treatments-xml/data/9A/22/25/9A222597BC0C159C77FD3828B3BC4F4C.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.524.5990" ID-GBIF-Dataset="44bd24bf-bbc8-4368-a1c8-d63f14a1f5d4" ID-PMC="PMC4602289" ID-Pensoft-Pub="1313-2970-524-17" ID-PubMed="26478704" ID-ZBK="B1F1F6385E5C48BA9B1573F4E94B5C0E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-524-17" ModsDocOrigin="ZooKeys 524" ModsDocTitle="Taxonomy and distribution of some katydids (OrthopteraTettigoniidae) from tropical Africa" checkinTime="1451243934978" checkinUser="pensoft" docAuthor="Massa, Bruno" docDate="2015" docId="9A222597BC0C159C77FD3828B3BC4F4C" docLanguage="en" docName="ZooKeys 524: 17-44" docOrigin="ZooKeys 524" docSource="http://dx.doi.org/10.3897/zookeys.524.5990" docTitle="Horatosphaga aethiopica Massa, 2015, sp. n." docType="treatment" docUuid="C13E52EB-70F7-4D4F-AA03-BD41E1D416DA" docUuidSource="ZooBank" docVersion="4" lastPageNumber="22" masterDocId="835988306165E3338C795A09FFA2FFC5" masterDocTitle="Taxonomy and distribution of some katydids (OrthopteraTettigoniidae) from tropical Africa" masterLastPageNumber="44" masterPageNumber="17" pageNumber="22" updateTime="1668160808894" updateUser="ExternalLinkService">
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<mods:title>Taxonomy and distribution of some katydids (OrthopteraTettigoniidae) from tropical Africa</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Massa, Bruno</mods:namePart>
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<mods:date>2015</mods:date>
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<mods:number>524</mods:number>
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<mods:start>17</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.524.5990</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.524.5990</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-524-17</mods:identifier>
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<treatment ID-GBIF-Taxon="152062535" LSID="urn:lsid:zoobank.org:act:C13E52EB-70F7-4D4F-AA03-BD41E1D416DA" httpUri="http://treatment.plazi.org/id/9A222597BC0C159C77FD3828B3BC4F4C" lastPageNumber="22" pageId="5" pageNumber="22">
<subSubSection pageId="5" pageNumber="22" type="multiple">
<paragraph pageId="5" pageNumber="22">Taxon classification Animalia Orthoptera Tettigoniidae</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="nomenclature">
<paragraph pageId="5" pageNumber="22">
<taxonomicName LSID="http://zoobank.org/C13E52EB-70F7-4D4F-AA03-BD41E1D416DA" class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="22">sp. n.</taxonomicNameLabel>
Figs 17-21, 22-24
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="materials_examined">
<paragraph pageId="5" pageNumber="22">Material examined and depository.</paragraph>
<paragraph pageId="5" pageNumber="22">
Ethiopia, Omo river, El Dire
<geoCoordinate direction="north" orientation="latitude" precision="1" value="5.1059585">5°06'21.45&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="36.852436">36°51'08.77&quot;E</geoCoordinate>
(950 m) 21.V.1939 (Expedition E. Zavattari) (♂ holotype, ♀ allotype, ♂ paratype); Ethiopia, Omo river, El Dire (950 m) 19.V.1939 (Expedition E. Zavattari) (♀ paratype); Ethiopia, Omo river, Calam
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.688911">4°41'20.08&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="35.66624">35°39'58.46&quot;E</geoCoordinate>
(370 m) 14.VIII.1939 (Expedition E. Zavattari) (♀ paratype)) (MZR).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="colour">
<paragraph pageId="5" pageNumber="22">Colour.</paragraph>
<paragraph pageId="5" pageNumber="22">Yellowish (alive specimens may show different colour) (Figs 17, 22).</paragraph>
<caption pageId="5" pageNumber="22">
<paragraph pageId="5" pageNumber="22">
Figures 17-21.
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
sp. n. Dorsal view of male (17); lateral view of last abdominal tergites of male (18); sub-genital plate of male (19); dorsal view of last abdominal tergite, cerci and appendices of the sub-genital plate of male (20); left tegmen of male (21).
</paragraph>
</caption>
<caption pageId="5" pageNumber="22">
<paragraph pageId="5" pageNumber="22">
Figures 22-25. Dorsal view of head, pronotum and tegmina of female of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
sp. n. (22); lateral view of the ovipositor (23); stridulatory file of male of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
sp. n. (24); stridulatory file of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga leggei" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="leggei">Horatosphaga leggei</taxonomicName>
(25).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="description">
<paragraph pageId="5" pageNumber="22">Description.</paragraph>
<paragraph pageId="5" pageNumber="22">Male. Head and antennae: fastigium of vertex very narrow, furrowed above, separated from the tuberculated fastigium of frons. Eyes rounded, well projecting. Legs long. Fore coxae unarmed. Fore tibiae furrowed on upper margin, distinctly widening above tympanum, which is closed on inner and on outer sides, tympanic auricles inflated. Fore femora unarmed, fore tibiae with 11 spines plus 1 spur on inner and outer ventral margins, 1 spur on inner and outer dorsal margins, mid femora unarmed, mid tibiae with 14 spines on inner and outer ventral margins, plus 1 spur on each side and 1 spur on both sides of dorsal margins, hind femora unarmed, hind tibiae with many spines on ventral and dorsal margins and 3 spurs on each side. Thorax: pronotum little narrowing anteriorly, little raised posteriorly, anterior margin straight, posterior margin rounded, humeral sinus absent, lobes of pronotum rounded and low. Tegmina shorter than abdomen, with pointed apices, their web-like venation very simple, cross-veins of area MA are arranged in a parallel fashion, a bit arcuate in inner part (Fig. 21). Hind wings rudimentary and linked to metanotum. Stridulatory region of left tegmen short and inflated, stridulatory file composed of 20-25 widely spaced teeth, of different size (Fig. 24). Abdomen: tenth tergite greatly enlarged and completely concealing supra-anal plate, in lateral view similar to a raptor beak (Figs 18-20); sub-genital plate long, up-curved and deeply divided into two lobes reaching margin of the tenth tergite and curved a bit backwards at end (Fig. 19); styli absent. Cerci short, in-curved, with a small apical spine (Fig. 20).</paragraph>
<paragraph pageId="5" pageNumber="22">Female. As the male, but tegmina reduced to two small overlapping scales, not exceeding first abdominal tergite, but showing a residual web-like venation (Fig. 22). Cerci conical and pointed. Ovipositor up-curved and provided with many denticles on upper and lower margins (Fig. 23).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="measurements">
<paragraph pageId="5" pageNumber="22">Measurements.</paragraph>
<paragraph pageId="5" pageNumber="22">
See Table 2.
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
is characterized by its very small size, compared to related species, in particular in the length of tegmina and ovipositor.
</paragraph>
<caption pageId="5" pageNumber="22">
<paragraph pageId="5" pageNumber="22">
Table 2. Measurements of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
sp. n. and the related three species
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga ruspolii" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="ruspolii">Horatosphaga ruspolii</taxonomicName>
,
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga vicina" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="vicina">Horatosphaga vicina</taxonomicName>
and
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga diminuta" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="diminuta">Horatosphaga diminuta</taxonomicName>
(after
<bibRefCitation author="Ragge, DR" journalOrPublisher="Bulletin of the British Museum (Natural History) Entomology" pageId="16" pageNumber="33" pagination="269 - 333" title="The Acrometopae of the Ethiopian Region: a revision, with notes on the sexual dimorphism shown by the group (Orthoptera: Tettigoniidae)." url="http://www.biodiversitylibrary.org/page/2245318page/386/mode/1up" volume="8" year="1960">Ragge 1960</bibRefCitation>
). For all species the min-max range is reported.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="22">
<table pageId="5" pageNumber="22">
<tr pageId="5" pageNumber="22">
<th colspan="1" pageId="5" pageNumber="22" rowspan="1">Species</th>
<th colspan="1" pageId="5" pageNumber="22" rowspan="1">Total length</th>
<th colspan="1" pageId="5" pageNumber="22" rowspan="1">Pronotum length</th>
<th colspan="1" pageId="5" pageNumber="22" rowspan="1">Length of hind femora</th>
<th colspan="1" pageId="5" pageNumber="22" rowspan="1">Length of tegmina</th>
<th colspan="1" pageId="5" pageNumber="22" rowspan="1">Length of ovipositor</th>
</tr>
<tr pageId="5" pageNumber="22">
<td colspan="1" pageId="5" pageNumber="22" rowspan="1">
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga ruspolii" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="ruspolii">Horatosphaga ruspolii</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="22">
<td colspan="1" pageId="5" pageNumber="22" rowspan="1">
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga vicina" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="vicina">Horatosphaga vicina</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="22">
<td colspan="1" pageId="5" pageNumber="22" rowspan="1">
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga diminuta" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="diminuta">Horatosphaga diminuta</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="22">
<td colspan="1" pageId="5" pageNumber="22" rowspan="1">
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="diagnosis">
<paragraph pageId="5" pageNumber="22">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="22">
Concerning the affinities between this and related species, there are only three
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">Horatosphaga</taxonomicName>
with the male having the tenth abdominal tergite greatly enlarged and completely concealing the supra-anal plate, namely
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga ruspolii" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="ruspolii">Horatosphaga ruspolii</taxonomicName>
(Shulthess, 1898),
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga diminuta" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="diminuta">Horatosphaga diminuta</taxonomicName>
(Chopard, 1954) and
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga vicina" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="vicina">Horatosphaga vicina</taxonomicName>
(Chopard, 1954), all described from Kenya.
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga ruspolii" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="ruspolii">Horatosphaga ruspolii</taxonomicName>
has fully developed wings, while the other two have reduced fore wings and rudimentary hind wings. According to
<bibRefCitation author="Ragge, DR" journalOrPublisher="Bulletin of the British Museum (Natural History) Entomology" pageId="16" pageNumber="33" pagination="269 - 333" title="The Acrometopae of the Ethiopian Region: a revision, with notes on the sexual dimorphism shown by the group (Orthoptera: Tettigoniidae)." url="http://www.biodiversitylibrary.org/page/2245318page/386/mode/1up" volume="8" year="1960">Ragge (1960)</bibRefCitation>
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga ruspolii" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="ruspolii">Horatosphaga ruspolii</taxonomicName>
lives also in the eastern Ethiopia, in the area between Ethiopia and Somalia, and in Uganda. Its external genitalia are very variable, but appendices of the sub-genital plate are simply up-curved and shorter than those of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
sp. n. However,
<bibRefCitation author="Ragge, DR" journalOrPublisher="Bulletin of the British Museum (Natural History) Entomology" pageId="16" pageNumber="33" pagination="269 - 333" title="The Acrometopae of the Ethiopian Region: a revision, with notes on the sexual dimorphism shown by the group (Orthoptera: Tettigoniidae)." url="http://www.biodiversitylibrary.org/page/2245318page/386/mode/1up" volume="8" year="1960">Ragge (1960)</bibRefCitation>
suspected that
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga diminuta" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="diminuta">Horatosphaga diminuta</taxonomicName>
could be a brachypterous form of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga ruspolii" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="ruspolii">Horatosphaga ruspolii</taxonomicName>
, and considered also that
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga vicina" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="vicina">Horatosphaga vicina</taxonomicName>
could be a large form of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga diminuta" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="diminuta">Horatosphaga diminuta</taxonomicName>
, that also Chopard (in
<bibRefCitation author="Chopard, L" journalOrPublisher="Transactions Royal Entomological Society London" pageId="16" pageNumber="33" pagination="315 - 353" title="Orthoptera-Ensifera from northern Kenya and Jubaland." volume="105" year="1954">Chopard and Mc Kevan 1954</bibRefCitation>
) considered almost an exact repetition of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga vicina" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="vicina">Horatosphaga vicina</taxonomicName>
on a rather smaller scale.
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
sp. n., which is smaller than the above three species (see
<bibRefCitation author="Ragge, DR" journalOrPublisher="Bulletin of the British Museum (Natural History) Entomology" pageId="16" pageNumber="33" pagination="269 - 333" title="The Acrometopae of the Ethiopian Region: a revision, with notes on the sexual dimorphism shown by the group (Orthoptera: Tettigoniidae)." url="http://www.biodiversitylibrary.org/page/2245318page/386/mode/1up" volume="8" year="1960">Ragge 1960</bibRefCitation>
and Table 2), differs from the previous species not only by its external male genitalia, but also by the high reduction of wings, mainly in the female, a character that forces individuals to an important isolation. The reduction of wings probably was also the cause for a very reduced stridulatory file. Regarding the reduction of the stridulatory file related to the wing reduction, it is possible that also
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga diminuta" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="diminuta">Horatosphaga diminuta</taxonomicName>
has a different stridulatory file compared to the fully developed wings of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga ruspolii" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="ruspolii">Horatosphaga ruspolii</taxonomicName>
, and this should result in a different song, an important specific barrier.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="etymology">
<paragraph pageId="5" pageNumber="22">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="22">The Latin name aethiopica is a female adjective meaning &quot;living in Ethiopia&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="22" type="discussion">
<paragraph pageId="5" pageNumber="22">Discussion.</paragraph>
<paragraph pageId="5" pageNumber="22">
In 1939, between March and September,
<bibRefCitation author="Zavattari, E" journalOrPublisher="Rivista Biologia Coloniale, Roma" pageId="16" pageNumber="33" pagination="97 - 108" title="La missione biologica Sagan-Omo (1939)." volume="3" year="1943">Zavattari (1943)</bibRefCitation>
carried out an expedition to the territory of the Omo river in Ethiopia. During that trip, participants reached the northern part of the Turkana lake, where they collected also some
<taxonomicName lsidName="Ensifera" order="Orthoptera" pageId="5" pageNumber="22" rank="order">Orthoptera</taxonomicName>
. Among them there was the series of
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">Horatosphaga</taxonomicName>
specimens listed above. This new taxon is remarkably different from all related taxa. Indeed, in none of the species known till now, the tegmina of the female are so much reduced to two small scales, as in
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga aethiopica" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="aethiopica">Horatosphaga aethiopica</taxonomicName>
sp. n. Concerning the tribe
<taxonomicName lsidName="" pageId="5" pageNumber="22" rank="tribe" tribe="Acrometopini">Acrometopini</taxonomicName>
, the characters of this
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Horatosphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatosphaga" order="Orthoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">Horatosphaga</taxonomicName>
species and those of other related species as provided by
<bibRefCitation author="Hemp, C" journalOrPublisher="Zootaxa" pageId="16" pageNumber="33" pagination="63 - 68" title="Faunistic relations between the ranges of the Eastern Arc mountains: relocation of Peronurahildebrandtiana Karsch, 1889 (Tettigoniidae: Phaneropterinae), the description of the male and implications from its phylogeographical pattern." volume="3006" year="2011">Hemp (2011)</bibRefCitation>
and
<bibRefCitation author="Hemp, C" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="16" pageNumber="33" pagination="66 - 82" title="A new genus of African Acrometopini (Tettigoniidae: Phaneropterinae) based on morphology, chromosomes, acoustics, distribution, and molecular data, and the description of a new species." url="10.1111/j.1096-3642.2009.00542.x" volume="158" year="2010">Hemp et al. (2010)</bibRefCitation>
modify and update the key in
<bibRefCitation author="Ragge, DR" journalOrPublisher="Bulletin of the British Museum (Natural History) Entomology" pageId="16" pageNumber="33" pagination="269 - 333" title="The Acrometopae of the Ethiopian Region: a revision, with notes on the sexual dimorphism shown by the group (Orthoptera: Tettigoniidae)." url="http://www.biodiversitylibrary.org/page/2245318page/386/mode/1up" volume="8" year="1960">Ragge (1960)</bibRefCitation>
partially based on the ratio between the length of the pronotum to that of the tegmina.
</paragraph>
</subSubSection>
</treatment>
</document>