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<document id="6FC788A95F7F7ABD3D2FC18961A103E8" ID-DOI="10.5281/zenodo.13588506" ID-ISSN="2295-0214" ID-Zenodo-Dep="13588506" ID-ZooBank="08799CEE-C597-4D1C-90F2-569C42E9FAEB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1723128466313" checkinUser="felipe" docAuthor="Grootaert, Patrick, Raemdonck, Hugo &amp; O. D., Alain Drumont" docDate="2020" docId="03D487E4FFBD23068951FE48FC09C3B2" docLanguage="en" docName="BelgJEntomol.104.1-18.pdf" docOrigin="Belgian Journal of Entomology 104" docStyle="DocumentStyle:1C24E9253A34C90F9A25106B2B38F3D3.1:BelgJEntomol.2015-2021.journal_article.type1" docStyleId="1C24E9253A34C90F9A25106B2B38F3D3" docStyleName="BelgJEntomol.2015-2021.journal_article.type1" docStyleVersion="1" docTitle="Archicera avarorum Szilady 1934" docType="treatment" docVersion="3" lastPageNumber="18" masterDocId="FFEDFF9CFFBB23018B08FFF6FFB6C519" masterDocTitle="The Rhagionidae or Snipeflies of the Botanical Garden Jean Massart (Brussels-Capital Region, Belgium) with notes on the identity of the rare European species Archicera avarorum Szilády, 1934 and Ptiolina obscura (Fallén, 1814) (Diptera: Rhagionidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1725004488485" updateUser="ExternalLinkService">
<mods:mods id="918F6882A50C48E8F8C520EECEAE31CF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="43F1A9629D3693FBE7C0B2CB8F5600FA">
<mods:title id="C2623DDA61BC04AC664FA5DD5FD77654">The Rhagionidae or Snipeflies of the Botanical Garden Jean Massart (Brussels-Capital Region, Belgium) with notes on the identity of the rare European species Archicera avarorum Szilády, 1934 and Ptiolina obscura (Fallén, 1814) (Diptera: Rhagionidae)</mods:title>
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<mods:namePart id="55D4BE98C3BF3BF7273751293D68E58E">Grootaert, Patrick</mods:namePart>
<mods:affiliation id="D388B69025AF67D91AA47D6D91EC0158">O. D. Taxonomy and Phylogeny-Entomology, Royal Belgian Institute of Natural Sciences, Brussels, Vautier Street 29, B- 1000 Brussels.</mods:affiliation>
<mods:nameIdentifier id="A625C7C5D22AEB4386847E7400F0D912" type="email">pgrootaert@yahoo.co.uk</mods:nameIdentifier>
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<mods:namePart id="1E267A466B83B91B1AC46ED24E0B32B3">Raemdonck, Hugo</mods:namePart>
<mods:affiliation id="B8B31675F9A5EA6A3425735A2F8A63E8">O. D. Taxonomy and Phylogeny-Entomology, Royal Belgian Institute of Natural Sciences, Brussels, Vautier Street 29, B- 1000 Brussels.</mods:affiliation>
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<mods:namePart id="DFFE112E385209CED340A1F5B099C11D">O. D., Alain Drumont</mods:namePart>
<mods:affiliation id="CCB45441C20736E6284F0711443AC207">O. D. Taxonomy and Phylogeny-Entomology, Royal Belgian Institute of Natural Sciences, Brussels, Vautier Street 29, B- 1000 Brussels.</mods:affiliation>
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<mods:title id="DD5154F77BEB8A0B9DE7BA0406135563">Belgian Journal of Entomology</mods:title>
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<mods:date id="81987AB086CFF5D0E1027213711E8BE2">2020</mods:date>
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<mods:number id="7D0D3B40B6DEE75471BD222847197E8F">2020-09-17</mods:number>
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<mods:number id="71B4C0F06DF1B5388A58A968EFA8B90D">104</mods:number>
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<mods:identifier id="130E73D800DEB109F7597E377B57D5A5" type="ISSN">2295-0214</mods:identifier>
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<treatment id="03D487E4FFBD23068951FE48FC09C3B2" LSID="urn:lsid:plazi:treatment:03D487E4FFBD23068951FE48FC09C3B2" httpUri="http://treatment.plazi.org/id/03D487E4FFBD23068951FE48FC09C3B2" lastPageId="7" pageId="6">
<subSubSection id="C3676579FFBD23078951FE48FBB1C4C0" box="[601,1031,446,473]" pageId="6" type="nomenclature">
<paragraph id="8BC236F2FFBD23078951FE48FBB1C4C0" blockId="6.[601,1031,446,509]" box="[601,1031,446,473]" pageId="6">
<taxonomicName id="4C7D4D71FFBD23078951FE48FBB1C4C0" ID-CoL="G8KC" authority="Szilady, 1934" authorityName="Szilady" authorityYear="1934" box="[601,1031,446,473]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="avarorum">
<emphasis id="B909EAE0FFBD23078951FE48FBB1C4C0" bold="true" box="[601,1031,446,473]" pageId="6">
<emphasis id="B909EAE0FFBD23078951FE48FCE0C4C0" bold="true" box="[601,854,446,473]" italics="true" pageId="6">Archicera avarorum</emphasis>
Szilády, 1934
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3676579FFBD230789FAFE14FCD8C4E4" box="[754,878,482,509]" pageId="6" type="description">
<paragraph id="8BC236F2FFBD230789FAFE14FCD8C4E4" blockId="6.[601,1031,446,509]" box="[754,878,482,509]" pageId="6">
(
<figureCitation id="13462A77FFBD230789F2FE14FCD3C4E4" box="[762,869,482,509]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[247,284,1527,1549]" captionStartId-1="7.[247,283,1420,1442]" captionStartId-2="8.[247,284,1634,1656]" captionTargetBox-0="[318,1312,741,1506]" captionTargetBox-1="[257,1352,853,1364]" captionTargetBox-2="[247,1375,534,1613]" captionTargetId-0="figure-286@6.[317,1313,740,1507]" captionTargetId-1="figure-420@7.[246,1365,804,1400]" captionTargetId-2="figure-169@8.[246,1379,521,1614]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Fig. 3. Archicera avarorum Szilády, 1934. Female habitus. © Patrick Grootaert." captionText-1="Fig. 4. Archicera avarorum Szilády, 1934. Female: A. Antenna of left side; B. Antenna of right side of same specimen. Scale: 0.1 mm. © Patrick Grootaert." captionText-2="Fig. 5. Archicera avarorum Szilády, 1934. Female A. Tip of abdomen, dorsal view; B. Tip of abdomen, ventral view; C. Spermatheca; D. Palpus, proboscis and labella, with detail central tube of pseudotrachaea, lateral view; E. Labellae, ventral view. Scales 0.1 mm. © Patrick Grootaert." figureDoi-0="http://doi.org/10.5281/zenodo.13588524" figureDoi-1="http://doi.org/10.5281/zenodo.13588528" figureDoi-2="http://doi.org/10.5281/zenodo.13588534" httpUri-0="https://zenodo.org/record/13588524/files/figure.png" httpUri-1="https://zenodo.org/record/13588528/files/figure.png" httpUri-2="https://zenodo.org/record/13588534/files/figure.png" pageId="6">Figs 3–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3676579FFBD23078BFFFDE9FE6BC7C8" pageId="6" type="materials_examined">
<paragraph id="8BC236F2FFBD23078BFFFDE9FB1CC745" blockId="6.[247,1385,542,604]" pageId="6">
<materialsCitation id="3B153CAFFFBD23078BFFFDE9FCB0C745" collectingDate="2015-05-21" collectingDateMax="2015-05-28" collectingDateMin="2015-05-21" collectionCode="MT" country="Belgium" county="Auderghem" location="Jean Massart" municipality="Jardin" pageId="6" specimenCode="MT2" specimenCount="2" specimenCount-female="2" stateProvince="Region de Bruxelles-Capitale">
<smallCapsWord id="8D24A02EFFBD23078BFFFDE9FECEC72E" baselines="562,562" box="[247,376,543,569]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Material" pageId="6">MATERIAL</smallCapsWord>
<smallCapsWord id="8D24A02EFFBD23078A77FDD4FE48C72E" baselines="562" box="[383,510,546,567]" lowerCaseFontSize="9" mainFontSize="11" normCase="lower" normString="examined" pageId="6">EXAMINED</smallCapsWord>
:
<collectingCountry id="F36A7662FFBD23078918FDE8FDCDC721" box="[528,635,542,568]" name="Belgium" pageId="6">Belgium</collectingCountry>
,
<collectingRegion id="49B9F810FFBD23078981FDE8FC03C721" box="[649,949,542,568]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="6">Brussels-Capital Region</collectingRegion>
:
<collectingCounty id="62A34E7EFFBD230788CAFDE8FBE5C721" box="[962,1107,542,568]" pageId="6">Auderghem</collectingCounty>
,
<collectingMunicipality id="6BA6AC88FFBD23078F69FDE8FB1CC721" box="[1121,1194,542,568]" pageId="6">Jardin</collectingMunicipality>
botanique
<location id="8EA26029FFBD23078E3CFDE9FEEFC745" LSID="urn:lsid:plazi:treatment:03D487E4FFBD23068951FE48FC09C3B2:8EA26029FFBD23078E3CFDE9FEEFC745" country="Belgium" county="Auderghem" municipality="Jardin" name="Jean Massart" pageId="6" stateProvince="Region de Bruxelles-Capitale">Jean Massart</location>
,
<specimenCount id="9D7BFD7BFFBD23078A6EFDB4FE6BC742" box="[358,477,577,604]" count="2" pageId="6" type="female">2 females</specimenCount>
,
<date id="FFC31032FFBD23078AE1FDB4FD0FC745" box="[489,697,577,604]" pageId="6" value="2015-05-21" valueMax="2015-05-28" valueMin="2015-05-21">
<collectingDate id="EF87E9DAFFBD23078AE1FDB4FD0FC745" box="[489,697,577,604]" pageId="6" value="2015-05-21" valueMax="2015-05-28" valueMin="2015-05-21">21–28 May 2015</collectingDate>
</date>
,
<specimenCode id="DBDB9E89FFBD230789CEFDB4FCB0C745" box="[710,774,578,604]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="6">MT2</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B153CAFFFBD2307881AFDB4FB13C745" box="[786,1189,577,604]" collectingDate="2015-05-13" collectingDateMax="2015-05-21" collectingDateMin="2015-05-13" collectionCode="MT" country="Belgium" county="Auderghem" isTailEnumeration="true" location="Jean Massart" municipality="Jardin" pageId="6" specimenCode="MT1" specimenCount="1" specimenCount-female="1" stateProvince="Region de Bruxelles-Capitale">
<specimenCount id="9D7BFD7BFFBD2307881AFDB4FCCBC742" box="[786,893,577,604]" count="1" pageId="6" type="female">1 female</specimenCount>
,
<date id="FFC31032FFBD23078881FDB4FBECC745" box="[905,1114,577,604]" pageId="6" value="2015-05-13" valueMax="2015-05-21" valueMin="2015-05-13">
<collectingDate id="EF87E9DAFFBD23078881FDB4FBECC745" box="[905,1114,577,604]" pageId="6" value="2015-05-13" valueMax="2015-05-21" valueMin="2015-05-13">13–21 May 2015</collectingDate>
</date>
,
<specimenCode id="DBDB9E89FFBD23078F6EFDB4FB13C745" box="[1126,1189,578,604]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="6">MT1</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC236F2FFBD23078BFFFD87FE6BC7C8" blockId="6.[247,1386,625,722]" pageId="6">
<smallCapsWord id="8D24A02EFFBD23078BFFFD87FE26C793" baselines="644,645" box="[247,400,625,651]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Additional" pageId="6">ADDITIONAL</smallCapsWord>
<smallCapsWord id="8D24A02EFFBD23078A9FFD83FDFAC793" baselines="645" box="[407,588,629,650]" lowerCaseFontSize="9" mainFontSize="11" normCase="lower" normString="observations" pageId="6">OBSERVATIONS</smallCapsWord>
:
<materialsCitation id="3B153CAFFFBD23078955FD87FBB7C792" box="[605,1025,625,652]" collectingDate="2017-05-11" collectingDateMax="2017-05-17" collectingDateMin="2017-05-11" collectionCode="MT" pageId="6" specimenCode="MT6" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D7BFD7BFFBD23078955FD87FD63C792" box="[605,725,625,651]" count="2" pageId="6" type="female">2 females</specimenCount>
,
<date id="FFC31032FFBD230789E9FD87FC02C792" box="[737,948,625,652]" pageId="6" value="2017-05-11" valueMax="2017-05-17" valueMin="2017-05-11">
<collectingDate id="EF87E9DAFFBD230789E9FD87FC02C792" box="[737,948,625,652]" pageId="6" value="2017-05-11" valueMax="2017-05-17" valueMin="2017-05-11">11–17 May 2017</collectingDate>
</date>
,
<specimenCode id="DBDB9E89FFBD230788C9FD87FBB7C792" box="[961,1025,625,651]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="6">MT6</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B153CAFFFBD23078F06FD87FE81C7B7" collectingDate="2017-05-11" collectingDateMax="2017-05-17" collectingDateMin="2017-05-11" collectionCode="MT" pageId="6" specimenCode="MT2" specimenCount="5" specimenCount-female="5">
<specimenCount id="9D7BFD7BFFBD23078F06FD87FB30C792" box="[1038,1158,625,652]" count="5" pageId="6" type="female">5 females</specimenCount>
,
<date id="FFC31032FFBD23078F9AFD87FAD2C792" box="[1170,1380,625,652]" pageId="6" value="2017-05-11" valueMax="2017-05-17" valueMin="2017-05-11">
<collectingDate id="EF87E9DAFFBD23078F9AFD87FAD2C792" box="[1170,1380,625,652]" pageId="6" value="2017-05-11" valueMax="2017-05-17" valueMin="2017-05-11">11–17 May 2017</collectingDate>
</date>
,
<specimenCode id="DBDB9E89FFBD23078BFFFD62FE81C7B7" box="[247,311,660,686]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="6">MT2</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B153CAFFFBD23078A4CFD62FD69C7B6" box="[324,735,660,687]" collectingDate="2017-05-11" collectingDateMax="2017-05-17" collectingDateMin="2017-05-11" collectionCode="MT" pageId="6" specimenCode="MT5" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D7BFD7BFFBD23078A4CFD62FE07C7B7" box="[324,433,660,686]" count="1" pageId="6" type="female">1 female</specimenCount>
,
<date id="FFC31032FFBD23078AB6FD62FD27C7B7" box="[446,657,660,687]" pageId="6" value="2017-05-11" valueMax="2017-05-17" valueMin="2017-05-11">
<collectingDate id="EF87E9DAFFBD23078AB6FD62FD27C7B7" box="[446,657,660,687]" pageId="6" value="2017-05-11" valueMax="2017-05-17" valueMin="2017-05-11">11–17 May 2017</collectingDate>
</date>
,
<specimenCode id="DBDB9E89FFBD23078997FD62FD69C7B6" box="[671,735,660,687]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="6">MT5</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B153CAFFFBD230789E4FD62FE6FC7C8" collectingDate="2017-05-17" collectingDateMax="2017-05-24" collectingDateMin="2017-05-17" collectionCode="MT" collectorName="A. Drumont &amp; H. Raemdonck" pageId="6" specimenCode="MT2" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D7BFD7BFFBD230789E4FD62FCD3C7B7" box="[748,869,660,686]" count="2" pageId="6" type="female">2 females</specimenCount>
,
<date id="FFC31032FFBD2307887AFD62FBF3C7B7" box="[882,1093,660,687]" pageId="6" value="2017-05-17" valueMax="2017-05-24" valueMin="2017-05-17">
<collectingDate id="EF87E9DAFFBD2307887AFD62FBF3C7B7" box="[882,1093,660,687]" pageId="6" value="2017-05-17" valueMax="2017-05-24" valueMin="2017-05-17">17–24 May 2017</collectingDate>
</date>
,
<specimenCode id="DBDB9E89FFBD23078F5AFD62FB38C7B7" box="[1106,1166,660,686]" collectionCode="MT" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="6">MT2</specimenCode>
(leg.
<collectorName id="26885324FFBD23078FDFFD62FADCC7B6" box="[1239,1386,660,687]" pageId="6">A. Drumont</collectorName>
&amp;
<collectorName id="26885324FFBD23078A1EFD4EFE67C7C8" box="[278,465,695,722]" pageId="6">H. Raemdonck</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF02667AFFBD23078BFFFA01FBAAC315" ID-DOI="http://doi.org/10.5281/zenodo.13588524" ID-Zenodo-Dep="13588524" box="[247,1052,1526,1549]" httpUri="https://zenodo.org/record/13588524/files/figure.png" pageId="6" startId="6.[247,284,1527,1549]" targetBox="[318,1312,741,1506]" targetPageId="6" targetType="figure">
<paragraph id="8BC236F2FFBD23078BFFFA01FBAAC315" blockId="6.[247,1052,1526,1549]" box="[247,1052,1526,1549]" pageId="6">
Fig. 3.
<taxonomicName id="4C7D4D71FFBD23078A36FA01FD29C315" authority="Szilady, 1934" authorityName="Szilady" authorityYear="1934" box="[318,671,1527,1549]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="avarorum">
<emphasis id="B909EAE0FFBD23078A36FA01FDBAC315" box="[318,524,1527,1548]" italics="true" pageId="6">Archicera avarorum</emphasis>
Szilády, 1934
</taxonomicName>
. Female habitus. © Patrick Grootaert.
</paragraph>
</caption>
<subSubSection id="C3676579FFBD23068BFFF9D8FC09C3B2" lastPageId="7" pageId="6" type="discussion">
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<paragraph id="8BC236F2FFBD23078BFFF9A8FADFC277" blockId="6.[247,1386,1630,1902]" pageId="6">
A few more specimens of this very rare species were found in the year following the first sampling at the Garden, and thus examined and incorporated in the study. At first this species was confused with
<taxonomicName id="4C7D4D71FFBD23078AE9F952FC87C3A7" authority="Meigen, 1830" authorityName="Meigen" authorityYear="1830" box="[481,817,1700,1726]" class="Insecta" family="Rhagionidae" genus="Spania" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B909EAE0FFBD23078AE9F952FDCBC3A7" box="[481,637,1700,1726]" italics="true" pageId="6">Spania nigra</emphasis>
Meigen, 1830
</taxonomicName>
. However, in
<taxonomicName id="4C7D4D71FFBD230788D2F952FB8CC3A7" authorityName="Meigen" authorityYear="1830" box="[986,1082,1700,1726]" class="Insecta" family="Rhagionidae" genus="Spania" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B909EAE0FFBD230788D2F952FB8CC3A7" box="[986,1082,1700,1726]" italics="true" pageId="6">S. nigra</emphasis>
</taxonomicName>
the flagellomere or third antennal segment has a rectangular base and bears a much thinner “arista” -like projection at the ventral corner of its apex. In all
<specimenCount id="9D7BFD7BFFBD230789ECF91CFCD9C21D" box="[740,879,1770,1796]" count="13" pageId="6" type="female">13 females</specimenCount>
examined from the Garden, the entire flagellomere is broad throughout (
<figureCitation id="13462A77FFBD230789BDF8FBFCB3C23E" box="[693,773,1805,1831]" captionStart="Fig" captionStartId="6.[247,284,1527,1549]" captionTargetBox="[318,1312,741,1506]" captionTargetId="figure-286@6.[317,1313,740,1507]" captionTargetPageId="6" captionText="Fig. 3. Archicera avarorum Szilády, 1934. Female habitus. © Patrick Grootaert." figureDoi="http://doi.org/10.5281/zenodo.13588524" httpUri="https://zenodo.org/record/13588524/files/figure.png" pageId="6">Fig. 3</figureCitation>
), also with a rather square base and no other segmentation is visible on the flagellomere. Such morphology of the antenna is typical for
<emphasis id="B909EAE0FFBD23078BFFF8A2FE4EC277" box="[247,504,1876,1902]" italics="true" pageId="6">
<taxonomicName id="4C7D4D71FFBD23078BFFF8A2FE43C277" authorityName="Szilady" authorityYear="1934" box="[247,501,1876,1902]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" phylum="Arthropoda" rank="species" species="avarorum">Archicera avarorum</taxonomicName>
,
</emphasis>
a species and genus described by SZILÁDY (1934) on the base of two
</paragraph>
<paragraph id="8BC236F2FFBC23068BFFFF49FC7CC4EB" blockId="7.[247,1386,191,498]" pageId="7">
<specimenCount id="9D7BFD7BFFBC23068BFFFF49FEEFC5C0" box="[247,345,191,217]" count="7" pageId="7" type="female">females</specimenCount>
.
<specimenCount id="9D7BFD7BFFBC23068A6CFF49FE59C5C0" box="[356,495,191,217]" count="1" pageId="7" type="female">One female</specimenCount>
was found in Jasenak (
<collectingCountry id="F36A7662FFBC2306880EFF49FCD4C5C0" box="[774,866,191,217]" name="Croatia" pageId="7">Croatia</collectingCountry>
) and another in Spital (
<collectingRegion id="49B9F810FFBC23068F73FF49FB71C5C0" box="[1147,1223,191,217]" country="Austria" name="Steiermark" pageId="7">Styria</collectingRegion>
, Steirermark,
<collectingRegion id="49B9F810FFBC23068BFFFF14FE1BC5E5" box="[247,429,226,252]" country="Austria" name="Oberoesterreich" pageId="7">Oberösterreich</collectingRegion>
,
<collectingCountry id="F36A7662FFBC23068AB3FF14FDAFC5E5" box="[443,537,226,252]" name="Austria" pageId="7">Austria</collectingCountry>
). It was originally described as a subgenus of
<taxonomicName id="4C7D4D71FFBC23068F5DFF14FB00C5E5" authorityName="Zetterstedt" authorityYear="1842" box="[1109,1206,226,252]" class="Insecta" family="Rhagionidae" genus="Ptiolina" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAE0FFBC23068F5DFF14FB00C5E5" box="[1109,1206,226,252]" italics="true" pageId="7">Ptiolina</emphasis>
</taxonomicName>
and only later considered as a distinct genus. Eventually the genus
<taxonomicName id="4C7D4D71FFBC23068875FEF3FC43C406" authorityName="Szilady" authorityYear="1934" box="[893,1013,261,287]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAE0FFBC23068875FEF3FC43C406" box="[893,1013,261,287]" italics="true" pageId="7">Archicera</emphasis>
</taxonomicName>
was considered as a synonym of
<taxonomicName id="4C7D4D71FFBC23068A12FEDEFED8C45B" authorityName="Meigen" authorityYear="1830" box="[282,366,296,322]" class="Insecta" family="Rhagionidae" genus="Spania" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAE0FFBC23068A12FEDEFED8C45B" box="[282,366,296,322]" italics="true" pageId="7">Spania</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFEC4B03FFBC23068A96FEDEFDC9C45B" author="NAGATOMI A." box="[414,639,296,322]" pageId="7" pagination="31 - 70" refId="ref7347" refString="NAGATOMI A., 1982. - The genera of Rhagionidae (Diptera). Journal of Natural History, 16: 31 - 70." type="journal article" year="1982">
<smallCapsWord id="8D24A02EFFBC23068A96FEDEFD93C458" baselines="315,316" box="[414,549,296,322]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Nagatomi" pageId="7">NAGATOMI</smallCapsWord>
(1982)
</bibRefCitation>
and by
<bibRefCitation id="EFEC4B03FFBC230689ECFEDEFC34C45B" author="KERR P." box="[740,898,296,322]" pageId="7" pagination="1 - 133" refId="ref6927" refString="KERR P., 2010. - Phylogeny and classification of Rhagionidae, with implications for Tabanomorpha (Diptera: Brachycera). Zootaxa, 2592: 1 - 133." type="journal article" year="2010">
<smallCapsWord id="8D24A02EFFBC230689ECFEDEFC9EC458" baselines="315,316" box="[740,808,296,322]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Kerr" pageId="7">KERR</smallCapsWord>
(2010)
</bibRefCitation>
because the shape of the third antennal segment fell into the variability of
<taxonomicName id="4C7D4D71FFBC2306899EFEBDFCF9C47C" box="[662,847,331,357]" class="Insecta" family="Rhagionidae" genus="Spania" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B909EAE0FFBC2306899EFEBDFD5FC47C" box="[662,745,331,357]" italics="true" pageId="7">Spania</emphasis>
species.
</taxonomicName>
Since the
<specimenCount id="9D7BFD7BFFBC230688C3FEBAFBDFC47F" box="[971,1129,332,358]" count="2" pageId="7" type="generic" typeStatus="syntype">two syntypes</specimenCount>
of
<taxonomicName id="4C7D4D71FFBC23068F98FEBDFA9EC47C" authorityName="Szilady" authorityYear="1934" box="[1168,1320,331,357]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="avarorum">
<emphasis id="B909EAE0FFBC23068F98FEBDFA9EC47C" box="[1168,1320,331,357]" italics="true" pageId="7">A. avarorum</emphasis>
</taxonomicName>
were destroyed,
<bibRefCitation id="EFEC4B03FFBC23068A75FE99FDA5C491" author="PAPP L." box="[381,531,366,393]" pageId="7" pagination="189 - 194" refId="ref7409" refString="PAPP L., 2018. - Archicera Szilady, 1934: rediscovered and redescribed (Diptera: Rhagionidae). Folia entomologica hungarica, 7: 189 - 194." type="journal article" year="2018">
<smallCapsWord id="8D24A02EFFBC23068A75FE99FE0CC49E" baselines="386,386" box="[381,442,367,393]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Papp" pageId="7">PAPP</smallCapsWord>
(2018)
</bibRefCitation>
designated a female recently found in
<collectingCountry id="F36A7662FFBC230688FCFE98FBD7C491" box="[1012,1121,366,392]" name="Romania" pageId="7">Romania</collectingCountry>
as
<typeStatus id="54C68850FFBC23068F85FE99FB46C490" box="[1165,1264,367,393]" pageId="7" type="neotype">neotype</typeStatus>
.
<smallCapsWord id="8D24A02EFFBC23068FF5FE99FA8FC49E" baselines="386,386" box="[1277,1337,367,393]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Papp" pageId="7">PAPP</smallCapsWord>
did not wish to express himself on the status of
<taxonomicName id="4C7D4D71FFBC23068812FE67FC24C4B2" authorityName="Szilady" authorityYear="1934" box="[794,914,401,427]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAE0FFBC23068812FE67FC24C4B2" box="[794,914,401,427]" italics="true" pageId="7">Archicera</emphasis>
</taxonomicName>
and we follow his view and leave the question unanswered. His re-description fits entirely to our specimens and that is why we consider our specimens to be conspecific with
<taxonomicName id="4C7D4D71FFBC23068822FE2EFC74C4E8" authorityName="Szilady" authorityYear="1934" box="[810,962,472,498]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="avarorum">
<emphasis id="B909EAE0FFBC23068822FE2EFC74C4E8" box="[810,962,472,498]" italics="true" pageId="7">A. avarorum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC236F2FFBC23068BFFFDFEFE7AC60E" blockId="7.[247,1386,519,791]" pageId="7">
<taxonomicName id="4C7D4D71FFBC23068BFFFDFEFE39C738" authorityName="Szilady" authorityYear="1934" box="[247,399,520,546]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="avarorum">
<emphasis id="B909EAE0FFBC23068BFFFDFEFE39C738" box="[247,399,520,546]" italics="true" pageId="7">A. avarorum</emphasis>
</taxonomicName>
is a variable species. The shape of the flagellomere is slightly variable even on the same specimen (
<figureCitation id="13462A77FFBC23068AC9FDDCFDBDC75D" box="[449,523,554,580]" captionStart="Fig" captionStartId="7.[247,283,1420,1442]" captionTargetBox="[257,1352,853,1364]" captionTargetId="figure-420@7.[246,1365,804,1400]" captionTargetPageId="7" captionText="Fig. 4. Archicera avarorum Szilády, 1934. Female: A. Antenna of left side; B. Antenna of right side of same specimen. Scale: 0.1 mm. © Patrick Grootaert." figureDoi="http://doi.org/10.5281/zenodo.13588528" httpUri="https://zenodo.org/record/13588528/files/figure.png" pageId="7">Fig. 4</figureCitation>
) but it is always broad and flattened. The discal cell has 3 veins on its apex. The third vein generally does not reach the wing margin, which is a generic character for both
<taxonomicName id="4C7D4D71FFBC23068A3CFD87FE3EC792" authorityName="Meigen" authorityYear="1830" box="[308,392,625,651]" class="Insecta" family="Rhagionidae" genus="Spania" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAE0FFBC23068A3CFD87FE3EC792" box="[308,392,625,651]" italics="true" pageId="7">Spania</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7D4D71FFBC23068ACAFD86FD8CC793" authorityName="Szilady" authorityYear="1934" box="[450,570,624,650]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAE0FFBC23068ACAFD86FD8CC793" box="[450,570,624,650]" italics="true" pageId="7">Archicera</emphasis>
</taxonomicName>
. However, we found a specimen in which the third vein reaches the wing margin in one wing (
<figureCitation id="13462A77FFBC2306893CFD65FDCAC7B7" box="[564,636,659,686]" captionStart="Fig" captionStartId="6.[247,284,1527,1549]" captionTargetBox="[318,1312,741,1506]" captionTargetId="figure-286@6.[317,1313,740,1507]" captionTargetPageId="6" captionText="Fig. 3. Archicera avarorum Szilády, 1934. Female habitus. © Patrick Grootaert." figureDoi="http://doi.org/10.5281/zenodo.13588524" httpUri="https://zenodo.org/record/13588524/files/figure.png" pageId="7">Fig. 3</figureCitation>
) but not on the other wing. In another specimen the discal cell was apically open while there was also a specimen in which the two apical veins of the discal cell arose from the tip of the discal cell in one wing as was illustrated by
<bibRefCitation id="EFEC4B03FFBC23068F90FD2DFA99C7ED" author="PAPP L." box="[1176,1327,730,757]" pageId="7" pagination="189 - 194" refId="ref7409" refString="PAPP L., 2018. - Archicera Szilady, 1934: rediscovered and redescribed (Diptera: Rhagionidae). Folia entomologica hungarica, 7: 189 - 194." type="journal article" year="2018">
<smallCapsWord id="8D24A02EFFBC23068F90FD2DFB62C7EA" baselines="750,750" box="[1176,1236,731,757]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Papp" pageId="7">PAPP</smallCapsWord>
(2018)
</bibRefCitation>
in a
<taxonomicName id="4C7D4D71FFBC23068BFFFD0BFEFDC60E" authorityName="Meigen" authorityYear="1830" box="[247,331,765,791]" class="Insecta" family="Rhagionidae" genus="Spania" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAE0FFBC23068BFFFD0BFEFDC60E" box="[247,331,765,791]" italics="true" pageId="7">Spania</emphasis>
</taxonomicName>
specimen.
</paragraph>
<caption id="DF02667AFFBC23068BFFFA7AFD62C0A6" ID-DOI="http://doi.org/10.5281/zenodo.13588528" ID-Zenodo-Dep="13588528" httpUri="https://zenodo.org/record/13588528/files/figure.png" pageId="7" startId="7.[247,283,1420,1442]" targetBox="[257,1352,853,1364]" targetPageId="7" targetType="figure">
<paragraph id="8BC236F2FFBC23068BFFFA7AFD62C0A6" blockId="7.[247,1383,1420,1472]" pageId="7">
Fig. 4.
<taxonomicName id="4C7D4D71FFBC23068A4CFA7AFD1AC0B8" authority="Szilady, 1934" authorityName="Szilady" authorityYear="1934" box="[324,684,1420,1442]" class="Insecta" family="Rhagionidae" genus="Archicera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" phylum="Arthropoda" rank="species" species="avarorum">
<emphasis id="B909EAE0FFBC23068A4CFA7AFDA2C0B8" box="[324,532,1420,1441]" italics="true" pageId="7">Archicera avarorum</emphasis>
Szilády, 1934
</taxonomicName>
. Female: A. Antenna of left side; B. Antenna of right side of same specimen. Scale: 0.1 mm. © Patrick Grootaert.
</paragraph>
</caption>
<paragraph id="8BC236F2FFBC23068BFFFA17FC09C3B2" blockId="7.[247,1386,1505,1708]" pageId="7">
The palp consists apparently of a single segment as no separation of palpomeres was visible (
<figureCitation id="13462A77FFBC23068A08F9F3FEE9C306" box="[256,351,1540,1567]" captionStart="Fig" captionStartId="8.[247,284,1634,1656]" captionTargetBox="[247,1375,534,1613]" captionTargetId="figure-169@8.[246,1379,521,1614]" captionTargetPageId="8" captionText="Fig. 5. Archicera avarorum Szilády, 1934. Female A. Tip of abdomen, dorsal view; B. Tip of abdomen, ventral view; C. Spermatheca; D. Palpus, proboscis and labella, with detail central tube of pseudotrachaea, lateral view; E. Labellae, ventral view. Scales 0.1 mm. © Patrick Grootaert." figureDoi="http://doi.org/10.5281/zenodo.13588534" httpUri="https://zenodo.org/record/13588534/files/figure.png" pageId="7">Fig. 5D</figureCitation>
) The palp is broad with a truncate tip, bearing two small bristles at the tip and a series of long bristles dorsally at the base (
<figureCitation id="13462A77FFBC230689B1F9DEFCAAC35A" box="[697,796,1576,1603]" captionStart="Fig" captionStartId="8.[247,284,1634,1656]" captionTargetBox="[247,1375,534,1613]" captionTargetId="figure-169@8.[246,1379,521,1614]" captionTargetPageId="8" captionText="Fig. 5. Archicera avarorum Szilády, 1934. Female A. Tip of abdomen, dorsal view; B. Tip of abdomen, ventral view; C. Spermatheca; D. Palpus, proboscis and labella, with detail central tube of pseudotrachaea, lateral view; E. Labellae, ventral view. Scales 0.1 mm. © Patrick Grootaert." figureDoi="http://doi.org/10.5281/zenodo.13588534" httpUri="https://zenodo.org/record/13588534/files/figure.png" pageId="7">Fig. 5D</figureCitation>
). The proboscis is tubiform and the labellae are large. The pseudotracheae are thin, long and coiled and some might be furcate so that it is not clear if 8 or 9 pseudotracheae are present (
<figureCitation id="13462A77FFBC230689FCF998FCC3C391" box="[756,885,1646,1673]" captionStart="Fig" captionStartId="8.[247,284,1634,1656]" captionTargetBox="[247,1375,534,1613]" captionTargetId="figure-169@8.[246,1379,521,1614]" captionTargetPageId="8" captionText="Fig. 5. Archicera avarorum Szilády, 1934. Female A. Tip of abdomen, dorsal view; B. Tip of abdomen, ventral view; C. Spermatheca; D. Palpus, proboscis and labella, with detail central tube of pseudotrachaea, lateral view; E. Labellae, ventral view. Scales 0.1 mm. © Patrick Grootaert." figureDoi="http://doi.org/10.5281/zenodo.13588534" httpUri="https://zenodo.org/record/13588534/files/figure.png" pageId="7">Fig. 5D, E</figureCitation>
).
<figureCitation id="13462A77FFBC23068882F998FC51C391" box="[906,999,1646,1673]" captionStart="Fig" captionStartId="8.[247,284,1634,1656]" captionTargetBox="[247,1375,534,1613]" captionTargetId="figure-169@8.[246,1379,521,1614]" captionTargetPageId="8" captionText="Fig. 5. Archicera avarorum Szilády, 1934. Female A. Tip of abdomen, dorsal view; B. Tip of abdomen, ventral view; C. Spermatheca; D. Palpus, proboscis and labella, with detail central tube of pseudotrachaea, lateral view; E. Labellae, ventral view. Scales 0.1 mm. © Patrick Grootaert." figureDoi="http://doi.org/10.5281/zenodo.13588534" httpUri="https://zenodo.org/record/13588534/files/figure.png" pageId="7">Fig. 5A</figureCitation>
illustrates the female terminalia in dorsal position while
<figureCitation id="13462A77FFBC23068912F967FDC3C3B5" box="[538,629,1681,1708]" captionStart="Fig" captionStartId="8.[247,284,1634,1656]" captionTargetBox="[247,1375,534,1613]" captionTargetId="figure-169@8.[246,1379,521,1614]" captionTargetPageId="8" captionText="Fig. 5. Archicera avarorum Szilády, 1934. Female A. Tip of abdomen, dorsal view; B. Tip of abdomen, ventral view; C. Spermatheca; D. Palpus, proboscis and labella, with detail central tube of pseudotrachaea, lateral view; E. Labellae, ventral view. Scales 0.1 mm. © Patrick Grootaert." figureDoi="http://doi.org/10.5281/zenodo.13588534" httpUri="https://zenodo.org/record/13588534/files/figure.png" pageId="7">Fig. 5B</figureCitation>
shows the ventral position.
</paragraph>
</subSubSection>
</treatment>
</document>
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