parapsidal line: absent. Parapsidal line distinctness: NOT CODED. Anteroadmedian line: present. Antero-admedian line length / mesoscutum median length: NOT CODED. Dorsomedian area
<pageBreakToken id="EE779DA179248C3D9C769F4300E37721" pageId="28" pageNumber="105" start="start">of</pageBreakToken>
mesoscutellar-axillar complex (disc of mesoscutellum+axillar foveae): coriaceous, foveolate laterally and posteriorly. Dorsomedian area of mesoscutellar-axillar complex (disc of mesoscutellum+axillar foveae) shape: trapezoid. Dorsomedian area of mesoscutellar-axillar complex (disc of mesoscutellum+axillar foveae) pilosity color: white. Dorsomedian area of mesoscutellar-axillar complex (disc of mesoscutellum+axillar foveae) pilosity density: rare. Mesoscutellar axillar complex posterior margin vs metanotum: overhanging. Mesoscutellar-axillar complex length / mesoscutellar disc width:&gt;1. Disc of mesoscutellum sculpture: rugose. Disc of mesoscutellum maximum width: in posterior 1/3. Scutellar fovea sculpture: smooth without longitudinal rugae. Scutellar fovea shape: transversely ovate. Scutellar fovea reflectivity: glossy. Foveal septum: narrow. Anterior pits on foveal septum: NOT CODED. Scutellar fovea maximum diameter / scutellar fovea minimum diameter: 1.7. Scutellar fovea minimum diameter / foveal septum width: NOT CODED. Postacetabular suclus count: NOT CODED. Mesopleuron sculpture: smooth transepisternal line is marked by few delicate sulci, area dorsal to transepisternal line smooth except few short transverse anterior striae. Mesopleuron reflectivity: matt; glossy. Mesopleuron pilosity: glabrous. Speculum sculpture: wrinkled. Speculum reflectivity: NOT CODED. Mesopleural triangle sculpture: smooth. Mesopleural triangle reflectivity: glossy. Mesopleural triangle pilosity: present. Mesopleural triangle pilosity color: NOT CODED. Mesopleural trinagle pilosity density: NOT CODED. Dorsoaxillar area sculpture: alutaceous with few rugae. Dorsoaxillar area reflectivity: NOT CODED. Dorsoaxillar area pilosity color: NOT CODED. Dorsoaxillar area pilosity density: NOT CODED. Lateroaxillar area sculpture: alutaceous with few rugae. Lateroaxillar area reflectivity: NOT CODED. Lateroaxillar area pilosity density: NOT CODED. Lateroaxillar area pilosity color: NOT CODED. Subaxillular bar sculpture: smooth. Subaxillular bar reflectivity: glossy. Posterior height of subaxillular bar / height of metanotal trough: 1.25. Metapleural sulcus anterior end: reaches mesometapleural suture in upper ⅓ of its length. Metascutellum sculpture: coriaceous. Metanotal trough sculpture: smooth. Metanotal trough reflectivity: glossy. Metanotal trough pilosity: absent. Metanotal trough pilosity density: NOT CODED. Ventral impressed area of metanotum sculpture: smooth without striae. Metascutellum height / ventral impressed area of metanotum height: 1. Central propodeal area sculpture: smooth with irregular rugae. Central propodeal area reflectivity: NOT CODED. Lateral propodeal carina shape: broad, high, lyre-shaped. Lateral propodeal area sculpture: smooth with irregular rugae. Lateral propodeal area pilosity: present. Lateral propodeal area pilosity color: NOT CODED. Lateral propodeal area pilosity density: NOT CODED. Nucha sculpture: with delicate longitudinal rugae dorsally and laterally. Radial cell length / radial cell width: 4.1. Rs+M vs basalis: reaches basalis in lower half of its height. Rs+M color: dark brown. Areolet: large, triangular, conspicuous. Marginal cilia: long. Rs distal end vs wing margin: adjacent to wing margin. R1 distal end vs wing margin: adjacent to wing margin. Basal lobe on metatarsal claw: absent. Basal lobe on tarsal claw shape: NOT CODED.
</paragraph>
<paragraph id="69B5C7291028BD8C99F628C7F46A49D7" lastPageId="29" lastPageNumber="106" pageId="28" pageNumber="105">
Metasoma length / head+mesosoma length: &lt;1. Metasoma lateral height / metasoma lateral length: &lt;1. 2nd metasomal tergite length dorsal view / length of meta
<pageBreakToken id="235285C6AFDFD0DAF06396184C39553D" pageId="29" pageNumber="106" start="start">soma</pageBreakToken>
dorsal view: 1/3. Second metasomal tergite pilosity: present mediolaterally. Second metasomal tergite sculpture: smooth. Second metasomal tergite reflectivity: NOT CODED. Second metasomal tergite pilosity density: rare. Metasomal tergites 3-6 sculpture: smooth, no micropunctures. Metasomal tergites 3-6 reflectivity: glossy. Metasomal tergites 3-6 pilosity: absent. Prominent part of ventral spine of hypopygium length ventral view / Prominent part of ventral spine of hypopygium width ventral view: 1.16. Hypopygial setae apical end: not extending beyond posterior end of ventral spine of hypopygium.
</paragraph>
<paragraph id="E7D73BBCA1C0918BE2BF1C17B7DD3780" pageId="29" pageNumber="106">Body length: 1.90-2.10 mm (n = 3).</paragraph>
<paragraph id="CC38AC14C9397A691E4A776642FE89A2" pageId="29" pageNumber="106">Male: Similar to female. Color: Head, mesosoma and metasoma dark brown; antenna brown. Eye size vs female eye size: Eye larger in male. Anterior tentorial pit size: NOT CODED. Diameter of torulus / intertorular distance: NOT CODED.Diameter of lateral ocellus vs diameter of female lateral ocellus: 1.76. Flagellomeres count: 13. Antenna length / body length: 1. F1 shape: F1 curved and broadened apically. F1 length / F2 length: 1.07. Flagellomeres relative length: F2&gt;F3&gt;F4&gt;F5&gt;F6&gt;F9&gt;F10&gt;F11&gt;F12. F13 length / F12 length:&gt;1. Placoid sensilla present on: F2-F13. Body length: 1.9 mm (n=1).</paragraph>
<paragraph id="D1104EACCBE477709A8D134E7D0B87CC" pageId="29" pageNumber="106">
Gall (Figs
<figureCitation id="4516EC64ACF0FADE6FA4E760365C7387" captionStart="Figures 6163" captionStartId="F11" captionText="Figures 61 - 63. Dryocosmus harrisonae sp. n. 61 female, metasoma, lateral view 62 - 63 galls (photos by C. - T. Tang)." figureDoi="10.3897/jhr.53.9890.figures61-63" httpUri="https://binary.pensoft.net/fig/115550" pageId="29" pageNumber="106">62-63</figureCitation>
): Galls are located on axillary buds or young leaves on young shoots. The cross-section of the gall is triangular, with a short stalk connected to the host-plant. The gall is greenish, with a brownish tip and a greenish-brown central region, 3.9-4.6 mm long and 1.7-2.0 mm broad (n=2).
</paragraph>
<caption id="0EB70B40FBDB09E0C8B00224116BAE4D" ID-Zenodo-Dep="984531" doi="10.3897/jhr.53.9890.figures48-55" httpUri="https://binary.pensoft.net/fig/115548" pageId="29" pageNumber="106" start="Figures 4855" startId="F9">
<paragraph id="471F005EB115C3758F2C0C1A1D4DE05A" pageId="29" pageNumber="106">
<emphasis id="618A50D6EE60E3749F336290EDA3A04E" bold="true" pageId="29" pageNumber="106">Figures 48-55.</emphasis>
<taxonomicName id="FA943B59FEDC4DE0CECE5985D007ED74" authorityName="Melika &amp; Tang" authorityYear="2016" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus harrisonae" order="Hymenoptera" pageId="29" pageNumber="106" phylum="Arthropoda" rank="species" species="harrisonae">
<emphasis id="DDF923DB1829C2D79F5D0B529D6C5194" italics="true" pageId="29" pageNumber="106">Dryocosmus harrisonae</emphasis>
</taxonomicName>
sp. n.
<emphasis id="40431EA1A3C5A1AA6341CB699800537D" bold="true" pageId="29" pageNumber="106">48-50</emphasis>
head, female:
<emphasis id="22B7EAE58A35C904B4C04AFADC1F7EF7" bold="true" pageId="29" pageNumber="106">48</emphasis>
frontal view
<emphasis id="2D7F1EBFF491CDDCD20D842FE9581478" bold="true" pageId="29" pageNumber="106">49</emphasis>
dorsal view
<emphasis id="B1502CB12DF74B823FEF266FA963B854" bold="true" pageId="29" pageNumber="106">50</emphasis>
posterior view
<emphasis id="4BA57C7371DE5EEEFAC34DE166F6DF87" bold="true" pageId="29" pageNumber="106">51-52</emphasis>
head, male:
<emphasis id="2D44F37CF75533B70F8BFA795661529F" bold="true" pageId="29" pageNumber="106">51</emphasis>
frontal view
<emphasis id="01CEE65904EFE5AAC51065D0E5059F57" bold="true" pageId="29" pageNumber="106">52</emphasis>
dorsal view
<emphasis id="ACF89A35AAB55EEA0FB53AE1B0057567" bold="true" pageId="29" pageNumber="106">53-54</emphasis>
antenna:
<emphasis id="60154F2657B42DBEFCDE57C7DEFC324E" bold="true" pageId="29" pageNumber="106">53</emphasis>
female
<emphasis id="61018529E0EBC87DBD13CF0DC396CF27" bold="true" pageId="29" pageNumber="106">54</emphasis>
male
<emphasis id="39B3CECEB7BE5940F63B53380B48FAD4" bold="true" pageId="29" pageNumber="106">55</emphasis>
mesosoma, female, lateral view (gen=gena, spe=speculum).
</paragraph>
</caption>
<caption id="779BF728BF3A803899CEF88D1EECAC48" ID-Zenodo-Dep="984469" doi="10.3897/jhr.53.9890.figures56-60" httpUri="https://binary.pensoft.net/fig/115549" pageId="29" pageNumber="106" start="Figures 5660" startId="F10">
<paragraph id="F09C74BF8A36E1B75235571F32DD8FF2" pageId="29" pageNumber="106">
<emphasis id="1A3D9B0913DD724D637FEE64C5717134" bold="true" pageId="29" pageNumber="106">Figures 56-60.</emphasis>
<taxonomicName id="EE74B6546ABABC7BD4F2246D2227ADCD" authorityName="Melika &amp; Tang" authorityYear="2016" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus harrisonae" order="Hymenoptera" pageId="29" pageNumber="106" phylum="Arthropoda" rank="species" species="harrisonae">
<emphasis id="62BA92A9BEABD190F4801503C81CB450" italics="true" pageId="29" pageNumber="106">Dryocosmus harrisonae</emphasis>
</taxonomicName>
, female sp. n.
<emphasis id="AC14549322F353745908C7B279871D7E" bold="true" pageId="29" pageNumber="106">56</emphasis>
mesoscutum, dorsal view
<emphasis id="36A391F66F81CB5FC8695594D35C6C60" bold="true" pageId="29" pageNumber="106">57</emphasis>
mesoscutellum, dorsal view
<emphasis id="FF427C88485D9CB38A1A74DD6C37F71E" bold="true" pageId="29" pageNumber="106">58</emphasis>
pronotum and propleuron, frontal view
<emphasis id="0C9DD8F1863C771D0397DFA8DACDF610" bold="true" pageId="29" pageNumber="106">59</emphasis>
metascutellum and propodeum, posterodorsal view
<emphasis id="621E01AB01B61A9B38DAC9A05F0FABEE" bold="true" pageId="29" pageNumber="106">60</emphasis>
fore wing, part.
</paragraph>
</caption>
<caption id="FBE8DC5E094F85417325B2A58AA6E35E" ID-Zenodo-Dep="984471" doi="10.3897/jhr.53.9890.figures61-63" httpUri="https://binary.pensoft.net/fig/115550" pageId="29" pageNumber="106" start="Figures 6163" startId="F11">
<paragraph id="F61DCC187BE7603D350518C1E63DDA80" pageId="29" pageNumber="106">
<emphasis id="C55470A5F1717A4468B50FB3F135BDA4" bold="true" pageId="29" pageNumber="106">Figures 61-63.</emphasis>
<taxonomicName id="91E5F6014E30874D25D2D9C0B005B060" authorityName="Melika &amp; Tang" authorityYear="2016" class="Insecta" family="Fagaceae" genus="Dryocosmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryocosmus harrisonae" order="Hymenoptera" pageId="29" pageNumber="106" phylum="Arthropoda" rank="species" species="harrisonae">
<emphasis id="52E0E8E4D94631730F7A2E022A29E593" italics="true" pageId="29" pageNumber="106">Dryocosmus harrisonae</emphasis>
</taxonomicName>
sp. n.
<emphasis id="9B8D075BE77D460FA28A34D9AFAC5A4F" bold="true" pageId="29" pageNumber="106">61</emphasis>
female, metasoma, lateral view
<emphasis id="9ADFDBC0B6C52A4C577BC6F465E30042" bold="true" pageId="29" pageNumber="106">62-63</emphasis>
galls (photos by C.-T. Tang).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="885605A95BAFB015DC88D51A427AB008" pageId="30" pageNumber="107" type="biology_ecology">
<paragraph id="CCD2967CBFF072E15A797AF803D06E8C" pageId="30" pageNumber="107">
<pageBreakToken id="5E726EE349A232435FDA62579FB038A8" pageId="30" pageNumber="107" start="start">Biology</pageBreakToken>
.
</paragraph>
<paragraph id="BCDD7C23FBAB657F6BE261732681560F" pageId="30" pageNumber="107">
Only the sexual generation is known. Galls are on young shoots of
<taxonomicName id="BC0EDFDDFE0BC278858192BFE8A1ED84" class="Hexapoda" family="Fagaceae" genus="Castanopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Castanopsis echinocarpa" order="Hymenoptera" pageId="30" pageNumber="107" phylum="Arthropoda" rank="species" species="echinocarpa">
<emphasis id="E698F65F9B87C605AE956DAF18B143B8" italics="true" pageId="30" pageNumber="107">Castanopsis echinocarpa</emphasis>
</taxonomicName>
. Galls were collected in April; adults emerged from galls under laboratory conditions immediately after field collection.
</paragraph>
</subSubSection>
<subSubSection id="A3A789B3098EC1576670DD9EEF77559E" pageId="30" pageNumber="107" type="distribution">
<paragraph id="454E43612464342718F6DEF920491B07" pageId="30" pageNumber="107">Distribution.</paragraph>
<paragraph id="3D47C719A6399BFF2D02B8E9084D28DC" pageId="30" pageNumber="107">China: Yunnan Province.</paragraph>
</subSubSection>
</treatment>
</document>