328 lines
37 KiB
XML
328 lines
37 KiB
XML
<document id="5D6528E7DF9A635DD3762B095F88013B" ID-CLB-Dataset="30822" ID-DOI="10.5852/ejt.2018.401" ID-GBIF-Dataset="49d45069-6150-4b6e-873c-fc8058025a39" ID-Zenodo-Dep="1167581" ID-ZooBank="9C9DB157-AAA5-40B7-BA0B-9A57779382C1" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517911480053" checkinUser="plazi" docAuthor="Salnitska, Maria & Solodovnikov, Alexey" docDate="2018" docId="03D5FE7E7042BF22FD95FA4AFACCFCBA" docLanguage="en" docName="ejt-401_salnitska_solodovnikov.pdf.imf" docOrigin="European Journal of Taxonomy 401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Quedius equus Smetana 2014" docType="treatment" docVersion="15" lastPageNumber="13" masterDocId="FFEC8606704BBF2EFFF0FFB0FFBBFFEB" masterDocTitle="Taxonomy of the poorly known Quedius mutilatus group of wingless montane species from Middle Asia (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="17" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547222236327" originalUpdateUser="yanell" pageNumber="10" updateTime="1698519557005" updateUser="ExternalLinkService">
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<mods:title id="4D7161BF869FC809128280AEBC157C8E">Taxonomy of the poorly known Quedius mutilatus group of wingless montane species from Middle Asia (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
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<mods:namePart id="9066DCADB4307C13ED79677376A29653">Salnitska, Maria</mods:namePart>
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<mods:namePart id="4B1C7168D05B658F85B90343BF5AC25A">Solodovnikov, Alexey</mods:namePart>
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<mods:typeOfResource id="BC63DE4654CEDF6F57A1754AC3EDDE9A">text</mods:typeOfResource>
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<mods:title id="3CA79E68620CC11CD3245A8860E1ADA1">European Journal of Taxonomy</mods:title>
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<mods:part id="8CF01EE9A9E5FB04019BBD2FD2DC7C21">
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<mods:date id="562E4CD1341BE90F17791F9B91FF5260">2018</mods:date>
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<mods:number id="9312EA0F8195A031EAA97EFE07493D6A">2018-02-06</mods:number>
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<treatment id="03D5FE7E7042BF22FD95FA4AFACCFCBA" ID-DOI="http://doi.org/10.5281/zenodo.5695868" ID-GBIF-Taxon="141125769" ID-Zenodo-Dep="5695868" LSID="urn:lsid:plazi:treatment:03D5FE7E7042BF22FD95FA4AFACCFCBA" httpUri="http://treatment.plazi.org/id/03D5FE7E7042BF22FD95FA4AFACCFCBA" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
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<subSubSection id="C3661CE37042BF27FD95FA4AFC74F9FF" box="[613,975,1530,1557]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BC34F687042BF27FD95FA4AFC74F9FF" blockId="9.[613,975,1530,1594]" box="[613,975,1530,1557]" pageId="9" pageNumber="10">
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<heading id="D08BF8047042BF27FD95FA4AFC74F9FF" box="[613,975,1530,1557]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
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<taxonomicName id="4C7C34EB7042BF27FD95FA4AFC74F9FF" ID-CoL="7929M" authority="Smetana, 2014" authorityName="Smetana" authorityYear="2014" box="[613,975,1530,1557]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="equus">
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<emphasis id="B908937A7042BF27FD95FA4AFCA2F9FE" bold="true" box="[613,793,1530,1557]" italics="true" pageId="9" pageNumber="10">Quedius equus</emphasis>
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<bibRefCitation id="EFED32997042BF27FCD0FA4AFC74F9FF" author="Smetana A." box="[800,975,1530,1556]" journalOrPublisher="Folia Heyrovskyana, Series A" pageId="9" pageNumber="10" pagination="1 - 39" part="21" refId="ref6707" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, Series A 21 (1 - 4): 1 - 39." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22" type="journal article" year="2014">Smetana, 2014</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3661CE37042BF27FD3AF990FCD2F9D1" box="[714,873,1568,1594]" pageId="9" pageNumber="10" type="description">
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<paragraph id="8BC34F687042BF27FD3AF990FCD2F9D1" blockId="9.[613,975,1530,1594]" box="[714,873,1568,1594]" pageId="9" pageNumber="10">
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<figureCitation id="134753ED7042BF27FD3AF990FCA8F9D1" box="[714,787,1568,1594]" captionStart="Fig.1" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[221,1375,624,1881]" captionTargetId="figure@4.[208,1379,622,1889]" captionTargetPageId="4" captionText="Fig.1.DistributionandaedeagusvariabilityoftheQuediusmutilatusgroupintheTien-ShanMountains. Numbers are specified in Table 1. Empty symbols = typespcimens (for Q. kalabi precise locality of the holotype unknown);? = ambiguously labelled or undetermined material. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1167583/files/figure.png" pageId="9" pageNumber="10">Figs 1</figureCitation>
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,
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<figureCitation id="134753ED7042BF27FCD1F990FCD2F9D1" box="[801,873,1568,1594]" captionStart="Fig. 4" captionStartId="11.[189,232,1858,1884]" captionTargetBox="[333,1256,265,1830]" captionTargetId="figure@11.[331,1256,265,1830]" captionTargetPageId="11" captionText="Fig. 4. Aedeagi in the Quedius mutilatus group. A–B. Quedius mutilatus Eppelsheim, 1888. C–D. Quedius kalabi Smetana, 1995. E–F. Quedius kungeicus sp. nov. G–N. Quedius equus Smetana, 2014 (Karkara Valley, Kazakhstan), variability of the structure of the aedeagus.A, C, E, G, I, K, M = median lobe, laterally. B, D, F, H, J, L, N = paramere (dissected from median lobe), underside (side facing median lobe). Scale bars: 1 mm." httpUri="https://zenodo.org/record/1167589/files/figure.png" pageId="9" pageNumber="10">4G–N</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3661CE37042BF27FF4DF9D4FC0FF929" pageId="9" pageNumber="10" type="reference_group">
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<paragraph id="8BC34F687042BF27FF4DF9D4FCEEF994" blockId="9.[189,853,1635,1663]" box="[189,853,1635,1663]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="AB6C68467042BF27FF4DF9D4FCEEF994" box="[189,853,1635,1663]" pageId="9" pageNumber="10">
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<taxonomicName id="4C7C34EB7042BF27FF4DF9D4FDF6F995" ID-CoL="7929M" authority="Smetana, 2014: 35" authorityName="Smetana" authorityPageNumber="35" authorityYear="2014" box="[189,589,1635,1662]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="equus">
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<emphasis id="B908937A7042BF27FF4DF9D4FED1F996" box="[189,362,1636,1662]" italics="true" pageId="9" pageNumber="10">Quedius equus</emphasis>
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<treatmentCitation id="0ADD69797042BF27FE82F9D4FDF6F995" author="Smetana" box="[370,589,1635,1662]" page="35" pageId="9" pageNumber="10" year="2014">
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<bibRefCitation id="EFED32997042BF27FE82F9D4FD9EF995" author="Smetana A." box="[370,549,1636,1662]" journalOrPublisher="Folia Heyrovskyana, Series A" pageId="9" pageNumber="10" pagination="1 - 39" part="21" refId="ref6707" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, Series A 21 (1 - 4): 1 - 39." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22" type="journal article" year="2014">Smetana, 2014</bibRefCitation>
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: 35
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</treatmentCitation>
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</taxonomicName>
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(original description).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC34F687042BF27FF4DF917FC0FF929" blockId="9.[189,948,1703,1730]" box="[189,948,1703,1730]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="AB6C68467042BF27FF4DF917FC0FF929" box="[189,948,1703,1730]" pageId="9" pageNumber="10">
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<taxonomicName id="4C7C34EB7042BF27FF4DF917FED1F92A" ID-CoL="7929M" authorityName="Smetana" authorityYear="2014" box="[189,362,1703,1729]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="equus">
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<emphasis id="B908937A7042BF27FF4DF917FED1F92A" box="[189,362,1703,1729]" italics="true" pageId="9" pageNumber="10">Quedius equus</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0ADD69797042BF27FE78F917FCB1F929" author="Solodovnikov" box="[392,778,1703,1730]" page="8" pageId="9" pageNumber="10" year="2016">
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<bibRefCitation id="EFED32997042BF27FE78F917FD4BF92A" author="Solodovnikov A. & Hansen A. K." box="[392,752,1703,1730]" journalOrPublisher="Zootaxa" pageId="9" pageNumber="10" pagination="475 - 490" part="4170" publicationUrl="https://doi.org/10.11646/zootaxa.4170.3.3" refId="ref6763" refString="Solodovnikov A. & Hansen A. K. 2016. Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini). Zootaxa 4170 (3): 475 - 490. https: // doi. org / 10.11646 / zootaxa. 4170.3.3" title="Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini)" type="journal article" year="2016">Solodovnikov & Hansen 2016</bibRefCitation>
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: 8
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</treatmentCitation>
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(distribution).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3661CE37042BF22FF4DF95BFDDAFE55" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="8BC34F687042BF27FF4DF95BFC00F8EE" blockId="9.[189,955,1771,1798]" box="[189,955,1771,1798]" pageId="9" pageNumber="10">
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<emphasis id="B908937A7042BF27FF4DF95BFED7F8EE" bold="true" box="[189,364,1771,1797]" pageId="9" pageNumber="10">
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<typeStatus id="54C7F1CA7042BF27FF4DF95BFF41F8EE" box="[189,250,1771,1797]" pageId="9" pageNumber="10">Type</typeStatus>
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material
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</emphasis>
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(not examined, listed according to
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<bibRefCitation id="EFED32997042BF27FCFAF95BFC08F8EE" author="Smetana A." box="[778,947,1771,1797]" journalOrPublisher="Folia Heyrovskyana, Series A" pageId="9" pageNumber="10" pagination="1 - 39" part="21" refId="ref6707" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, Series A 21 (1 - 4): 1 - 39." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22" type="journal article" year="2014">Smetana 2014</bibRefCitation>
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)
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</paragraph>
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<paragraph id="8BC34F687042BF27FF1AF8ACFEE7F8DD" blockId="9.[189,1399,1820,1922]" box="[234,348,1820,1846]" pageId="9" pageNumber="10">
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<emphasis id="B908937A7042BF27FF1AF8ACFEE7F8DD" bold="true" box="[234,348,1820,1846]" pageId="9" pageNumber="10">Holotype</emphasis>
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</paragraph>
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<paragraph id="8BC34F687042BF27FF4DF8F2FBDEF86A" blockId="9.[189,1399,1820,1922]" pageId="9" pageNumber="10">
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<materialsCitation id="3B1445357042BF27FF4DF8F2FBDEF86A" ID-GBIF-Occurrence="1821772571" collectedFrom="alp. meadows, screes" collectingDate="2009-05-31" collectingDateMax="2009-06-02" collectingDateMin="2009-05-31" collectionCode="NMW" collectorName="J. Kalab" country="China" elevation="3500" latitude="42.016666" location="NW-Xinjiang" longLatPrecision="1152" longitude="81.25" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Xinjiang" typeStatus="holotype">
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<collectingCountry id="F36B0FF87042BF27FF4DF8F2FEA4F8B7" box="[189,287,1858,1884]" name="China" pageId="9" pageNumber="10">CHINA</collectingCountry>
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:
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<specimenCount id="9D7A84E17042BF27FEDFF8F2FEFDF8B7" box="[303,326,1858,1884]" pageId="9" pageNumber="10" type="male">♂</specimenCount>
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, “CHINA:
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<collectingRegion id="49B8818A7042BF27FE27F8F2FDCCF8B7" box="[471,631,1858,1884]" pageId="9" pageNumber="10">NW-Xinjiang</collectingRegion>
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<locationDeviation id="9A71564A7042BF27FD73F8F2FB4BF8B7" box="[643,1264,1858,1885]" pageId="9" pageNumber="10">
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ca
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||
<quantity id="4C84E28D7042BF27FD5AF8F2FD42F8B7" box="[682,761,1858,1885]" metricMagnitude="4" metricUnit="m" metricValue="5.8" pageId="9" pageNumber="10" unit="km" value="58.0">58 km</quantity>
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WNW Baicheng right trib. of Terek riv
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</locationDeviation>
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.
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<geoCoordinate id="EE4829AF7042BF27FAF1F8F0FAC9F8B7" box="[1281,1394,1856,1884]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="42.016666">42°01′ N</geoCoordinate>
|
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,
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||
<geoCoordinate id="EE4829AF7042BF27FF4DF8D5FE98F869" box="[189,291,1893,1922]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="81.25">81°15′ E</geoCoordinate>
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/
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<collectingDate id="EF8690407042BF27FEC3F8D7FE4AF86A" box="[307,497,1895,1922]" pageId="9" pageNumber="10">
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||
31.5. –
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||
<date id="FFC269A87042BF27FE7BF8D7FE4AF86A" box="[395,497,1895,1921]" pageId="9" pageNumber="10" value="2009-06-02">2.6.2009</date>
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</collectingDate>
|
||
<quantity id="4C84E28D7042BF27FE09F8D7FDE9F869" box="[505,594,1895,1922]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="m" value="3500.0">
|
||
<elevation id="0051A85B7042BF27FE09F8D7FDE9F869" box="[505,594,1895,1922]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="m" value="3500.0">3500 m</elevation>
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||
</quantity>
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||
<collectedFrom id="38600C957042BF27FDA9F8D7FCEAF869" box="[601,849,1895,1922]" pageId="9" pageNumber="10">alp. meadows, screes</collectedFrom>
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leg.
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||
<collectorName id="26892ABE7042BF27FC7AF8D8FC56F86A" box="[906,1005,1895,1922]" pageId="9" pageNumber="10">J. Kalab</collectorName>
|
||
” (
|
||
<collectionCode id="ED6DD7AD7042BF27FBF7F8D7FBE2F86A" box="[1031,1113,1895,1921]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="10">NMW</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC34F687042BF27FF1AF81BFEDCF82E" blockId="9.[189,1002,1963,2027]" box="[234,359,1963,1989]" pageId="9" pageNumber="10">
|
||
<emphasis id="B908937A7042BF27FF1AF81BFEDCF82E" bold="true" box="[234,359,1963,1989]" pageId="9" pageNumber="10">Paratypes</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC34F687042BF27FF4DF861FC51F800" blockId="9.[189,1002,1963,2027]" box="[189,1002,2001,2027]" pageId="9" pageNumber="10">
|
||
<materialsCitation id="3B1445357042BF27FF4DF861FC51F800" ID-GBIF-Occurrence="1821772572" box="[189,1002,2001,2027]" collectedFrom="alp. meadows, screes" collectingDate="2009-05-31" collectingDateMax="2009-06-02" collectingDateMin="2009-05-31" collectionCode="NMW" collectorName="J. Kalab" country="China" elevation="3500" latitude="42.016666" location="NW-Xinjiang" longLatPrecision="1152" longitude="81.25" pageId="9" pageNumber="10" specimenCount="2" specimenCount-male="2" stateProvince="Xinjiang" typeStatus="paratype">
|
||
<collectingCountry id="F36B0FF87042BF27FF4DF861FEA4F800" box="[189,287,2001,2027]" name="China" pageId="9" pageNumber="10">CHINA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D7A84E17042BF27FED9F861FED6F800" box="[297,365,2001,2027]" pageId="9" pageNumber="10" type="male">2 ♂♂</specimenCount>
|
||
, same collecting data as for holotype (
|
||
<bibRefCitation id="EFED32997042BF27FCC0F861FC65F800" author="Smetana A." box="[816,990,2001,2027]" journalOrPublisher="Folia Heyrovskyana, Series A" pageId="9" pageNumber="10" pagination="1 - 39" part="21" refId="ref6707" refString="Smetana A. 2014. Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, Series A 21 (1 - 4): 1 - 39." title="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22" type="journal article" year="2014">Smetana 2014</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF031FE07041BF24FF4DF81CFE2BF801" httpUri="https://zenodo.org/record/1167587/files/figure.png" pageId="10" pageNumber="11" targetBox="[331,1257,336,1918]" targetPageId="10">
|
||
<paragraph id="8BC34F687041BF24FF4DF81CFE2BF801" blockId="10.[189,1399,1964,2026]" pageId="10" pageNumber="11">
|
||
<emphasis id="B908937A7041BF24FF4DF81CFEB3F82D" bold="true" box="[189,264,1964,1990]" pageId="10" pageNumber="11">Fig. 3.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C7C34EB7041BF24FE6CF81CFCBAF82C" authority="Smetana, 1995" authorityName="Smetana" authorityYear="1995" box="[412,769,1964,1991]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7041BF24FE6CF81CFDF7F82D" box="[412,588,1964,1990]" italics="true" pageId="10" pageNumber="11">Quedius kalabi</emphasis>
|
||
Smetana, 1995
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B908937A7041BF24FCF8F81CFCA6F82D" bold="true" box="[776,797,1964,1990]" pageId="10" pageNumber="11">A</emphasis>
|
||
. Habitus.
|
||
<emphasis id="B908937A7041BF24FC61F81DFC1EF82C" bold="true" box="[913,933,1965,1991]" pageId="10" pageNumber="11">B</emphasis>
|
||
. Permanent preparation of the aedeagus. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF031FE07040BF25FF4DF8F2FD32F802" httpUri="https://zenodo.org/record/1167589/files/figure.png" pageId="11" pageNumber="12" targetBox="[333,1256,265,1830]" targetPageId="11">
|
||
<paragraph id="8BC34F687040BF25FF4DF8F2FD32F802" blockId="11.[189,1399,1858,2026]" pageId="11" pageNumber="12">
|
||
<emphasis id="B908937A7040BF25FF4DF8F2FEAEF8B7" bold="true" box="[189,277,1858,1884]" pageId="11" pageNumber="12">Fig. 4.</emphasis>
|
||
Aedeagi in the
|
||
<taxonomicName id="4C7C34EB7040BF25FE08F8F2FD60F8B7" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[504,731,1858,1884]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7040BF25FE08F8F2FD60F8B7" box="[504,731,1858,1884]" italics="true" pageId="11" pageNumber="12">Quedius mutilatus</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
<emphasis id="B908937A7040BF25FCBAF8F2FC39F8B7" bold="true" box="[842,898,1858,1884]" pageId="11" pageNumber="12">A–B</emphasis>
|
||
.
|
||
<taxonomicName id="4C7C34EB7040BF25FC6BF8F2FAC8F8B7" authority="Eppelsheim, 1888" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[923,1395,1858,1884]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7040BF25FC6BF8F2FBC5F8B7" box="[923,1150,1858,1884]" italics="true" pageId="11" pageNumber="12">Quedius mutilatus</emphasis>
|
||
Eppelsheim, 1888
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B908937A7040BF25FF4DF8D5FF4DF894" bold="true" box="[189,246,1893,1919]" pageId="11" pageNumber="12">C–D</emphasis>
|
||
.
|
||
<taxonomicName id="4C7C34EB7040BF25FEF6F8D5FDC8F86B" authority="Smetana, 1995" authorityName="Smetana" authorityYear="1995" box="[262,627,1893,1920]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kalabi">
|
||
<emphasis id="B908937A7040BF25FEF6F8D5FE02F894" box="[262,441,1893,1919]" italics="true" pageId="11" pageNumber="12">Quedius kalabi</emphasis>
|
||
Smetana, 1995
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B908937A7040BF25FD8FF8D5FD08F894" bold="true" box="[639,691,1893,1919]" pageId="11" pageNumber="12">E–F</emphasis>
|
||
.
|
||
<taxonomicName id="4C7C34EB7040BF25FD32F8D5FC1AF894" authority="Salnitska & Solodovnikov, 2018" authorityName="Salnitska & Solodovnikov" authorityYear="2018" box="[706,929,1893,1919]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kungeicus" status="sp. nov.">
|
||
<emphasis id="B908937A7040BF25FD32F8D5FC1AF894" box="[706,929,1893,1919]" italics="true" pageId="11" pageNumber="12">Quedius kungeicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23B2E017040BF25FC59F8D6FBBFF86B" box="[937,1028,1894,1920]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B908937A7040BF25FBFBF8D5FBFEF894" bold="true" box="[1035,1093,1893,1919]" pageId="11" pageNumber="12">G–N</emphasis>
|
||
.
|
||
<taxonomicName id="4C7C34EB7040BF25FBA5F8D5FF43F849" authority="Smetana, 2014" authorityName="Smetana" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="equus">
|
||
<emphasis id="B908937A7040BF25FBA5F8D5FAB9F894" box="[1109,1282,1893,1919]" italics="true" pageId="11" pageNumber="12">Quedius equus</emphasis>
|
||
Smetana, 2014
|
||
</taxonomicName>
|
||
(Karkara Valley, Kazakhstan), variability of the structure of the aedeagus. A, C, E, G, I, K, M = median lobe, laterally. B, D, F, H, J, L, N = paramere (dissected from median lobe), underside (side facing median lobe). Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC34F687047BF22FF4DFEBBFE11FECE" blockId="12.[189,426,267,293]" box="[189,426,267,293]" pageId="12" pageNumber="13">
|
||
<heading id="D08BF8047047BF22FF4DFEBBFE11FECE" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="B908937A7047BF22FF4DFEBBFE11FECE" bold="true" box="[189,426,267,293]" pageId="12" pageNumber="13">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC34F687047BF22FF4DFE8AFDDAFE55" blockId="12.[189,1399,312,446]" pageId="12" pageNumber="13">
|
||
<materialsCitation id="3B1445357047BF22FF4DFE8AFE0FFE9C" ID-GBIF-Occurrence="1821772566" collectingDate="1989-07-18" collectingDateMax="1993-07-05" collectingDateMin="1984-07-22" collectionCode="ZIN" collectorName="I. Kabak" country="Kazakhstan" elevation="2250" latitude="42.618" location="Karkara Valley, Baiankol Valley" longLatPrecision="68" longitude="79.892" pageId="12" pageNumber="13" specimenCount="6" specimenCount-female="2" specimenCount-male="4">
|
||
<collectingCountry id="F36B0FF87047BF22FF4DFE8AFE33FEBF" box="[189,392,314,340]" name="Kazakhstan" pageId="12" pageNumber="13">KAZAKHSTAN</collectingCountry>
|
||
:
|
||
<specimenCount id="9D7A84E17047BF22FE62FE8AFE6EFEBF" box="[402,469,314,340]" pageId="12" pageNumber="13" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D7A84E17047BF22FE10FE8AFD99FEBF" box="[480,546,314,340]" pageId="12" pageNumber="13" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<location id="8EA319B37047BF22FDDFFE8AFD64FEBF" LSID="urn:lsid:plazi:treatment:03D5FE7E7042BF22FD95FA4AFACCFCBA:8EA319B37047BF22FDDFFE8AFD64FEBF" box="[559,735,314,340]" country="Kazakhstan" latitude="42.618" longLatPrecision="68" longitude="79.892" municipality="Baiankol Valley" name="Karkara Valley" pageId="12" pageNumber="13">Karkara Valley</location>
|
||
,
|
||
<quantity id="4C84E28D7047BF22FD1AFE89FC30FEBF" box="[746,907,313,340]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="12" pageNumber="13" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">
|
||
<elevation id="0051A85B7047BF22FD1AFE89FC30FEBF" box="[746,907,313,340]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="12" pageNumber="13" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">2000–2500 m</elevation>
|
||
</quantity>
|
||
[
|
||
<geoCoordinate id="EE4829AF7047BF22FC68FE88FB8BFEBF" box="[920,1072,312,340]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="42.655556">42°39′20″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4829AF7047BF22FBCBFE88FB76FEBF" box="[1083,1229,312,340]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="79.225">79°13′30″ E</geoCoordinate>
|
||
],
|
||
<date id="FFC269A87047BF22FB11FE8AFACAFEBF" box="[1249,1393,314,340]" pageId="12" pageNumber="13" value="1989-07-18">
|
||
<collectingDate id="EF8690407047BF22FB11FE8AFACAFEBF" box="[1249,1393,314,340]" pageId="12" pageNumber="13" value="1989-07-18">18 Jul. 1989</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26892ABE7047BF22FF4DFEEEFE9DFE9C" box="[189,294,349,376]" pageId="12" pageNumber="13">I. Kabak</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED6DD7AD7047BF22FE80FEEDFE1CFE9C" box="[368,423,349,375]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="13">ZIN</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1445357047BF22FE4FFEEDFE10FE70" ID-GBIF-Occurrence="1821772570" collectingDate="1993-07-05" collectingDateMax="1993-07-05" collectingDateMin="1984-07-22" collectionCode="ZIN" collectorName="I. Kabak" country="Kazakhstan" elevation="2750" latitude="42.618" location="Baiankol Valley" longLatPrecision="68" longitude="79.892" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="9D7A84E17047BF22FE4FFEEDFE4BFE93" box="[447,496,349,376]" pageId="12" pageNumber="13" type="male">1 ♂</specimenCount>
|
||
,
|
||
<location id="8EA319B37047BF22FDF2FEEDFD79FE9C" LSID="urn:lsid:plazi:treatment:03D5FE7E7042BF22FD95FA4AFACCFCBA:8EA319B37047BF22FDF2FEEDFD79FE9C" box="[514,706,349,375]" country="Kazakhstan" latitude="42.618" longLatPrecision="68" longitude="79.892" municipality="Baiankol Valley" name="Baiankol Valley" pageId="12" pageNumber="13">Baiankol Valley</location>
|
||
,
|
||
<quantity id="4C84E28D7047BF22FD22FEEDFCC3FE93" box="[722,888,349,376]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="12" pageNumber="13" unit="m" value="2750.0" valueMax="3000.0" valueMin="2500.0">
|
||
<elevation id="0051A85B7047BF22FD22FEEDFCC3FE93" box="[722,888,349,376]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="12" pageNumber="13" unit="m" value="2750.0" valueMax="3000.0" valueMin="2500.0">2500–3000 m</elevation>
|
||
</quantity>
|
||
[
|
||
<geoCoordinate id="EE4829AF7047BF22FC7AFEEBFB9DFE93" box="[906,1062,347,376]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="42.595833">42°35′45″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4829AF7047BF22FBC6FEEBFB75FE9C" box="[1078,1230,347,375]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="79.9675">79°58′03″ E</geoCoordinate>
|
||
],
|
||
<date id="FFC269A87047BF22FB16FEEDFACAFE9C" box="[1254,1393,349,376]" pageId="12" pageNumber="13" value="1993-07-05">
|
||
<collectingDate id="EF8690407047BF22FB16FEEDFACAFE9C" box="[1254,1393,349,376]" pageId="12" pageNumber="13" value="1993-07-05">5 Jul. 1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26892ABE7047BF22FF4DFE31FE98FE71" box="[189,291,384,411]" pageId="12" pageNumber="13">I. Kabak</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED6DD7AD7047BF22FE97FE31FE25FE70" box="[359,414,385,411]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="13">ZIN</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1445357047BF22FE43FE31FDDAFE55" ID-GBIF-Occurrence="1821772568" collectedFrom="alpine meadows" collectingDate="1984-07-22" collectingDateMax="1993-07-05" collectingDateMin="1984-07-22" collectionCode="ZIN" collectorName="B. Iskakov" country="Kazakhstan" elevation="3400" latitude="42.618" longLatPrecision="68" longitude="79.892" municipality="Baiankol Valley" pageId="12" pageNumber="13" specimenCount="2" specimenCount-male="2">
|
||
<specimenCount id="9D7A84E17047BF22FE43FE31FE43FE70" box="[435,504,385,411]" pageId="12" pageNumber="13" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<locationDeviation id="9A71564A7047BF22FDF6FE31FCA4FE71" box="[518,799,384,411]" pageId="12" pageNumber="13">
|
||
<quantity id="4C84E28D7047BF22FDF6FE31FDEBFE71" box="[518,592,384,411]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="13" unit="km" value="28.0">28 km</quantity>
|
||
SW of Narynkol
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4C84E28D7047BF22FCDCFE31FC3CFE70" box="[812,903,385,411]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="13" unit="m" value="3400.0">
|
||
<elevation id="0051A85B7047BF22FCDCFE31FC3CFE70" box="[812,903,385,411]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="13" unit="m" value="3400.0">3400 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="38600C957047BF22FC65FE30FBEFFE71" box="[917,1108,384,410]" pageId="12" pageNumber="13">alpine meadows</collectedFrom>
|
||
[
|
||
<geoCoordinate id="EE4829AF7047BF22FB92FE30FB59FE70" box="[1122,1250,384,411]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="55" value="42.618">42.618° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4829AF7047BF22FB1FFE31FAD2FE70" box="[1263,1385,384,411]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="55" value="79.892">79.892° E</geoCoordinate>
|
||
],
|
||
<date id="FFC269A87047BF22FF4DFE14FEEBFE55" box="[189,336,420,446]" pageId="12" pageNumber="13" value="1984-07-22">
|
||
<collectingDate id="EF8690407047BF22FF4DFE14FEEBFE55" box="[189,336,420,446]" pageId="12" pageNumber="13" value="1984-07-22">22 Jul. 1984</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26892ABE7047BF22FEACFE14FE60FE55" box="[348,475,420,446]" pageId="12" pageNumber="13">B. Iskakov</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED6DD7AD7047BF22FDEDFE14FDEFFE55" box="[541,596,420,446]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="13">ZIN</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3661CE37047BF22FF4DFE55FAC9FD57" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="8BC34F687047BF22FF4DFE55FDA8FE14" blockId="12.[189,531,485,512]" box="[189,531,485,512]" pageId="12" pageNumber="13">
|
||
<heading id="D08BF8047047BF22FF4DFE55FDA8FE14" bold="true" box="[189,531,485,512]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="B908937A7047BF22FF4DFE55FDA8FE14" bold="true" box="[189,531,485,512]" pageId="12" pageNumber="13">Distribution and bionomics</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC34F687047BF22FF4DFDA4FAC9FD57" blockId="12.[189,1399,532,701]" pageId="12" pageNumber="13">
|
||
Previously,
|
||
<emphasis id="B908937A7047BF22FEB6FDA5FED9FDC4" box="[326,354,533,559]" italics="true" pageId="12" pageNumber="13">Q.</emphasis>
|
||
equus was known from the
|
||
<typeStatus id="54C7F1CA7047BF22FD53FDA4FCB1FDC5" box="[675,778,532,558]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
and two
|
||
<typeStatus id="54C7F1CA7047BF22FC81FDA5FC5AFDC4" box="[881,993,533,559]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
|
||
(all males) only, collected together at the
|
||
<typeStatus id="54C7F1CA7047BF22FEF7FD88FE81FDB9" box="[263,314,568,594]" pageId="12" pageNumber="13">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="49B8818A7047BF22FE31FD88FD93FDB9" box="[449,552,568,594]" country="China" name="Xinjiang Uygur" pageId="12" pageNumber="13">Xinjiang</collectingRegion>
|
||
Province in
|
||
<collectingCountry id="F36B0FF87047BF22FD30FD88FCB1FDB9" box="[704,778,568,594]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
|
||
. New material examined here matches well with the original description and significantly expands the distribution of this species westwards (
|
||
<figureCitation id="134753ED7047BF22FB1FFDEBFAA0FD9E" box="[1263,1307,603,629]" captionStart="Fig.1" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[221,1375,624,1881]" captionTargetId="figure@4.[208,1379,622,1889]" captionTargetPageId="4" captionText="Fig.1.DistributionandaedeagusvariabilityoftheQuediusmutilatusgroupintheTien-ShanMountains. Numbers are specified in Table 1. Empty symbols = typespcimens (for Q. kalabi precise locality of the holotype unknown);? = ambiguously labelled or undetermined material. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1167583/files/figure.png" pageId="12" pageNumber="13">Fig</figureCitation>
|
||
. 1). All newly studied specimens were collected at high elevations around
|
||
<quantity id="4C84E28D7047BF22FC31FDCFFBD9FD72" box="[961,1122,639,665]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="3.4" metricValueMin="2.0" pageId="12" pageNumber="13" unit="m" value="2700.0" valueMax="3400.0" valueMin="2000.0">2000–3400 m</quantity>
|
||
in July. For only two of them does the label specifies alpine meadows as the habitat, which is the same as for the
|
||
<typeStatus id="54C7F1CA7047BF22FB20FD13FAB8FD56" box="[1232,1283,675,701]" pageId="12" pageNumber="13">type</typeStatus>
|
||
material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3661CE37047BF22FF4DFD54FACCFCBA" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="8BC34F687047BF22FF4DFD54FEE3FD15" blockId="12.[189,344,740,766]" box="[189,344,740,766]" pageId="12" pageNumber="13">
|
||
<heading id="D08BF8047047BF22FF4DFD54FEE3FD15" bold="true" box="[189,344,740,766]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
||
<emphasis id="B908937A7047BF22FF4DFD54FEE3FD15" bold="true" box="[189,344,740,766]" pageId="12" pageNumber="13">Comparison</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC34F687047BF22FF4DFCA3FACCFCBA" blockId="12.[189,1399,787,849]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C7C34EB7047BF22FF4DFCA3FED1FCC6" authorityName="Smetana" authorityYear="2014" box="[189,362,787,813]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="equus">
|
||
<emphasis id="B908937A7047BF22FF4DFCA3FED1FCC6" box="[189,362,787,813]" italics="true" pageId="12" pageNumber="13">Quedius equus</emphasis>
|
||
</taxonomicName>
|
||
differs from all other species in the
|
||
<taxonomicName id="4C7C34EB7047BF22FCE6FCA3FC1CFCC6" authorityName="Eppelsheim A. Smetana" authorityYear="1888" box="[790,935,787,813]" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mutilatus">
|
||
<emphasis id="B908937A7047BF22FCE6FCA3FC1CFCC6" box="[790,935,787,813]" italics="true" pageId="12" pageNumber="13">Q. mutilatus</emphasis>
|
||
</taxonomicName>
|
||
group by the deep incision on the apex of the paramere and by fewer (1–3) sensory peg setae in the lateral groups arranged in longitudinal rows.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |