293 lines
37 KiB
XML
293 lines
37 KiB
XML
<document id="9D106450CAA02A1AE9809F2F925E3B04" ID-CLB-Dataset="30822" ID-DOI="10.5852/ejt.2018.401" ID-GBIF-Dataset="49d45069-6150-4b6e-873c-fc8058025a39" ID-Zenodo-Dep="1167581" ID-ZooBank="9C9DB157-AAA5-40B7-BA0B-9A57779382C1" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517911480053" checkinUser="plazi" docAuthor="Salnitska, Maria & Solodovnikov, Alexey" docDate="2018" docId="03D5FE7E7043BF27FD93F98CFBC9FA50" docLanguage="en" docName="ejt-401_salnitska_solodovnikov.pdf.imf" docOrigin="European Journal of Taxonomy 401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Quedius kalabi Smetana 1995" docType="treatment" docVersion="15" lastPageNumber="10" masterDocId="FFEC8606704BBF2EFFF0FFB0FFBBFFEB" masterDocTitle="Taxonomy of the poorly known Quedius mutilatus group of wingless montane species from Middle Asia (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="17" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547222236327" originalUpdateUser="yanell" pageNumber="9" updateTime="1698519557005" updateUser="ExternalLinkService">
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<mods:title id="57271F87055E21A315119B49EDD65EBC">Taxonomy of the poorly known Quedius mutilatus group of wingless montane species from Middle Asia (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
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<mods:namePart id="0165FC4E66DD03C70FF5B01233D82051">Salnitska, Maria</mods:namePart>
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<mods:namePart id="BA390A69D7A73FAEB6026BF7044E3119">Solodovnikov, Alexey</mods:namePart>
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<mods:typeOfResource id="B215889939E27F9FD675DCBF8727B75D">text</mods:typeOfResource>
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<mods:title id="B2797398F2017BB632B1FB280D9BC288">European Journal of Taxonomy</mods:title>
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<mods:part id="BC680853D1740E3EA648F159736E9CA0">
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<mods:date id="B28D8EFBFAA927A1E35F191DC5A9AC1B">2018</mods:date>
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<mods:number id="59C4052657E94CE6B9027A4ED4CFD0F1">2018-02-06</mods:number>
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<mods:number id="17E4CCF432E3F7551EE41757F6CFC7DD">401</mods:number>
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<treatment id="03D5FE7E7043BF27FD93F98CFBC9FA50" ID-DOI="http://doi.org/10.5281/zenodo.5695866" ID-GBIF-Taxon="141125768" ID-Zenodo-Dep="5695866" LSID="urn:lsid:plazi:treatment:03D5FE7E7043BF27FD93F98CFBC9FA50" httpUri="http://treatment.plazi.org/id/03D5FE7E7043BF27FD93F98CFBC9FA50" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<subSubSection id="C3661CE37043BF26FD93F98CFC6AF9BC" box="[611,977,1596,1623]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BC34F687043BF26FD93F98CFC6AF9BC" blockId="8.[611,977,1596,1660]" box="[611,977,1596,1623]" pageId="8" pageNumber="9">
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<heading id="D08BF8047043BF26FD93F98CFC6AF9BC" box="[611,977,1596,1623]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
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<taxonomicName id="4C7C34EB7043BF26FD93F98CFC6AF9BC" ID-CoL="4R2Q7" authority="Smetana, 1995" authorityName="Smetana" authorityYear="1995" box="[611,977,1596,1623]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kalabi">
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<emphasis id="B908937A7043BF26FD93F98CFCA0F9BC" bold="true" box="[611,795,1596,1623]" italics="true" pageId="8" pageNumber="9">Quedius kalabi</emphasis>
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<bibRefCitation id="EFED32997043BF26FCD2F98DFC6AF9BC" author="Smetana A." box="[802,977,1596,1623]" journalOrPublisher="Elytra (Tokyo)" pageId="8" pageNumber="9" pagination="77 - 88" part="23" refId="ref6582" refString="Smetana A. 1995. Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Elytra (Tokyo) 23: 77 - 88." title="Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini)" type="journal article" year="1995">Smetana, 1995</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3661CE37043BF26FD4CF9D2FCCCF997" box="[700,887,1634,1660]" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BC34F687043BF26FD4CF9D2FCCCF997" blockId="8.[611,977,1596,1660]" box="[700,887,1634,1660]" pageId="8" pageNumber="9">
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<figureCitation id="134753ED7043BF26FD4CF9D2FCBEF997" box="[700,773,1634,1660]" captionStart="Fig.1" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[221,1375,624,1881]" captionTargetId="figure@4.[208,1379,622,1889]" captionTargetPageId="4" captionText="Fig.1.DistributionandaedeagusvariabilityoftheQuediusmutilatusgroupintheTien-ShanMountains. Numbers are specified in Table 1. Empty symbols = typespcimens (for Q. kalabi precise locality of the holotype unknown);? = ambiguously labelled or undetermined material. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1167583/files/figure.png" pageId="8" pageNumber="9">Figs 1</figureCitation>
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,
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<figureCitation id="134753ED7043BF26FCE4F9D2FC9FF997" box="[788,804,1634,1660]" captionStart="Fig.3" captionStartId="10.[189,232,1964,1990]" captionTargetBox="[331,1257,336,1918]" captionTargetId="figure@10.[331,1257,334,1918]" captionTargetPageId="10" captionText="Fig.3.HolotypeofQuediuskalabiSmetana,1995.A.Habitus.B.Permanentpreparationoftheaedeagus. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1167587/files/figure.png" pageId="8" pageNumber="9">3</figureCitation>
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,
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<figureCitation id="134753ED7043BF26FCC1F9D2FCCCF997" box="[817,887,1634,1660]" captionStart="Fig. 4" captionStartId="11.[189,232,1858,1884]" captionTargetBox="[333,1256,265,1830]" captionTargetId="figure@11.[331,1256,265,1830]" captionTargetPageId="11" captionText="Fig. 4. Aedeagi in the Quedius mutilatus group. A–B. Quedius mutilatus Eppelsheim, 1888. C–D. Quedius kalabi Smetana, 1995. E–F. Quedius kungeicus sp. nov. G–N. Quedius equus Smetana, 2014 (Karkara Valley, Kazakhstan), variability of the structure of the aedeagus.A, C, E, G, I, K, M = median lobe, laterally. B, D, F, H, J, L, N = paramere (dissected from median lobe), underside (side facing median lobe). Scale bars: 1 mm." httpUri="https://zenodo.org/record/1167589/files/figure.png" pageId="8" pageNumber="9">4C–D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3661CE37043BF26FF4DF916FE18F8C0" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="8BC34F687043BF26FF4DF916FCE1F92A" blockId="8.[189,858,1702,1729]" box="[189,858,1702,1729]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="AB6C68467043BF26FF4DF916FCE1F92A" box="[189,858,1702,1729]" pageId="8" pageNumber="9">
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<taxonomicName id="4C7C34EB7043BF26FF4DF916FDE9F92A" ID-CoL="4R2Q7" authority="Smetana, 1995: 77" authorityName="Smetana" authorityPageNumber="77" authorityYear="1995" box="[189,594,1702,1729]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kalabi">
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<emphasis id="B908937A7043BF26FF4DF916FED4F92B" box="[189,367,1702,1728]" italics="true" pageId="8" pageNumber="9">Quedius kalabi</emphasis>
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<treatmentCitation id="0ADD69797043BF26FE87F916FDE9F92A" author="Smetana" box="[375,594,1702,1729]" page="77" pageId="8" pageNumber="9" year="1995">
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<bibRefCitation id="EFED32997043BF26FE87F916FD92F92A" author="Smetana A." box="[375,553,1702,1729]" journalOrPublisher="Elytra (Tokyo)" pageId="8" pageNumber="9" pagination="77 - 88" part="23" refId="ref6582" refString="Smetana A. 1995. Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Elytra (Tokyo) 23: 77 - 88." title="Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini)" type="journal article" year="1995">Smetana, 1995</bibRefCitation>
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: 77
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</treatmentCitation>
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</taxonomicName>
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(original description).
|
||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8BC34F687043BF26FF4DF95AFE18F8C0" blockId="8.[189,1399,1770,1835]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="AB6C68467043BF26FF4DF95AFE18F8C0" pageId="8" pageNumber="9">
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<taxonomicName id="4C7C34EB7043BF26FF4DF95AFED5F8EF" ID-CoL="4R2Q7" authorityName="Smetana" authorityYear="1995" box="[189,366,1770,1796]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kalabi">
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<emphasis id="B908937A7043BF26FF4DF95AFED5F8EF" box="[189,366,1770,1796]" italics="true" pageId="8" pageNumber="9">Quedius kalabi</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0ADD69797043BF26FE7AF95AFDD2F8EF" author="Smetana" box="[394,617,1770,1796]" page="118" pageId="8" pageNumber="9" year="1998">
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||
<bibRefCitation id="EFED32997043BF26FE7AF95AFD8FF8EF" author="Smetana A." box="[394,564,1770,1796]" journalOrPublisher="Elytra (Tokyo)" pageId="8" pageNumber="9" pagination="115 - 128" part="26" refId="ref6615" refString="Smetana A. 1998. Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Part 2. Elytra (Tokyo) 26 (1): 115 - 128." title="Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Part 2" type="journal article" year="1998">Smetana 1998</bibRefCitation>
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||
: 118
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||
</treatmentCitation>
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||
(comparison with
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<taxonomicName id="4C7C34EB7043BF26FCB2F95BFC69F8EF" ID-CoL="4R2SL" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[834,978,1770,1797]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="mutilatus">
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<emphasis id="B908937A7043BF26FCB2F95BFC69F8EF" box="[834,978,1770,1797]" italics="true" pageId="8" pageNumber="9">Q. mutilatus</emphasis>
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</taxonomicName>
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||
). —
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<treatmentCitation id="0ADD69797043BF26FBFDF95AFF42F8C0" author="Solodovnikov" page="7" pageId="8" pageNumber="9" year="2016">
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||
<bibRefCitation id="EFED32997043BF26FBFDF95AFAC8F8EF" author="Solodovnikov A. & Hansen A. K." box="[1037,1395,1770,1797]" journalOrPublisher="Zootaxa" pageId="8" pageNumber="9" pagination="475 - 490" part="4170" publicationUrl="https://doi.org/10.11646/zootaxa.4170.3.3" refId="ref6763" refString="Solodovnikov A. & Hansen A. K. 2016. Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini). Zootaxa 4170 (3): 475 - 490. https: // doi. org / 10.11646 / zootaxa. 4170.3.3" title="Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini)" type="journal article" year="2016">Solodovnikov & Hansen 2016</bibRefCitation>
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: 7
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</treatmentCitation>
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(distribution).
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||
</treatmentCitationGroup>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C3661CE37043BF27FF4DF8E4FD12FE9A" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8BC34F687043BF26FF4DF8E4FE11F885" blockId="8.[189,426,1876,1902]" box="[189,426,1876,1902]" pageId="8" pageNumber="9">
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<heading id="D08BF8047043BF26FF4DF8E4FE11F885" bold="true" box="[189,426,1876,1902]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
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<emphasis id="B908937A7043BF26FF4DF8E4FE11F885" bold="true" box="[189,426,1876,1902]" pageId="8" pageNumber="9">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC34F687043BF26FF1AF835FE22F874" blockId="8.[189,1398,1925,2027]" box="[234,409,1925,1951]" pageId="8" pageNumber="9">
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<emphasis id="B908937A7043BF26FF1AF835FE22F874" bold="true" box="[234,409,1925,1951]" pageId="8" pageNumber="9">Type material</emphasis>
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</paragraph>
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<paragraph id="8BC34F687043BF26FF4DF81BFE61F800" blockId="8.[189,1398,1925,2027]" pageId="8" pageNumber="9">
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<materialsCitation id="3B1445357043BF26FF4DF81BFE61F800" ID-GBIF-Occurrence="1821772567" collectionCode="NMW" country="Kyrgyzstan" elevation="3300" location="SU - Tien Shan NE part of Terskey Ala Too [Terskei Alatau] ridge" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<collectingCountry id="F36B0FF87043BF26FF4DF81BFE3DF82E" box="[189,390,1963,1989]" name="Kyrgyzstan" pageId="8" pageNumber="9">KYRGYZSTAN</collectingCountry>
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:
|
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<typeStatus id="54C7F1CA7043BF26FE64F81BFDBFF82E" box="[404,516,1963,1989]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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,
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||
<specimenCount id="9D7A84E17043BF26FDE3F81BFD91F82E" box="[531,554,1963,1989]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
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, “
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<location id="8EA319B37043BF26FDB5F81BFACDF82E" LSID="urn:lsid:plazi:treatment:03D5FE7E7043BF27FD93F98CFBC9FA50:8EA319B37043BF26FDB5F81BFACDF82E" box="[581,1398,1962,1989]" country="Kyrgyzstan" name="SU - Tien Shan NE part of Terskey Ala Too [Terskei Alatau] ridge" pageId="8" pageNumber="9">SU – Tien Shan NE part of Terskey Ala Too [Terskei Alatau] ridge</location>
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<quantity id="4C84E28D7043BF26FF4DF861FEDAF800" box="[189,353,2001,2027]" metricMagnitude="3" metricUnit="m" metricValue="3.3" metricValueMax="3.6" metricValueMin="3.0" pageId="8" pageNumber="9" unit="m" value="3300.0" valueMax="3600.0" valueMin="3000.0">
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<elevation id="0051A85B7043BF26FF4DF861FEDAF800" box="[189,353,2001,2027]" metricMagnitude="3" metricUnit="m" metricValue="3.3" metricValueMax="3.6" metricValueMin="3.0" pageId="8" pageNumber="9" unit="m" value="3300.0" valueMax="3600.0" valueMin="3000.0">3000–3600 m</elevation>
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</quantity>
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” (
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<collectionCode id="ED6DD7AD7043BF26FE8CF861FE75F800" box="[380,462,2001,2027]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9">NMW</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC34F687042BF27FF1AFEBBFE5AFECE" blockId="9.[189,1398,267,369]" box="[234,481,267,293]" pageId="9" pageNumber="10">
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<emphasis id="B908937A7042BF27FF1AFEBBFE5AFECE" bold="true" box="[234,481,267,293]" pageId="9" pageNumber="10">Additional material</emphasis>
|
||
</paragraph>
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<paragraph id="8BC34F687042BF27FF4DFE81FD12FE9A" blockId="9.[189,1398,267,369]" pageId="9" pageNumber="10">
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<materialsCitation id="3B1445357042BF27FF4DFE81FD12FE9A" ID-GBIF-Occurrence="1821772623" collectingDate="1993-06-16" collectionCode="cSch" collectorName="S. Ovchinnikov" country="Kyrgyzstan" elevation="2600" latitude="42.44" location="E TerskeiAlatau" longLatPrecision="6" longitude="78.52" municipality="Teploklyuchenka Village" pageId="9" pageNumber="10" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
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<collectingCountry id="F36B0FF87042BF27FF4DFE81FE3DFEA0" box="[189,390,305,331]" name="Kyrgyzstan" pageId="9" pageNumber="10">KYRGYZSTAN</collectingCountry>
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:
|
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<specimenCount id="9D7A84E17042BF27FE7EFE81FE03FEA0" box="[398,440,305,331]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
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,
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||
<specimenCount id="9D7A84E17042BF27FE33FE81FE57FEA0" box="[451,492,305,331]" pageId="9" pageNumber="10" type="female">1 ♀</specimenCount>
|
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,
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||
<location id="8EA319B37042BF27FE08FE81FD03FEA1" LSID="urn:lsid:plazi:treatment:03D5FE7E7043BF27FD93F98CFBC9FA50:8EA319B37042BF27FE08FE81FD03FEA1" box="[504,696,304,331]" country="Kyrgyzstan" latitude="42.44" longLatPrecision="6" longitude="78.52" municipality="Teploklyuchenka Village" name="E TerskeiAlatau" pageId="9" pageNumber="10">E TerskeiAlatau</location>
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,
|
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<collectingMunicipality id="6BA7D5127042BF27FD31FE80FC5DFEA1" box="[705,998,304,330]" country="Kyrgyzstan" latitude="42.44" longLatPrecision="6" longitude="78.52" name="Teploklyuchenka Village" pageId="9" pageNumber="10">Teploklyuchenka Village</collectingMunicipality>
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,
|
||
<elevation id="0051A85B7042BF27FC1FFE80FBFEFEA0" box="[1007,1093,304,331]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="10" unit="m" value="2600.0">
|
||
<quantity id="4C84E28D7042BF27FC1FFE80FBFEFEA0" box="[1007,1093,304,331]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="10" unit="m" value="2600.0">2600 m</quantity>
|
||
</elevation>
|
||
[
|
||
<geoCoordinate id="EE4829AF7042BF27FBA0FE81FB60FEA0" box="[1104,1243,305,331]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="5" value="42.44">42.4400° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4829AF7042BF27FB14FE80FAD2FEA0" box="[1252,1385,304,331]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="5" value="78.52">78.5200° E</geoCoordinate>
|
||
],
|
||
<date id="FFC269A87042BF27FF4DFEE6FEEDFE9B" box="[189,342,342,369]" pageId="9" pageNumber="10" value="1993-06-16">
|
||
<collectingDate id="EF8690407042BF27FF4DFEE6FEEDFE9B" box="[189,342,342,369]" pageId="9" pageNumber="10" value="1993-06-16">16 Jun. 1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26892ABE7042BF27FE92FEE6FDA1FE9B" box="[354,538,342,368]" pageId="9" pageNumber="10">S. Ovchinnikov</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED6DD7AD7042BF27FDADFEE6FD27FE9A" box="[605,668,342,369]" pageId="9" pageNumber="10">cSch</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3661CE37042BF27FF4DFE2AFD9DFD4A" pageId="9" pageNumber="10" type="distribution">
|
||
<paragraph id="8BC34F687042BF27FF4DFE2AFDA8FE5F" blockId="9.[189,531,410,436]" box="[189,531,410,436]" pageId="9" pageNumber="10">
|
||
<heading id="D08BF8047042BF27FF4DFE2AFDA8FE5F" bold="true" box="[189,531,410,436]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B908937A7042BF27FF4DFE2AFDA8FE5F" bold="true" box="[189,531,410,436]" pageId="9" pageNumber="10">Distribution and bionomics</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC34F687042BF27FF4DFE7BFD9DFD4A" blockId="9.[189,1399,459,673]" pageId="9" pageNumber="10">
|
||
<taxonomicName id="4C7C34EB7042BF27FF4DFE7BFECFFE0E" authorityName="Smetana" authorityYear="1995" box="[189,372,459,485]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FF4DFE7BFECFFE0E" box="[189,372,459,485]" italics="true" pageId="9" pageNumber="10">Quedius kalabi</emphasis>
|
||
</taxonomicName>
|
||
is known from the three specimens examined here, collected in the Terskei Alatau mountain range at high elevations around
|
||
<quantity id="4C84E28D7042BF27FD4EFE40FCD8FDE0" box="[702,867,496,523]" metricMagnitude="3" metricUnit="m" metricValue="3.1" metricValueMax="3.6" metricValueMin="2.6" pageId="9" pageNumber="10" unit="m" value="3100.0" valueMax="3600.0" valueMin="2600.0">2600–3600 m</quantity>
|
||
(
|
||
<figureCitation id="134753ED7042BF27FC86FE40FC05FDE1" box="[886,958,496,522]" captionStart="Fig.1" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[221,1375,624,1881]" captionTargetId="figure@4.[208,1379,622,1889]" captionTargetPageId="4" captionText="Fig.1.DistributionandaedeagusvariabilityoftheQuediusmutilatusgroupintheTien-ShanMountains. Numbers are specified in Table 1. Empty symbols = typespcimens (for Q. kalabi precise locality of the holotype unknown);? = ambiguously labelled or undetermined material. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1167583/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
|
||
). Only two of them bear labels that specify the date and exact collecting locality (
|
||
<date id="FFC269A87042BF27FD12FDA6FCC4FDDB" box="[738,895,534,560]" pageId="9" pageNumber="10" value="1993-06-16">16 Jun. 1993</date>
|
||
, Teploklyuchenka Village). All previously published material for
|
||
<taxonomicName id="4C7C34EB7042BF27FE29FD8CFDF7FDBE" authorityName="Smetana" authorityYear="1995" box="[473,588,571,598]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FE29FD8CFDF7FDBE" box="[473,588,571,598]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
, except the
|
||
<typeStatus id="54C7F1CA7042BF27FD1BFD8BFCEEFDBE" box="[747,853,571,597]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
, but including the
|
||
<typeStatus id="54C7F1CA7042BF27FBB6FD8CFB15FDBD" box="[1094,1198,572,598]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
|
||
, was apparently misidentified (see below). Like other species of the
|
||
<taxonomicName id="4C7C34EB7042BF27FCECFDD2FC17FD90" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[796,940,609,636]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7042BF27FCECFDD2FC17FD90" box="[796,940,609,636]" italics="true" pageId="9" pageNumber="10">Q. mutilatus</emphasis>
|
||
</taxonomicName>
|
||
group,
|
||
<taxonomicName id="4C7C34EB7042BF27FBF7FDD2FBCFFD90" authorityName="Smetana" authorityYear="1995" box="[1031,1140,609,636]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FBF7FDD2FBCFFD90" box="[1031,1140,609,636]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
presumably occurs in talus-associated microhabitats.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3661CE37042BF27FF4DFD7BFBC9FA50" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="8BC34F687042BF27FF4DFD7BFDBDFD0F" blockId="9.[189,518,714,741]" box="[189,518,714,741]" pageId="9" pageNumber="10">
|
||
<heading id="D08BF8047042BF27FF4DFD7BFDBDFD0F" bold="true" box="[189,518,714,741]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B908937A7042BF27FF4DFD7BFDBDFD0F" bold="true" box="[189,518,714,741]" pageId="9" pageNumber="10">Notes on the type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC34F687042BF27FF4DFD4BFBC9FA50" blockId="9.[189,1399,763,1467]" pageId="9" pageNumber="10">
|
||
<bibRefCitation id="EFED32997042BF27FF4DFD4BFEC0FCFD" author="Smetana A." box="[189,379,763,790]" journalOrPublisher="Elytra (Tokyo)" pageId="9" pageNumber="10" pagination="77 - 88" part="23" refId="ref6582" refString="Smetana A. 1995. Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Elytra (Tokyo) 23: 77 - 88." title="Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini)" type="journal article" year="1995">Smetana (1995)</bibRefCitation>
|
||
based his description of
|
||
<taxonomicName id="4C7C34EB7042BF27FD5CFD4CFCA7FCFE" authorityName="Smetana" authorityYear="1995" box="[684,796,763,790]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FD5CFD4CFCA7FCFE" box="[684,796,763,790]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
on the
|
||
<typeStatus id="54C7F1CA7042BF27FC8BFD4BFC59FCFE" box="[891,994,763,789]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
(male, see above for details) and one
|
||
<typeStatus id="54C7F1CA7042BF27FF01FC91FEEDFCD0" box="[241,342,801,827]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
|
||
(female, for information see the undetermined or ambiguous material at the end of this paper), both from the north-eastern part of Terskei Alatau. The precise geographic origin of the
|
||
<typeStatus id="54C7F1CA7042BF27FAE0FCF6FACCFC8B" box="[1296,1399,838,864]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
is unknown, while the
|
||
<typeStatus id="54C7F1CA7042BF27FE3FFCDDFD8FFC6C" box="[463,564,877,903]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
|
||
comes from a locality called Jety Oguz. The aedeagus preparation of the
|
||
<typeStatus id="54C7F1CA7042BF27FF19FC22FEEBFC47" box="[233,336,914,940]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7C34EB7042BF27FE89FC22FE5CFC47" authorityName="Smetana" authorityYear="1995" box="[377,487,914,940]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FE89FC22FE5CFC47" box="[377,487,914,940]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F36B0FF87042BF27FDFFFC22FDDCFC47" box="[527,615,914,940]" name="Canada" pageId="9" pageNumber="10">Canada</collectingCountry>
|
||
balsam allows an examination of its lateral side, not illustrated in
|
||
<bibRefCitation id="EFED32997042BF27FF4DFC07FEC2FC39" author="Smetana A." box="[189,377,951,978]" journalOrPublisher="Elytra (Tokyo)" pageId="9" pageNumber="10" pagination="77 - 88" part="23" refId="ref6582" refString="Smetana A. 1995. Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Elytra (Tokyo) 23: 77 - 88." title="Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini)" type="journal article" year="1995">Smetana (1995)</bibRefCitation>
|
||
(
|
||
<figureCitation id="134753ED7042BF27FE7AFC07FE5DFC3A" box="[394,486,951,977]" captionStart="Fig.3" captionStartId="10.[189,232,1964,1990]" captionTargetBox="[331,1257,336,1918]" captionTargetId="figure@10.[331,1257,334,1918]" captionTargetPageId="10" captionText="Fig.3.HolotypeofQuediuskalabiSmetana,1995.A.Habitus.B.Permanentpreparationoftheaedeagus. Scale bars: 1 mm." httpUri="https://zenodo.org/record/1167587/files/figure.png" pageId="9" pageNumber="10">Fig. 3B</figureCitation>
|
||
). It confirms our identification of
|
||
<taxonomicName id="4C7C34EB7042BF27FC8AFC08FC5CFC3A" authorityName="Smetana" authorityYear="1995" box="[890,999,951,978]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FC8AFC08FC5CFC3A" box="[890,999,951,978]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
, which differs from other species of the
|
||
<taxonomicName id="4C7C34EB7042BF27FEFCFC6DFE24FC1C" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[268,415,989,1015]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7042BF27FEFCFC6DFE24FC1C" box="[268,415,989,1015]" italics="true" pageId="9" pageNumber="10">Q. mutilatus</emphasis>
|
||
</taxonomicName>
|
||
group by its narrower and somewhat curved apical portion of the median lobe of the aedeagus, with a relatively short blade of its subapical tooth (aedeagus in lateral view). In the shape of the apical portion of the paramere and degree of its incision,
|
||
<taxonomicName id="4C7C34EB7042BF27FC5CFB98FBA2FBA9" authorityName="Smetana" authorityYear="1995" box="[940,1049,1064,1090]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FC5CFB98FBA2FBA9" box="[940,1049,1064,1090]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
displays a transition between
|
||
<taxonomicName id="4C7C34EB7042BF27FF4DFBFEFEF7FB83" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[189,332,1102,1128]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7042BF27FF4DFBFEFEF7FB83" box="[189,332,1102,1128]" italics="true" pageId="9" pageNumber="10">Q. mutilatus</emphasis>
|
||
</taxonomicName>
|
||
with a less incised paramere bearing more lateral peg setae, and
|
||
<taxonomicName id="4C7C34EB7042BF27FBDBFBFEFB2AFB8C" authorityName="Smetana" authorityYear="2014" box="[1067,1169,1102,1128]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="equus">
|
||
<emphasis id="B908937A7042BF27FBDBFBFEFB2AFB8C" box="[1067,1169,1102,1128]" italics="true" pageId="9" pageNumber="10">Q. equus</emphasis>
|
||
</taxonomicName>
|
||
with a more incised paramere bearing fewer lateral peg setae. The characters indicated by
|
||
<bibRefCitation id="EFED32997042BF27FC07FBC3FB09FB66" author="Smetana A." box="[1015,1202,1139,1165]" journalOrPublisher="Elytra (Tokyo)" pageId="9" pageNumber="10" pagination="115 - 128" part="26" refId="ref6615" refString="Smetana A. 1998. Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Part 2. Elytra (Tokyo) 26 (1): 115 - 128." title="Taxonomic and faunistic contributions to the knowledge of Palearctic Quediina (Coleoptera, Staphylinidae, Staphylinini). Part 2" type="journal article" year="1998">Smetana (1988)</bibRefCitation>
|
||
in a comparison of
|
||
<taxonomicName id="4C7C34EB7042BF27FF2EFB29FEF7FB58" authorityName="Smetana" authorityYear="1995" box="[222,332,1177,1203]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FF2EFB29FEF7FB58" box="[222,332,1177,1203]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C7C34EB7042BF27FE62FB29FD9FFB58" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[402,548,1177,1203]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7042BF27FE62FB29FD9FFB58" box="[402,548,1177,1203]" italics="true" pageId="9" pageNumber="10">Q. mutilatus</emphasis>
|
||
</taxonomicName>
|
||
were based, as revealed here, on non-conspecific males and females of both species. They do not align with the more extensive material examined here. The collecting locality of the
|
||
<typeStatus id="54C7F1CA7042BF27FEF8FB54FED6FB15" box="[264,365,1252,1278]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7C34EB7042BF27FE64FB54FDBAFB15" authorityName="Smetana" authorityYear="1995" box="[404,513,1252,1278]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FE64FB54FDBAFB15" box="[404,513,1252,1278]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
is, in fact, located closer to the distribution area of
|
||
<taxonomicName id="4C7C34EB7042BF27FBACFB54FB56FB15" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[1116,1261,1252,1278]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7042BF27FBACFB54FB56FB15" box="[1116,1261,1252,1278]" italics="true" pageId="9" pageNumber="10">Q. mutilatus</emphasis>
|
||
</taxonomicName>
|
||
. Except for the
|
||
<typeStatus id="54C7F1CA7042BF27FF17FAB9FEEAFAC8" box="[231,337,1289,1315]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
, the distributional records of
|
||
<taxonomicName id="4C7C34EB7042BF27FD51FABAFCB7FAC8" authorityName="Smetana" authorityYear="1995" box="[673,780,1289,1316]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FD51FABAFCB7FAC8" box="[673,780,1289,1316]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
summarized in
|
||
<bibRefCitation id="EFED32997042BF27FC34FAB9FA8CFACF" author="Solodovnikov A. & Hansen A. K." box="[964,1335,1289,1316]" journalOrPublisher="Zootaxa" pageId="9" pageNumber="10" pagination="475 - 490" part="4170" publicationUrl="https://doi.org/10.11646/zootaxa.4170.3.3" refId="ref6763" refString="Solodovnikov A. & Hansen A. K. 2016. Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini). Zootaxa 4170 (3): 475 - 490. https: // doi. org / 10.11646 / zootaxa. 4170.3.3" title="Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini)" type="journal article" year="2016">Solodovnikov & Hansen (2016)</bibRefCitation>
|
||
were based on misidentified specimens of
|
||
<taxonomicName id="4C7C34EB7042BF27FD86FA9FFCB3FAA2" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[630,776,1327,1353]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7042BF27FD86FA9FFCB3FAA2" box="[630,776,1327,1353]" italics="true" pageId="9" pageNumber="10">Q. mutilatus</emphasis>
|
||
</taxonomicName>
|
||
. The specimens of
|
||
<taxonomicName id="4C7C34EB7042BF27FC04FA9FFBD9FAA2" authorityName="Smetana" authorityYear="1995" box="[1012,1122,1327,1353]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FC04FA9FFBD9FAA2" box="[1012,1122,1327,1353]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
from Teploklyuchenka Village, examined here and compared with the
|
||
<typeStatus id="54C7F1CA7042BF27FD00FAE4FCE1FA85" box="[752,858,1364,1390]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
, are the first precisely georeferenced material for this species. Based on the shape of aedeagus, three possibly mislabeled specimens from Toksanbai in Dzungarian Alatau seem to belong to
|
||
<taxonomicName id="4C7C34EB7042BF27FD66FA10FCB8FA51" authorityName="Smetana" authorityYear="1995" box="[662,771,1440,1466]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7042BF27FD66FA10FCB8FA51" box="[662,771,1440,1466]" italics="true" pageId="9" pageNumber="10">Q. kalabi</emphasis>
|
||
</taxonomicName>
|
||
as well (for details see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |