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<document id="4300D2F4DC33E9E26538457C929C3843" ID-DOI="10.6620/ZS.2016.55-11" ID-ISSN="1810-522X" ID-PMC="PMC6511828" ID-PubMed="31966156" ID-Zenodo-Dep="12956889" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721941832834" checkinUser="felipe" docAuthor="Camerik, Anne M., Magowski, Wojciech Ł., Hawkes, Peter G., Ueckermann, Edward A., Ochoa, Ronald &amp; Bauchan, Gary R." docDate="2016" docId="03D487E09538B857FF43FD2BCBBEFF3E" docLanguage="en" docName="ZoolStud.55.11.1-21.pdf" docOrigin="Zoological Studies 55 (11)" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.4:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="4" docTitle="Zambedania sekhukhunensis Camerik, Magowski, Hawkes, Ueckermann, Ochoa &amp; Bauchan, 2016, sp. n." docType="treatment" docUuid="5AC608F6-69A9-4689-BEBA-818DF0A4FF39" docUuidSource="ZooBank" docVersion="4" lastPageNumber="18" masterDocId="FFEDFF98953CB846FF82FFACCA7AFF81" masterDocTitle="A New Species of Zambedania (Acari: Heterostigmatina: Pygmephoridae) from the Two Rivers Platinum Mine in South Africa and Notes on the Life-cycle of the Genus" masterLastPageNumber="21" masterPageNumber="1" pageNumber="5" updateTime="1722006054037" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B56F16E2C4663C90AD8BAC6559B0F247">A New Species of Zambedania (Acari: Heterostigmatina: Pygmephoridae) from the Two Rivers Platinum Mine in South Africa and Notes on the Life-cycle of the Genus</mods:title>
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<mods:namePart id="A7B2B6BDA4F67B37D1B4855543BB500E">Camerik, Anne M.</mods:namePart>
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<mods:namePart id="2B72C1629F95D626A3FC09BB6EDD498F">Magowski, Wojciech Ł.</mods:namePart>
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<mods:namePart id="FECEBBB99B26AC39DDDCB9DB0791D1D0">Hawkes, Peter G.</mods:namePart>
<mods:affiliation id="08B8675D1ED8FBF4F8AFDB084A52C8EB">AfriBugs CC, 341 27 th Avenue, Villieria, Pretoria, 0186, South Africa &amp; Department of Zoology and Entomology, University of Pretoria, Pretoria, Gauteng Province, 0002, South Africa.</mods:affiliation>
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<mods:namePart id="4E109CD663211042959F4627A2B8052F">Ueckermann, Edward A.</mods:namePart>
<mods:affiliation id="FB393BFBB04C0E7CF033C9074C0DA354">Agricultural Research Council - Plant Protection Research Institute, Pretoria, South Africa &amp; School of Biological Sciences / Zoology, North-West University, Potchefstroom 2520, South Africa. E-mail: UeckermannE @ arc. agric. za</mods:affiliation>
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<mods:namePart id="804C2863D4316F647355FCA70EF9A6BE">Ochoa, Ronald</mods:namePart>
<mods:affiliation id="879FF2D1AA604C4CEC174584807AFB33">United States Department of Agriculture, Agricultural Research Service, Systematic Entomology Laboratory, Beltsville, MD 20705, USA.</mods:affiliation>
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<taxonomicName id="4C7D4D759538B842FF43FD2BC84CFD21" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[193,566,647,672]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49538B842FF43FD2BC84CFD21" bold="true" box="[193,566,647,672]" italics="true" pageId="4" pageNumber="5">Zambedania sekhukhunensis</emphasis>
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Camerik and Magowski
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(
<figureCitation id="13462A739538B842FE1FFD6BC862FD61" box="[413,536,711,736]" captionStart-0="Fig" captionStart-1="Fig" captionStart-10="Fig" captionStart-11="Fig" captionStart-12="Fig" captionStart-13="Fig" captionStart-14="Fig" captionStart-15="Fig" captionStart-16="Fig" captionStart-17="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="5.[136,170,1950,1970]" captionStartId-1="6.[174,208,1950,1970]" captionStartId-10="12.[850,885,1925,1945]" captionStartId-11="13.[136,171,1926,1946]" captionStartId-12="13.[813,848,1926,1946]" captionStartId-13="14.[174,208,1953,1973]" captionStartId-14="15.[137,171,1953,1973]" captionStartId-15="16.[174,208,1951,1971]" captionStartId-16="17.[137,171,1950,1970]" captionStartId-17="18.[174,208,1953,1973]" captionStartId-2="7.[136,170,1950,1970]" captionStartId-3="8.[174,209,1923,1943]" captionStartId-4="8.[850,885,1923,1943]" captionStartId-5="9.[136,170,1949,1969]" captionStartId-6="10.[174,208,1264,1284]" captionStartId-7="11.[136,170,1159,1179]" captionStartId-8="11.[136,170,1951,1971]" captionStartId-9="12.[174,209,1925,1945]" captionTargetBox-0="[151,1401,822,1921]" captionTargetBox-1="[251,1357,1008,1915]" captionTargetBox-10="[102,1452,779,1899]" captionTargetBox-11="[138,736,432,1900]" captionTargetBox-12="[814,1413,638,1898]" captionTargetBox-13="[171,1451,705,1917]" captionTargetBox-14="[159,1402,1038,1928]" captionTargetBox-15="[292,1330,709,1916]" captionTargetBox-2="[129,1412,1054,1921]" captionTargetBox-3="[175,1451,842,1897]" captionTargetBox-6="[176,1449,252,1233]" captionTargetBox-7="[134,1414,249,1131]" captionTargetBox-8="[137,1416,1266,1881]" captionTargetId-0="figure-288@5.[136,1413,813,1934]" captionTargetId-1="figure-443@6.[241,1384,991,2075]" captionTargetId-10="figure-224@12.[87,907,775,1912]" captionTargetId-11="figure-121@13.[125,740,424,1922]" captionTargetId-12="figure-135@13.[802,1450,617,1912]" captionTargetId-13="figure-227@14.[159,1461,689,1927]" captionTargetId-14="figure-401@15.[136,1442,1029,1949]" captionTargetId-15="figure-149@16.[281,1344,690,1927]" captionTargetId-2="figure-344@7.[106,1433,1043,1973]" captionTargetId-3="figure-1@8.[748,1470,824,1909]" captionTargetId-6="figure-305@10.[174,1451,249,1235]" captionTargetId-7="figure-45@11.[121,1430,242,1139]" captionTargetId-8="figure-51@11.[123,1429,1248,1962]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-10="12" captionTargetPageId-11="13" captionTargetPageId-12="13" captionTargetPageId-13="14" captionTargetPageId-14="15" captionTargetPageId-15="16" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="11" captionText-0="Fig. 3. Zambedania sekhukhunensis sp. n. Female holotype. Idiosoma: A- dorsum, B- venter; sculpture omitted." captionText-1="Fig. 4. Zambedania sekhukhunensis sp. n. Female holotype. Gnathosoma A- dorsum, B- venter, C- pharyngeal pumps." captionText-10="Fig. 13. Zambedania sekhukhunensis sp. n. Male paratype. Leg II (right): dorsal view." captionText-11="Fig. 14. Zambedania sekhukhunensis sp. n. Male paratype. Leg III (right): dorsal view." captionText-12="Fig. 15. Zambedania sekhukhunensis sp. n. Male paratype. Leg IV (left): dorsal view." captionText-13="Fig. 16. Zambedania sekhukhunensis sp. n. Larva paratype. Idiosoma: A- dorsum, B- venter." captionText-14="Fig. 17. Zambedania sekhukhunensis sp. n. Larva paratype. Gnathosoma: A- dorsal view, B. ventral view, C- pharyngeal pump system." captionText-15="Fig. 18. Zambedania sekhukhunensis sp. n. Larva paratype. Leg I (right): dorsal view." captionText-16="Fig. 19. Zambedania sekhukhunensis sp. n. Larva paratype. Leg II (right): dorsal view." captionText-17="Fig. 20. Zambedania sekhukhunensis sp. n. Larva paratype. Leg III (right): dorsal view." captionText-2="Fig. 5. Zambedania sekhukhunensis sp. n. Female holotype. Leg I A- (left): antaxial view, B- tibiotarsus (left): paraxial view." captionText-3="Fig. 6. Zambedania sekhukhunensis sp. n. Female holotype. Leg II (right): dorsal view." captionText-4="Fig. 7. Zambedania sekhukhunensis sp. n. Female holotype. Leg III (right): dorsal view." captionText-5="Fig. 8. Zambedania sekhukhunensis sp. n. Female holotype. Leg IV (right): paraxial view." captionText-6="Fig. 9. Zambedania sekhukhunensis sp. n. Female. A- dorsal view; B- caput of bothridial seta, dorsal view; in front “pilose” setae; C- Leg I (right). Note TiTa in dorsal view. At distal end “turret” carrying claw with at its base dorsal pinnaculum of the “locking system”, flanked by part of counterpiece. Eupathidia dorsad on pinnaculi. The cluster d and k associated with the three solenidia. Ventro-laterad near to counter piece seta s followed by feathered pv and v. D- detail distal end of left TiTa I in lateral view, claw and butterfly-shaped counter piece made up of setae u’ - u”, below seta s, followed by “feathered” primiventrals pv’ and pv” (photos by G. Bauchan and R. Ochoa)." captionText-7="Fig. 10. Zambedania sekhukhunensis sp. n. Male paratype. Idiosoma: A- dorsum, B- venter; gnathosoma: C- dorsum, D- venter; ascalebar for idiosoma, b- scalebar for gnathosoma; sculpture omitted." captionText-8="Fig. 11. Zambedania sekhukhunensis sp. n. Male paratype. Genital capsule: A- dorsal view, B. ventral view." captionText-9="Fig. 12. Zambedania sekhukhunensis sp. n. Male paratype. Leg I (right): dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.12956919" figureDoi-1="http://doi.org/10.5281/zenodo.12956925" figureDoi-10="http://doi.org/10.5281/zenodo.12956973" figureDoi-11="http://doi.org/10.5281/zenodo.12956981" figureDoi-12="http://doi.org/10.5281/zenodo.12956989" figureDoi-13="http://doi.org/10.5281/zenodo.12956993" figureDoi-14="http://doi.org/10.5281/zenodo.12956997" figureDoi-15="http://doi.org/10.5281/zenodo.12957009" figureDoi-2="http://doi.org/10.5281/zenodo.12956931" figureDoi-3="http://doi.org/10.5281/zenodo.12956941" figureDoi-6="http://doi.org/10.5281/zenodo.12956951" figureDoi-7="http://doi.org/10.5281/zenodo.12956957" figureDoi-8="http://doi.org/10.5281/zenodo.12956967" httpUri-0="https://zenodo.org/record/12956919/files/figure.png" httpUri-1="https://zenodo.org/record/12956925/files/figure.png" httpUri-10="https://zenodo.org/record/12956973/files/figure.png" httpUri-11="https://zenodo.org/record/12956981/files/figure.png" httpUri-12="https://zenodo.org/record/12956989/files/figure.png" httpUri-13="https://zenodo.org/record/12956993/files/figure.png" httpUri-14="https://zenodo.org/record/12956997/files/figure.png" httpUri-15="https://zenodo.org/record/12957009/files/figure.png" httpUri-2="https://zenodo.org/record/12956931/files/figure.png" httpUri-3="https://zenodo.org/record/12956941/files/figure.png" httpUri-6="https://zenodo.org/record/12956951/files/figure.png" httpUri-7="https://zenodo.org/record/12956957/files/figure.png" httpUri-8="https://zenodo.org/record/12956967/files/figure.png" pageId="4" pageNumber="5">Figs. 3-20</figureCitation>
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urn:lsid:zoobank.org:act: 
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<emphasis id="B909EAE49538B842FF68FCF6CB1AFCF2" box="[234,352,858,883]" italics="true" pageId="4" pageNumber="5">Diagnosis</emphasis>
: The female of 
<taxonomicName id="4C7D4D759538B842FDA6FCF6C97DFCF2" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[548,775,858,883]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sekhukhunensis">
<emphasis id="B909EAE49538B842FDA6FCF6C97DFCF2" box="[548,775,858,883]" italics="true" pageId="4" pageNumber="5">Z. sekhukhunensis</emphasis>
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is most similar to that of 
<taxonomicName id="4C7D4D759538B842FE78FCD6C8FFFC12" authorityName="Mahunka" authorityYear="1972" box="[506,645,890,915]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B909EAE49538B842FE78FCD6C8FFFC12" box="[506,645,890,915]" italics="true" pageId="4" pageNumber="5">Z. africana</emphasis>
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in having leg IV terminating in two claws, the stigmata being undivided and aggenital plate smooth (Ag). However, the species is uniquely different from 
<taxonomicName id="4C7D4D759538B842FF72FC56CB0CFB92" authorityName="Mahunka" authorityYear="1972" box="[240,374,1018,1043]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B909EAE49538B842FF72FC56CB0CFB92" box="[240,374,1018,1043]" italics="true" pageId="4" pageNumber="5">Z. africana</emphasis>
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by the following combination of characteristics: stigmatal peritremes about half (vs. about one third) the length of the prodorsal shield (PdS); except for setae 
<emphasis id="B909EAE49538B842FE44FBF6CBB7FBF2" box="[454,461,1114,1139]" italics="true" pageId="4" pageNumber="5">f</emphasis>
, no other idiosomal dorsal setae on protuberances (vs. setae 
<emphasis id="B909EAE49538B842FDCEFBD6C823FB12" box="[588,601,1146,1171]" italics="true" pageId="4" pageNumber="5">c</emphasis>
, 
<emphasis id="B909EAE49538B842FDEBFBD6C802FB12" box="[617,632,1146,1171]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, 
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, 
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and 
<emphasis id="B909EAE49538B842FD6BFBD6C97DFB12" box="[745,775,1146,1171]" italics="true" pageId="4" pageNumber="5">h1</emphasis>
on protuberances); setae 
<emphasis id="B909EAE49538B842FE64FB36CB8FFB32" box="[486,501,1178,1203]" italics="true" pageId="4" pageNumber="5">e</emphasis>
about 0.9x as long as setae 
<emphasis id="B909EAE49538B842FF7BFB16CB7AFB52" box="[249,256,1210,1235]" italics="true" pageId="4" pageNumber="5">f</emphasis>
(vs 0.4 as long); 
<emphasis id="B909EAE49538B842FE5AFB16CB8CFB52" box="[472,502,1210,1235]" italics="true" pageId="4" pageNumber="5">h1</emphasis>
about 1.5x as long as 
<emphasis id="B909EAE49538B842FF2CFB76CAB6FB72" box="[174,204,1242,1267]" italics="true" pageId="4" pageNumber="5">h2</emphasis>
(vs. subequal in length); tibiotarsal solenidion ω is stemmed and about 1.3x the length of Φ1, 
<emphasis id="B909EAE49538B842FD7AFB56C97DFA92" box="[760,775,1274,1299]" italics="true" pageId="4" pageNumber="5">2</emphasis>
(vs. sessile and 1.7x); TaII: 
<emphasis id="B909EAE49538B842FD92FAB6C849FAB2" box="[528,563,1306,1331]" italics="true" pageId="4" pageNumber="5">pv’</emphasis>
, 
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barbed (vs. feathered).
</paragraph>
<paragraph id="8BC236F69538B842FF68FAF6CB81FA12" blockId="4.[171,775,858,1492]" pageId="4" pageNumber="5">
<emphasis id="B909EAE49538B842FF68FAF6CBFDFAF2" box="[234,391,1370,1395]" italics="true" pageId="4" pageNumber="5">D e s c r i p t i o n</emphasis>
: (measurements- 
<tableCitation id="C6FF034D9538B842FD0EFAF6C88BFAF2" box="[652,753,1370,1395]" captionStart="Table 1" captionStartId="3.[136,204,253,278]" captionText="Table 1. Measurements of Zambedania sekhukhunensis sp. n. (holotype and paratypes phoretic females, males and larva)" pageId="4" pageNumber="5">Table 1</tableCitation>
). Female. (
<figureCitation id="13462A739538B842FE9FFAD6CBF0FA12" box="[285,394,1402,1427]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="5.[136,170,1950,1970]" captionStartId-1="6.[174,208,1950,1970]" captionStartId-2="7.[136,170,1950,1970]" captionStartId-3="8.[174,209,1923,1943]" captionStartId-4="8.[850,885,1923,1943]" captionStartId-5="9.[136,170,1949,1969]" captionStartId-6="10.[174,208,1264,1284]" captionTargetBox-0="[151,1401,822,1921]" captionTargetBox-1="[251,1357,1008,1915]" captionTargetBox-2="[129,1412,1054,1921]" captionTargetBox-3="[175,1451,842,1897]" captionTargetBox-6="[176,1449,252,1233]" captionTargetId-0="figure-288@5.[136,1413,813,1934]" captionTargetId-1="figure-443@6.[241,1384,991,2075]" captionTargetId-2="figure-344@7.[106,1433,1043,1973]" captionTargetId-3="figure-1@8.[748,1470,824,1909]" captionTargetId-6="figure-305@10.[174,1451,249,1235]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-6="10" captionText-0="Fig. 3. Zambedania sekhukhunensis sp. n. Female holotype. Idiosoma: A- dorsum, B- venter; sculpture omitted." captionText-1="Fig. 4. Zambedania sekhukhunensis sp. n. Female holotype. Gnathosoma A- dorsum, B- venter, C- pharyngeal pumps." captionText-2="Fig. 5. Zambedania sekhukhunensis sp. n. Female holotype. Leg I A- (left): antaxial view, B- tibiotarsus (left): paraxial view." captionText-3="Fig. 6. Zambedania sekhukhunensis sp. n. Female holotype. Leg II (right): dorsal view." captionText-4="Fig. 7. Zambedania sekhukhunensis sp. n. Female holotype. Leg III (right): dorsal view." captionText-5="Fig. 8. Zambedania sekhukhunensis sp. n. Female holotype. Leg IV (right): paraxial view." captionText-6="Fig. 9. Zambedania sekhukhunensis sp. n. Female. A- dorsal view; B- caput of bothridial seta, dorsal view; in front “pilose” setae; C- Leg I (right). Note TiTa in dorsal view. At distal end “turret” carrying claw with at its base dorsal pinnaculum of the “locking system”, flanked by part of counterpiece. Eupathidia dorsad on pinnaculi. The cluster d and k associated with the three solenidia. Ventro-laterad near to counter piece seta s followed by feathered pv and v. D- detail distal end of left TiTa I in lateral view, claw and butterfly-shaped counter piece made up of setae u’ - u”, below seta s, followed by “feathered” primiventrals pv’ and pv” (photos by G. Bauchan and R. Ochoa)." figureDoi-0="http://doi.org/10.5281/zenodo.12956919" figureDoi-1="http://doi.org/10.5281/zenodo.12956925" figureDoi-2="http://doi.org/10.5281/zenodo.12956931" figureDoi-3="http://doi.org/10.5281/zenodo.12956941" figureDoi-6="http://doi.org/10.5281/zenodo.12956951" httpUri-0="https://zenodo.org/record/12956919/files/figure.png" httpUri-1="https://zenodo.org/record/12956925/files/figure.png" httpUri-2="https://zenodo.org/record/12956931/files/figure.png" httpUri-3="https://zenodo.org/record/12956941/files/figure.png" httpUri-6="https://zenodo.org/record/12956951/files/figure.png" pageId="4" pageNumber="5">Figs. 3-9</figureCitation>
, 
<tableCitation id="C6FF034D9538B842FE17FAD6CB95FA12" box="[405,495,1402,1427]" captionStart="Table 2" captionStartId="3.[136,204,1598,1623]" captionText="Table 2. Leg chaeto- and solenidiotaxy of the genus Zambedania sekhukhunensis sp. n. females SEGMENT LEG I LEG II LEG III LEG IV Nr SIGLA Nr SIGLA Nr SIGLA Nr SIGLA TROCHANTER 1 v' 1 v' 1 v' 1 v' FEMUR 3 d,l',v&quot; 3 d,l',v&quot; 2 d,v' 2 d,v&quot; GENU 4 l',l&quot;,v', v&quot; 3 l',l&quot;,v&quot; 2 l',v.&quot; 1 v' TIBIOTARSUS Leg I only 13 d,l',l&quot;,v',v&quot;,k,pl',pl&quot;,pv',pv&quot;,u',u&quot;,s TIBIA: 4 d,l',v',v&quot; 4 d,l',v',v&quot; 4 d,l',v',v&quot; Eupathidia 5 tc',tc&quot;,p&quot;,ft',ft&quot; 0 0 0 Solenidia 3 ω1,Φ1,Φ2 1 Φ 1 Φ 1 Φ TARSUS 1 Claws 2 2 1 or 2 Tactile setae 6 pl&quot;,tc',tc&quot;,pv',pv&quot;,u 6 pl&quot;,tc',tc&quot;,pv',pv&quot;,u 6 pl&quot;,tc',tc&quot;,pv',pv&quot;,u Solenidia 1 ω 0 0" pageId="4" pageNumber="5">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BC236F69538B842FF68FA36CFD1FF3E" blockId="4.[171,775,858,1492]" lastBlockId="4.[1325,1451,168,191]" pageId="4" pageNumber="5">
Gnathosoma (
<figureCitation id="13462A739538B842FE28FA36C837FA32" box="[426,589,1434,1459]" captionStart="Fig" captionStartId="6.[174,208,1950,1970]" captionTargetBox="[251,1357,1008,1915]" captionTargetId="figure-443@6.[241,1384,991,2075]" captionTargetPageId="6" captionText="Fig. 4. Zambedania sekhukhunensis sp. n. Female holotype. Gnathosoma A- dorsum, B- venter, C- pharyngeal pumps." figureDoi="http://doi.org/10.5281/zenodo.12956925" httpUri="https://zenodo.org/record/12956925/files/figure.png" pageId="4" pageNumber="5">Figs. 4A, B</figureCitation>
). Orientation: hypognathous; shape ovoid. Cheliceral setae page 5 of 21
</paragraph>
<paragraph id="8BC236F69538B842FCD0FF51CF7EFDB7" blockId="4.[850,1452,253,1494]" pageId="4" pageNumber="5">
(
<emphasis id="B909EAE49538B842FCDEFF51C9F2FE97" box="[860,904,253,278]" italics="true" pageId="4" pageNumber="5">ch1</emphasis>
smooth, 
<emphasis id="B909EAE49538B842FB83FF51CE57FE97" box="[1025,1069,253,278]" italics="true" pageId="4" pageNumber="5">ch2</emphasis>
barbed) setiform, subequal in size; 
<emphasis id="B909EAE49538B842FC1BFEB1C9BAFEB4" box="[921,960,285,310]" italics="true" pageId="4" pageNumber="5">
ch 
<subScript id="17F934B39538B842FC35FE8AC9BAFEB4" attach="left" box="[951,960,294,309]" fontSize="6" pageId="4" pageNumber="5">1</subScript>
</emphasis>
located antero- mediad of 
<emphasis id="B909EAE49538B842FAA0FEB1CF45FEB7" box="[1314,1343,285,310]" italics="true" pageId="4" pageNumber="5">ch</emphasis>
<subScript id="17F934B39538B842FAC2FE8ACF33FEB4" attach="left" box="[1344,1353,294,309]" fontSize="6" pageId="4" pageNumber="5">2</subScript>
. Palpal supracoxal setae (
<emphasis id="B909EAE49538B842FBCBFE91CE12FED7" box="[1097,1128,317,342]" italics="true" pageId="4" pageNumber="5">pp</emphasis>
) present. Dorso-median apodeme prominent. The antero-laterad pedipalpal segments are fused, bearing smooth setae 
<emphasis id="B909EAE49538B842FAFEFED1CFD1FE17" box="[1404,1451,381,406]" italics="true" pageId="4" pageNumber="5">dFe</emphasis>
and 
<emphasis id="B909EAE49538B842FC08FE31C9C7FE37" box="[906,957,413,438]" italics="true" pageId="4" pageNumber="5">dGe</emphasis>
. Subcapitulum (= infracapitulum) with a pair of slender, simple, smooth setae (
<emphasis id="B909EAE49538B842FAB6FE11CF2BFE57" box="[1332,1361,445,470]" italics="true" pageId="4" pageNumber="5">su</emphasis>
) and a mushroom-shaped accessory setigenous structure (
<emphasis id="B909EAE49538B842FCD9FE51C9FFFD97" box="[859,901,509,534]" italics="true" pageId="4" pageNumber="5">ass</emphasis>
) located anteromedially and laterally flanked by a clavate, non-striated solenidion.
</paragraph>
<paragraph id="8BC236F69538B842FC0DFD91CF15FCF7" blockId="4.[850,1452,253,1494]" pageId="4" pageNumber="5">
Pharyngeal system (
<figureCitation id="13462A739538B842FB19FD91CF77FDD6" box="[1179,1293,573,599]" captionStart="Fig" captionStartId="6.[174,208,1950,1970]" captionTargetBox="[251,1357,1008,1915]" captionTargetId="figure-443@6.[241,1384,991,2075]" captionTargetPageId="6" captionText="Fig. 4. Zambedania sekhukhunensis sp. n. Female holotype. Gnathosoma A- dorsum, B- venter, C- pharyngeal pumps." figureDoi="http://doi.org/10.5281/zenodo.12956925" httpUri="https://zenodo.org/record/12956925/files/figure.png" pageId="4" pageNumber="5">Figs. 4C</figureCitation>
) subdivided into three striate, muscular pumps; in ventral view extending from the gnathosomal foramen to beyond apodeme 2. The first (anterior) pump is suboval, rather short and is immediately attached to the second pump. The second is narrow, rectangular and the longest of the three pumps. A short stretch of pharyngeal tube connects the second to the third pump, which is the widest and sclerotized at its anterior and posterior edges.
</paragraph>
<paragraph id="8BC236F69538B842FC0DFCD1CF05FA57" blockId="4.[850,1452,253,1494]" pageId="4" pageNumber="5">
Idiosomal dorsum (
<figureCitation id="13462A739538B842FB09FCD1CF7AFC17" box="[1163,1280,893,918]" captionStart="Fig" captionStartId="5.[136,170,1950,1970]" captionTargetBox="[151,1401,822,1921]" captionTargetId="figure-288@5.[136,1413,813,1934]" captionTargetPageId="5" captionText="Fig. 3. Zambedania sekhukhunensis sp. n. Female holotype. Idiosoma: A- dorsum, B- venter; sculpture omitted." figureDoi="http://doi.org/10.5281/zenodo.12956919" httpUri="https://zenodo.org/record/12956919/files/figure.png" pageId="4" pageNumber="5">Figs. 3A</figureCitation>
, 
<figureCitation id="13462A739538B842FA93FCD1CF43FC17" box="[1297,1337,893,918]" captionStart="Fig" captionStartId="10.[174,208,1264,1284]" captionTargetBox="[176,1449,252,1233]" captionTargetId="figure-305@10.[174,1451,249,1235]" captionTargetPageId="10" captionText="Fig. 9. Zambedania sekhukhunensis sp. n. Female. A- dorsal view; B- caput of bothridial seta, dorsal view; in front “pilose” setae; C- Leg I (right). Note TiTa in dorsal view. At distal end “turret” carrying claw with at its base dorsal pinnaculum of the “locking system”, flanked by part of counterpiece. Eupathidia dorsad on pinnaculi. The cluster d and k associated with the three solenidia. Ventro-laterad near to counter piece seta s followed by feathered pv and v. D- detail distal end of left TiTa I in lateral view, claw and butterfly-shaped counter piece made up of setae u’ - u”, below seta s, followed by “feathered” primiventrals pv’ and pv” (photos by G. Bauchan and R. Ochoa)." figureDoi="http://doi.org/10.5281/zenodo.12956951" httpUri="https://zenodo.org/record/12956951/files/figure.png" pageId="4" pageNumber="5">9A</figureCitation>
). Cuticle strongly sclerotized with uniform sparse punctate sculpturing. Prodorsal shield (PdS) with long, slitlike, obliquely placed stigmata reaching its anterior edge; capitate sensillae (
<emphasis id="B909EAE49538B842FBF9FC51CEDEFB97" box="[1147,1188,1021,1046]" italics="true" pageId="4" pageNumber="5">sc1</emphasis>
) with tiny barbs (
<figureCitation id="13462A739538B842FAF2FC51C90DFBB7" captionStart="Fig" captionStartId="5.[136,170,1950,1970]" captionTargetBox="[151,1401,822,1921]" captionTargetId="figure-288@5.[136,1413,813,1934]" captionTargetPageId="5" captionText="Fig. 3. Zambedania sekhukhunensis sp. n. Female holotype. Idiosoma: A- dorsum, B- venter; sculpture omitted." figureDoi="http://doi.org/10.5281/zenodo.12956919" httpUri="https://zenodo.org/record/12956919/files/figure.png" pageId="4" pageNumber="5">Figs. 3A</figureCitation>
, 
<figureCitation id="13462A739538B842FC07FBB1C9DDFBB7" box="[901,935,1053,1078]" captionStart="Fig" captionStartId="10.[174,208,1264,1284]" captionTargetBox="[176,1449,252,1233]" captionTargetId="figure-305@10.[174,1451,249,1235]" captionTargetPageId="10" captionText="Fig. 9. Zambedania sekhukhunensis sp. n. Female. A- dorsal view; B- caput of bothridial seta, dorsal view; in front “pilose” setae; C- Leg I (right). Note TiTa in dorsal view. At distal end “turret” carrying claw with at its base dorsal pinnaculum of the “locking system”, flanked by part of counterpiece. Eupathidia dorsad on pinnaculi. The cluster d and k associated with the three solenidia. Ventro-laterad near to counter piece seta s followed by feathered pv and v. D- detail distal end of left TiTa I in lateral view, claw and butterfly-shaped counter piece made up of setae u’ - u”, below seta s, followed by “feathered” primiventrals pv’ and pv” (photos by G. Bauchan and R. Ochoa)." figureDoi="http://doi.org/10.5281/zenodo.12956951" httpUri="https://zenodo.org/record/12956951/files/figure.png" pageId="4" pageNumber="5">9B</figureCitation>
) are placed in circular bothridia; two pairs of well-developed tapered, setiform, barbed setae located posteriad of the stigmata; of these, vertical (
<emphasis id="B909EAE49538B842FCDEFBD1C902FB17" box="[860,888,1149,1174]" italics="true" pageId="4" pageNumber="5">v2</emphasis>
) anterolaterad and shorter than the long and more robust scapular setae (
<emphasis id="B909EAE49538B842FB43FB31CE96FB37" box="[1217,1260,1181,1206]" italics="true" pageId="4" pageNumber="5">sc2</emphasis>
). Opisthosoma elliptical, five segmented with four dorsal segments: C, D, EF (coalesced) and H; the fifth segment, pseudanal shield (Ps) located caudoventrally. All segments bear two pairs of setae, except D which has one pair only. All dorsal setae very long, tapered, setiform and barbed, except setae 
<emphasis id="B909EAE49538B842FC19FAD1C9C2FA17" box="[923,952,1405,1430]" italics="true" pageId="4" pageNumber="5">v2</emphasis>
which is similar in form to the other dorsal setae but less than one third as long. Only seta 
<emphasis id="B909EAE49538B842FA26FA31CFD1FA37" box="[1444,1451,1437,1462]" italics="true" pageId="4" pageNumber="5">f</emphasis>
on a short protuberance. No cupules observed.
</paragraph>
<caption id="DF02667E9538B842FF2CF98CCEBCF824" pageId="4" pageNumber="5" startId="4.[174,242,1568,1593]" targetType="table">
<paragraph id="8BC236F69538B842FF2CF98CCF66F9B8" blockId="4.[174,1427,1568,1957]" box="[174,1308,1568,1593]" pageId="4" pageNumber="5">
<emphasis id="B909EAE49538B842FF2CF98CCB75F9B8" bold="true" box="[174,271,1568,1593]" pageId="4" pageNumber="5">Table 3.</emphasis>
Leg chaeto- and solenidiotaxy of the genus 
<taxonomicName id="4C7D4D759538B842FCA0F98CCE07F9B8" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[802,1149,1568,1593]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49538B842FCA0F98CCE07F9B8" box="[802,1149,1568,1593]" italics="true" pageId="4" pageNumber="5">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9538B842FB06F98CCEBFF9B8" box="[1156,1221,1568,1593]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
males.
</paragraph>
<paragraph id="8BC236F69538B842FF30F9CDCF18F9F4" blockId="4.[174,1427,1568,1957]" box="[178,1378,1633,1653]" pageId="4" pageNumber="5">SEGMENT LEG I LEG II LEG III LEG IV</paragraph>
<paragraph id="8BC236F69538B842FF30F935CE93F949" blockId="4.[174,1427,1568,1957]" pageId="4" pageNumber="5">
SEGMENT Nr SIGLA Nr SIGLA Nr SIGLA Nr SIGLA TROCHANTER 1 
<emphasis id="B909EAE49538B842FE5AF918CB9CF949" box="[472,486,1716,1736]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
1 
<emphasis id="B909EAE49538B842FD5BF918C89DF949" box="[729,743,1716,1736]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
1 
<emphasis id="B909EAE49538B842FC58F918C992F949" box="[986,1000,1716,1736]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
1 
<emphasis id="B909EAE49538B842FB59F918CE93F949" box="[1243,1257,1716,1736]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
</paragraph>
<paragraph id="8BC236F69538B842FF30F97CCE87F965" blockId="4.[174,1427,1568,1957]" box="[178,1277,1744,1764]" pageId="4" pageNumber="5">
FEMUR 3 
<emphasis id="B909EAE49538B842FE5AF97CCB99F965" box="[472,483,1744,1764]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, 
<emphasis id="B909EAE49538B842FE6BF97CCB8BF965" box="[489,497,1744,1764]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FE75F97CC872F965" box="[503,520,1744,1764]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
3 
<emphasis id="B909EAE49538B842FD5BF97CC89EF965" box="[729,740,1744,1764]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, 
<emphasis id="B909EAE49538B842FD68F97CC888F965" box="[746,754,1744,1764]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FD7AF97CC973F965" box="[760,777,1744,1764]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
2 
<emphasis id="B909EAE49538B842FC58F97CC99FF965" box="[986,997,1744,1764]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, 
<emphasis id="B909EAE49538B842FC69F97CC983F965" box="[1003,1017,1744,1764]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
2 
<emphasis id="B909EAE49538B842FB59F97CCE9CF965" box="[1243,1254,1744,1764]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, 
<emphasis id="B909EAE49538B842FB6EF97CCE87F965" box="[1260,1277,1744,1764]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
</paragraph>
<paragraph id="8BC236F69538B842FF30F947CE93F881" blockId="4.[174,1427,1568,1957]" box="[178,1257,1771,1792]" pageId="4" pageNumber="5">
GENU 4 
<emphasis id="B909EAE49538B842FE5AF940CB9AF881" box="[472,480,1772,1792]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FE64F940CB88F881" box="[486,498,1772,1792]" italics="true" pageId="4" pageNumber="5">l&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FE75F940C87FF881" box="[503,517,1772,1792]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
, 
<emphasis id="B909EAE49538B842FD92F940C85BF881" box="[528,545,1772,1792]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
3 
<emphasis id="B909EAE49538B842FD5BF940C89BF881" box="[729,737,1772,1792]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FD65F940C889F881" box="[743,755,1772,1792]" italics="true" pageId="4" pageNumber="5">l&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FD7AF940C973F881" box="[760,777,1772,1792]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
2 
<emphasis id="B909EAE49538B842FC58F940C998F881" box="[986,994,1772,1792]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FC6AF940C983F881" box="[1000,1017,1772,1792]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
1 
<emphasis id="B909EAE49538B842FB59F940CE93F881" box="[1243,1257,1772,1792]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
</paragraph>
<paragraph id="8BC236F69538B842FF30F8ABCF64F89A" blockId="4.[174,1427,1568,1957]" box="[178,1310,1799,1819]" pageId="4" pageNumber="5">
TIBIA 6 
<emphasis id="B909EAE49538B842FE5AF8ABCB9AF89A" box="[472,480,1799,1819]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FE64F8ABCB88F89A" box="[486,498,1799,1819]" italics="true" pageId="4" pageNumber="5">l&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FE75F8ABC87FF89A" box="[503,517,1799,1819]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
, 
<emphasis id="B909EAE49538B842FD89F8ABC866F89A" box="[523,540,1799,1819]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FDA3F8ABC851F89A" box="[545,555,1799,1819]" italics="true" pageId="4" pageNumber="5">k</emphasis>
, 
<emphasis id="B909EAE49538B842FDB3F8ABC846F89A" box="[561,572,1799,1819]" italics="true" pageId="4" pageNumber="5">d</emphasis>
4 
<emphasis id="B909EAE49538B842FD5BF8ABC89EF89A" box="[729,740,1799,1819]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, 
<emphasis id="B909EAE49538B842FD68F8ABC888F89A" box="[746,754,1799,1819]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FD7AF8ABC97CF89A" box="[760,774,1799,1819]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
, 
<emphasis id="B909EAE49538B842FC89F8ABC966F89A" box="[779,796,1799,1819]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
4 
<emphasis id="B909EAE49538B842FC58F8ABC99FF89A" box="[986,997,1799,1819]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, 
<emphasis id="B909EAE49538B842FC69F8ABC989F89A" box="[1003,1011,1799,1819]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FC7BF8ABCE7DF89A" box="[1017,1031,1799,1819]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
, 
<emphasis id="B909EAE49538B842FB8EF8ABCE67F89A" box="[1036,1053,1799,1819]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
4 
<emphasis id="B909EAE49538B842FB5EF8ABCE9DF89A" box="[1244,1255,1799,1819]" italics="true" pageId="4" pageNumber="5">d</emphasis>
, 
<emphasis id="B909EAE49538B842FB6EF8ABCE8EF89A" box="[1260,1268,1799,1819]" italics="true" pageId="4" pageNumber="5">l'</emphasis>
, 
<emphasis id="B909EAE49538B842FB78F8ABCF72F89A" box="[1274,1288,1799,1819]" italics="true" pageId="4" pageNumber="5">v'</emphasis>
, 
<emphasis id="B909EAE49538B842FA8FF8ABCF64F89A" box="[1293,1310,1799,1819]" italics="true" pageId="4" pageNumber="5">v&quot;</emphasis>
</paragraph>
<paragraph id="8BC236F69538B842FF30F88ECE99F8B4" blockId="4.[174,1427,1568,1957]" box="[178,1251,1826,1847]" pageId="4" pageNumber="5">
Solenidia 2 Φ 
<emphasis id="B909EAE49538B842FE61F88FCB94F8B6" box="[483,494,1827,1847]" italics="true" pageId="4" pageNumber="5">1</emphasis>
, Φ 
<emphasis id="B909EAE49538B842FD86F88FC875F8B6" box="[516,527,1827,1847]" italics="true" pageId="4" pageNumber="5">2</emphasis>
1 Φ 1 Φ 1 Φ
</paragraph>
<paragraph id="8BC236F69538B842FF30F892CEBDF8EF" blockId="4.[174,1427,1568,1957]" pageId="4" pageNumber="5">
TARSUS 7 
<emphasis id="B909EAE49538B842FE5AF892CB91F8D3" box="[472,491,1854,1874]" italics="true" pageId="4" pageNumber="5">pl'</emphasis>
, 
<emphasis id="B909EAE49538B842FE73F892C872F8D3" box="[497,520,1854,1874]" italics="true" pageId="4" pageNumber="5">pl&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FD8CF892C85DF8D3" box="[526,551,1854,1874]" italics="true" pageId="4" pageNumber="5">pv'</emphasis>
, 
<emphasis id="B909EAE49538B842FDAEF892C832F8D3" box="[556,584,1854,1874]" italics="true" pageId="4" pageNumber="5">pv&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FDCCF892C827F8D3" box="[590,605,1854,1874]" italics="true" pageId="4" pageNumber="5">u'</emphasis>
, 
<emphasis id="B909EAE49538B842FDE0F892C80EF8D3" box="[610,628,1854,1874]" italics="true" pageId="4" pageNumber="5">u&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FDF8F892C8FEF8D3" box="[634,644,1854,1874]" italics="true" pageId="4" pageNumber="5">s</emphasis>
7 
<emphasis id="B909EAE49538B842FD5BF892C88BF8D3" box="[729,753,1854,1874]" italics="true" pageId="4" pageNumber="5">pl&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FD74F892C973F8D3" box="[758,777,1854,1874]" italics="true" pageId="4" pageNumber="5">tc'</emphasis>
, 
<emphasis id="B909EAE49538B842FC8DF892C95FF8D3" box="[783,805,1854,1874]" italics="true" pageId="4" pageNumber="5">tc&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FCA9F892C93EF8D3" box="[811,836,1854,1874]" italics="true" pageId="4" pageNumber="5">pv'</emphasis>
, 
<emphasis id="B909EAE49538B842FCCBF892C91FF8D3" box="[841,869,1854,1874]" italics="true" pageId="4" pageNumber="5">pv&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FCE9F892C900F8D3" box="[875,890,1854,1874]" italics="true" pageId="4" pageNumber="5">u'</emphasis>
, 
<emphasis id="B909EAE49538B842FC02F892C9EBF8D3" box="[896,913,1854,1874]" italics="true" pageId="4" pageNumber="5">u&quot;</emphasis>
7 
<emphasis id="B909EAE49538B842FC58F892C988F8D3" box="[986,1010,1854,1874]" italics="true" pageId="4" pageNumber="5">pl&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FC75F892CE70F8D3" box="[1015,1034,1854,1874]" italics="true" pageId="4" pageNumber="5">tc'</emphasis>
, 
<emphasis id="B909EAE49538B842FB92F892CE5CF8D3" box="[1040,1062,1854,1874]" italics="true" pageId="4" pageNumber="5">tc&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FBAEF892CE3FF8D3" box="[1068,1093,1854,1874]" italics="true" pageId="4" pageNumber="5">pv'</emphasis>
, 
<emphasis id="B909EAE49538B842FBC8F892CE1CF8D3" box="[1098,1126,1854,1874]" italics="true" pageId="4" pageNumber="5">pv&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FBEEF892CE01F8D3" box="[1132,1147,1854,1874]" italics="true" pageId="4" pageNumber="5">u'</emphasis>
, 
<emphasis id="B909EAE49538B842FB03F892CEE8F8D3" box="[1153,1170,1854,1874]" italics="true" pageId="4" pageNumber="5">u&quot;</emphasis>
6 
<emphasis id="B909EAE49538B842FB59F892CE89F8D3" box="[1243,1267,1854,1874]" italics="true" pageId="4" pageNumber="5">pl&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FB7AF892CF71F8D3" box="[1272,1291,1854,1874]" italics="true" pageId="4" pageNumber="5">tc'</emphasis>
, 
<emphasis id="B909EAE49538B842FA93F892CF5DF8D3" box="[1297,1319,1854,1874]" italics="true" pageId="4" pageNumber="5">tc&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FAAFF892CF3CF8D3" box="[1325,1350,1854,1874]" italics="true" pageId="4" pageNumber="5">pv'</emphasis>
, 
<emphasis id="B909EAE49538B842FAC9F892CF1DF8D3" box="[1355,1383,1854,1874]" italics="true" pageId="4" pageNumber="5">pv&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FAEFF892CF06F8D3" box="[1389,1404,1854,1874]" italics="true" pageId="4" pageNumber="5">u'</emphasis>
, 
<emphasis id="B909EAE49538B842FA00F892CFE9F8D3" box="[1410,1427,1854,1874]" italics="true" pageId="4" pageNumber="5">u&quot;</emphasis>
Eupathidia 6 
<emphasis id="B909EAE49538B842FE5AF8F6CB91F8EF" box="[472,491,1882,1902]" italics="true" pageId="4" pageNumber="5">tc'</emphasis>
, 
<emphasis id="B909EAE49538B842FE75F8F6C877F8EF" box="[503,525,1882,1902]" italics="true" pageId="4" pageNumber="5">tc&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FD9BF8F6C852F8EF" box="[537,552,1882,1902]" italics="true" pageId="4" pageNumber="5">p'</emphasis>
, 
<emphasis id="B909EAE49538B842FDB1F8F6C83FF8EF" box="[563,581,1882,1902]" italics="true" pageId="4" pageNumber="5">p&quot;</emphasis>
, 
<emphasis id="B909EAE49538B842FDD2F8F6C825F8EF" box="[592,607,1882,1902]" italics="true" pageId="4" pageNumber="5">ft'</emphasis>
, 
<emphasis id="B909EAE49538B842FDE8F8F6C806F8EF" box="[618,636,1882,1902]" italics="true" pageId="4" pageNumber="5">ft&quot;</emphasis>
0 0 0
</paragraph>
<paragraph id="8BC236F69538B842FF30F8D9CEBCF808" blockId="4.[174,1427,1568,1957]" box="[178,1222,1909,1929]" pageId="4" pageNumber="5">
Solenidia 2 ω 
<emphasis id="B909EAE49538B842FE64F8D9CB8BF808" box="[486,497,1909,1929]" italics="true" pageId="4" pageNumber="5">1</emphasis>
, ω 
<emphasis id="B909EAE49538B842FD92F8D9C861F808" box="[528,539,1909,1929]" italics="true" pageId="4" pageNumber="5">2</emphasis>
1 ω 1 ω 0
</paragraph>
<paragraph id="8BC236F69538B842FF30F83DCEBCF824" blockId="4.[174,1427,1568,1957]" box="[178,1222,1937,1957]" pageId="4" pageNumber="5">Claws 1 2 2 1</paragraph>
</caption>
<paragraph id="8BC236F69539B843FFFFFF04CBBEFF3E" blockId="5.[125,452,168,191]" box="[125,452,168,191]" pageId="5" pageNumber="6">
<bibCitation id="5E573E9A9539B843FFFFFF04CBBEFF3E" box="[125,452,168,191]" journal="Zoological Studies" page="11" pageId="5" pageNumber="6" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49539B843FFFFFF04CB4DFF3E" box="[125,311,168,191]" italics="true" pageId="5" pageNumber="6">Zoological Studies</emphasis>
<emphasis id="B909EAE49539B843FEBFFF04CB24FF3E" bold="true" box="[317,350,168,191]" pageId="5" pageNumber="6">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69539B843FF47FF51C89BFD77" blockId="5.[136,737,253,758]" pageId="5" pageNumber="6">
Idiosomal venter (
<figureCitation id="13462A739539B843FE21FF51C879FE97" box="[419,515,253,278]" captionStart="Fig" captionStartId="5.[136,170,1950,1970]" captionTargetBox="[151,1401,822,1921]" captionTargetId="figure-288@5.[136,1413,813,1934]" captionTargetPageId="5" captionText="Fig. 3. Zambedania sekhukhunensis sp. n. Female holotype. Idiosoma: A- dorsum, B- venter; sculpture omitted." figureDoi="http://doi.org/10.5281/zenodo.12956919" httpUri="https://zenodo.org/record/12956919/files/figure.png" pageId="5" pageNumber="6">Fig. 3B</figureCitation>
). All ventral setae short (at least half the size of dorsal opisthosomal setae) and slightly barbed. Ap. 1, 2 complete, well-sclerotized, originating at the anterior margin of coxisternal plates I and II respectively. Ap. 1 forming an Y-shaped juncture with prosternal apodeme (appr). Ap. 2 slightly oblique, originating from the anterior base of legs II, extending across plates I, II and uniting postero-medially with the appr. The appr is incomplete, not fused with the well developed and strongly sclerotized sejugal apodeme (apsej). Ap. 3, 4 complete, anteriorly arched, medially joined to the poststernal apodeme (appo). Distribution of coxisternal setae typical for the genus. Setae 
<emphasis id="B909EAE49539B843FE15FD11CBCFFD57" box="[407,437,701,726]" italics="true" pageId="5" pageNumber="6">2b</emphasis>
and 
<emphasis id="B909EAE49539B843FE7BFD11C86DFD57" box="[505,535,701,726]" italics="true" pageId="5" pageNumber="6">4b</emphasis>
are the longest coxisternal setae. Ps with three pairs of setae,
</paragraph>
<paragraph id="8BC236F69539B843FA8AFF04CFFFFF3E" blockId="5.[1288,1413,168,191]" box="[1288,1413,168,191]" pageId="5" pageNumber="6">page 6 of 21</paragraph>
<paragraph id="8BC236F69539B843FCAFFF51CF01FE97" blockId="5.[813,1415,253,758]" box="[813,1403,253,278]" pageId="5" pageNumber="6">
<emphasis id="B909EAE49539B843FCAFFF51C933FE97" box="[813,841,253,278]" italics="true" pageId="5" pageNumber="6">ps</emphasis>
<subScript id="17F934B39539B843FCCBFEAAC90FFE94" attach="left" box="[841,885,262,277]" fontSize="6" pageId="5" pageNumber="6">
<emphasis id="B909EAE49539B843FCCBFEAAC928FE94" box="[841,850,262,277]" italics="true" pageId="5" pageNumber="6">1</emphasis>
, 
<emphasis id="B909EAE49539B843FCD9FEAAC91EFE94" box="[859,868,262,277]" italics="true" pageId="5" pageNumber="6">2</emphasis>
, 
<emphasis id="B909EAE49539B843FCEEFEAAC90FFE94" box="[876,885,262,277]" italics="true" pageId="5" pageNumber="6">3</emphasis>
</subScript>
, of which 
<emphasis id="B909EAE49539B843FC6FFF51CE68FE94" box="[1005,1042,253,278]" italics="true" pageId="5" pageNumber="6">
ps 
<subScript id="17F934B39539B843FB8BFEAACE68FE94" attach="left" box="[1033,1042,262,277]" fontSize="6" pageId="5" pageNumber="6">2</subScript>
</emphasis>
is the longest pseudanal seta.
</paragraph>
<paragraph id="8BC236F69539B843FCEBFEB1C9C6FE57" blockId="5.[813,1415,253,758]" pageId="5" pageNumber="6">
Legs (
<figureCitation id="13462A739539B843FC36FEB1CE43FEB7" box="[948,1081,285,310]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="7.[136,170,1950,1970]" captionStartId-1="8.[174,209,1923,1943]" captionStartId-2="8.[850,885,1923,1943]" captionStartId-3="9.[136,170,1949,1969]" captionTargetBox-0="[129,1412,1054,1921]" captionTargetBox-1="[175,1451,842,1897]" captionTargetId-0="figure-344@7.[106,1433,1043,1973]" captionTargetId-1="figure-1@8.[748,1470,824,1909]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 5. Zambedania sekhukhunensis sp. n. Female holotype. Leg I A- (left): antaxial view, B- tibiotarsus (left): paraxial view." captionText-1="Fig. 6. Zambedania sekhukhunensis sp. n. Female holotype. Leg II (right): dorsal view." captionText-2="Fig. 7. Zambedania sekhukhunensis sp. n. Female holotype. Leg III (right): dorsal view." captionText-3="Fig. 8. Zambedania sekhukhunensis sp. n. Female holotype. Leg IV (right): paraxial view." figureDoi-0="http://doi.org/10.5281/zenodo.12956931" figureDoi-1="http://doi.org/10.5281/zenodo.12956941" httpUri-0="https://zenodo.org/record/12956931/files/figure.png" httpUri-1="https://zenodo.org/record/12956941/files/figure.png" pageId="5" pageNumber="6">Figs. 5 - 8</figureCitation>
). The first two pairs of legs are subequal in length; legs III, IV are distinctly longer. All tectal leg setae are setiform, tapering, distally acute and more or less barbed, pilose or one-sided barbed with long barbs (feathered, see 
<figureCitation id="13462A739539B843FCAFFE11C9CAFE57" box="[813,944,445,471]" captionStart="Fig" captionStartId="10.[174,208,1264,1284]" captionTargetBox="[176,1449,252,1233]" captionTargetId="figure-305@10.[174,1451,249,1235]" captionTargetPageId="10" captionText="Fig. 9. Zambedania sekhukhunensis sp. n. Female. A- dorsal view; B- caput of bothridial seta, dorsal view; in front “pilose” setae; C- Leg I (right). Note TiTa in dorsal view. At distal end “turret” carrying claw with at its base dorsal pinnaculum of the “locking system”, flanked by part of counterpiece. Eupathidia dorsad on pinnaculi. The cluster d and k associated with the three solenidia. Ventro-laterad near to counter piece seta s followed by feathered pv and v. D- detail distal end of left TiTa I in lateral view, claw and butterfly-shaped counter piece made up of setae u’ - u”, below seta s, followed by “feathered” primiventrals pv’ and pv” (photos by G. Bauchan and R. Ochoa)." figureDoi="http://doi.org/10.5281/zenodo.12956951" httpUri="https://zenodo.org/record/12956951/files/figure.png" pageId="5" pageNumber="6">Fig. 9 C, D</figureCitation>
).
</paragraph>
<paragraph id="8BC236F69539B843FCEBFE71CFFCFD77" blockId="5.[813,1415,253,758]" pageId="5" pageNumber="6">
Leg I. (
<figureCitation id="13462A739539B843FC44FE71CE26FE77" box="[966,1116,477,502]" captionStart="Fig" captionStartId="7.[136,170,1950,1970]" captionTargetBox="[129,1412,1054,1921]" captionTargetId="figure-344@7.[106,1433,1043,1973]" captionTargetPageId="7" captionText="Fig. 5. Zambedania sekhukhunensis sp. n. Female holotype. Leg I A- (left): antaxial view, B- tibiotarsus (left): paraxial view." figureDoi="http://doi.org/10.5281/zenodo.12956931" httpUri="https://zenodo.org/record/12956931/files/figure.png" pageId="5" pageNumber="6">Figs. 5A, B</figureCitation>
; 
<figureCitation id="13462A739539B843FBEEFE71CEC0FE77" box="[1132,1210,477,503]" captionStart="Fig" captionStartId="10.[174,208,1264,1284]" captionTargetBox="[176,1449,252,1233]" captionTargetId="figure-305@10.[174,1451,249,1235]" captionTargetPageId="10" captionText="Fig. 9. Zambedania sekhukhunensis sp. n. Female. A- dorsal view; B- caput of bothridial seta, dorsal view; in front “pilose” setae; C- Leg I (right). Note TiTa in dorsal view. At distal end “turret” carrying claw with at its base dorsal pinnaculum of the “locking system”, flanked by part of counterpiece. Eupathidia dorsad on pinnaculi. The cluster d and k associated with the three solenidia. Ventro-laterad near to counter piece seta s followed by feathered pv and v. D- detail distal end of left TiTa I in lateral view, claw and butterfly-shaped counter piece made up of setae u’ - u”, below seta s, followed by “feathered” primiventrals pv’ and pv” (photos by G. Bauchan and R. Ochoa)." figureDoi="http://doi.org/10.5281/zenodo.12956951" httpUri="https://zenodo.org/record/12956951/files/figure.png" pageId="5" pageNumber="6">9C, D</figureCitation>
). Tr: 
<emphasis id="B909EAE49539B843FA82FE71CF6EFE77" box="[1280,1300,477,502]" italics="true" pageId="5" pageNumber="6">v’</emphasis>
&lt;Fe 
<emphasis id="B909EAE49539B843FAE4FE71CF07FE77" box="[1382,1405,477,502]" italics="true" pageId="5" pageNumber="6">v”</emphasis>
; Fe: long 
<emphasis id="B909EAE49539B843FC23FE51C990FD97" box="[929,1002,509,534]" italics="true" pageId="5" pageNumber="6">l’&gt; v”</emphasis>
, 
<emphasis id="B909EAE49539B843FC7CFE51CE77FD97" box="[1022,1037,509,534]" italics="true" pageId="5" pageNumber="6">d</emphasis>
strongly barbed longest seta on segment; Ge: 
<emphasis id="B909EAE49539B843FB8BFDB1CE6FFDB7" box="[1033,1045,541,566]" italics="true" pageId="5" pageNumber="6">l’</emphasis>
strongly barbed, ~ Fe 
<emphasis id="B909EAE49539B843FAB5FDB1CF3CFDB7" box="[1335,1350,541,566]" italics="true" pageId="5" pageNumber="6">d</emphasis>
; 
<emphasis id="B909EAE49539B843FADBFDB1C90DFDD7" italics="true" pageId="5" pageNumber="6">v’&gt; l”&gt; v”</emphasis>
. TiTa with a long and strongly sclerotized, single, longitudinally ribbed apical claw closing into a butterfly-shaped counterpiece made up of the setae 
<emphasis id="B909EAE49539B843FCF4FD31C9F1FD37" box="[886,907,669,694]" italics="true" pageId="5" pageNumber="6">u’</emphasis>
and 
<emphasis id="B909EAE49539B843FC4BFD31C99AFD37" box="[969,992,669,694]" italics="true" pageId="5" pageNumber="6">u”</emphasis>
(
<figureCitation id="13462A739539B843FC72FD31CE37FD37" box="[1008,1101,669,694]" captionStart="Fig" captionStartId="10.[174,208,1264,1284]" captionTargetBox="[176,1449,252,1233]" captionTargetId="figure-305@10.[174,1451,249,1235]" captionTargetPageId="10" captionText="Fig. 9. Zambedania sekhukhunensis sp. n. Female. A- dorsal view; B- caput of bothridial seta, dorsal view; in front “pilose” setae; C- Leg I (right). Note TiTa in dorsal view. At distal end “turret” carrying claw with at its base dorsal pinnaculum of the “locking system”, flanked by part of counterpiece. Eupathidia dorsad on pinnaculi. The cluster d and k associated with the three solenidia. Ventro-laterad near to counter piece seta s followed by feathered pv and v. D- detail distal end of left TiTa I in lateral view, claw and butterfly-shaped counter piece made up of setae u’ - u”, below seta s, followed by “feathered” primiventrals pv’ and pv” (photos by G. Bauchan and R. Ochoa)." figureDoi="http://doi.org/10.5281/zenodo.12956951" httpUri="https://zenodo.org/record/12956951/files/figure.png" pageId="5" pageNumber="6">Fig. 9D</figureCitation>
). Claw with small auxiliary blade located inwards near its base, and similarly small blunt protrusion even more proximally at its
</paragraph>
<caption id="DF02667E9539B843FF0AF832CED6F833" ID-DOI="http://doi.org/10.5281/zenodo.12956919" ID-Zenodo-Dep="12956919" box="[136,1196,1950,1970]" httpUri="https://zenodo.org/record/12956919/files/figure.png" pageId="5" pageNumber="6" startId="5.[136,170,1950,1970]" targetBox="[151,1401,822,1921]" targetPageId="5" targetType="figure">
<paragraph id="8BC236F69539B843FF0AF832CED6F833" blockId="5.[136,1196,1950,1970]" box="[136,1196,1950,1970]" pageId="5" pageNumber="6">
<emphasis id="B909EAE49539B843FF0AF832CABCF833" bold="true" box="[136,198,1950,1970]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="4C7D4D759539B843FF50F832CB9DF833" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[210,487,1950,1970]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49539B843FF50F832CB9DF833" box="[210,487,1950,1970]" italics="true" pageId="5" pageNumber="6">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9539B843FE6FF832C858F833" box="[493,546,1950,1970]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
Female holotype. Idiosoma: A- dorsum, B- venter; sculpture omitted.
</paragraph>
</caption>
<paragraph id="8BC236F6953AB840FF2CFF04CB8EFF3E" blockId="6.[174,500,168,191]" box="[174,500,168,191]" pageId="6" pageNumber="7">
<bibCitation id="5E573E9A953AB840FF2CFF04CB8EFF3E" box="[174,500,168,191]" journal="Zoological Studies" page="11" pageId="6" pageNumber="7" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE4953AB840FF2CFF04CB1DFF3E" box="[174,359,168,191]" italics="true" pageId="6" pageNumber="7">Zoological Studies</emphasis>
<emphasis id="B909EAE4953AB840FEEFFF04CBF4FF3E" bold="true" box="[365,398,168,191]" pageId="6" pageNumber="7">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F6953AB840FF2CFF51C816FD37" blockId="6.[171,776,253,919]" pageId="6" pageNumber="7">
base. The “lock” on the dorsal side of tarsal part of the segment at the base of the flexed claw is well exposed in figure 9C. Barbed seta 
<emphasis id="B909EAE4953AB840FDCAFE91C82FFED7" box="[584,597,317,342]" italics="true" pageId="6" pageNumber="7">k</emphasis>
&lt;
<emphasis id="B909EAE4953AB840FDF6FE91C8F9FED7" box="[628,643,317,342]" italics="true" pageId="6" pageNumber="7">d</emphasis>
. Setae 
<emphasis id="B909EAE4953AB840FD5CFE91CABEFEF7" italics="true" pageId="6" pageNumber="7">v’ ~ v”</emphasis>
, both feathered; 
<emphasis id="B909EAE4953AB840FE0FFEF1CBBEFEF7" box="[397,452,349,374]" italics="true" pageId="6" pageNumber="7">l’ ~ l”</emphasis>
, both barbed; feathered 
<emphasis id="B909EAE4953AB840FD64FEF1CA96FE17" italics="true" pageId="6" pageNumber="7">pv’ &lt;pv”</emphasis>
typically brush-like barbed at distal end (
<figureCitation id="13462A73953AB840FD5BFED1CAA8FE37" captionStart="Fig" captionStartId="7.[136,170,1950,1970]" captionTargetBox="[129,1412,1054,1921]" captionTargetId="figure-344@7.[106,1433,1043,1973]" captionTargetPageId="7" captionText="Fig. 5. Zambedania sekhukhunensis sp. n. Female holotype. Leg I A- (left): antaxial view, B- tibiotarsus (left): paraxial view." figureDoi="http://doi.org/10.5281/zenodo.12956931" httpUri="https://zenodo.org/record/12956931/files/figure.png" pageId="6" pageNumber="7">Fig. 5B</figureCitation>
); 
<emphasis id="B909EAE4953AB840FF67FE31CB42FE37" box="[229,312,413,438]" italics="true" pageId="6" pageNumber="7">pl’ &lt;pl”</emphasis>
either barbed. Seta 
<emphasis id="B909EAE4953AB840FDA6FE31C84BFE37" box="[548,561,413,438]" italics="true" pageId="6" pageNumber="7">s</emphasis>
is situated anterolaterad of the claw, but difficult to observe with a light microscope because of the accumulation of setae there. However 
<emphasis id="B909EAE4953AB840FE3DFE51CBB6FD97" box="[447,460,509,534]" italics="true" pageId="6" pageNumber="7">s</emphasis>
is clearly visible in figure 9D and easily observed on the larva and male. Three striated solenidia: stemmed solenidion ω&gt; Φ 
<subScript id="17F934B3953AB840FF3EFDCACAA2FDF4" attach="right" box="[188,216,614,629]" fontSize="6" pageId="6" pageNumber="7">1, 2</subScript>
. Striation of Φ 
<emphasis id="B909EAE4953AB840FE02FDCACBF3FDF4" box="[384,393,614,629]" italics="true" pageId="6" pageNumber="7">
<subScript id="17F934B3953AB840FE02FDCACBF3FDF4" attach="left" box="[384,393,614,629]" fontSize="6" pageId="6" pageNumber="7">2</subScript>
</emphasis>
wider apart than both of Φ 
<emphasis id="B909EAE4953AB840FD4AFDCAC8ABFDF4" box="[712,721,614,629]" italics="true" pageId="6" pageNumber="7">
<subScript id="17F934B3953AB840FD4AFDCAC8ABFDF4" attach="left" box="[712,721,614,629]" fontSize="6" pageId="6" pageNumber="7">1</subScript>
</emphasis>
and ω. Eupathidia on pinnaculi (
<emphasis id="B909EAE4953AB840FE78FDD1C875FD17" box="[506,527,637,662]" italics="true" pageId="6" pageNumber="7">p’</emphasis>
missing): 
<emphasis id="B909EAE4953AB840FD08FDD1C8D8FD17" box="[650,674,637,662]" italics="true" pageId="6" pageNumber="7">p”</emphasis>
, 
<emphasis id="B909EAE4953AB840FD31FDD1C885FD17" box="[691,767,637,662]" italics="true" pageId="6" pageNumber="7">ft’ &lt;ft”</emphasis>
. Distalmost seta on the verticil 
<emphasis id="B909EAE4953AB840FD92FD31C81CFD37" box="[528,614,669,694]" italics="true" pageId="6" pageNumber="7">tc’ &lt;tc”</emphasis>
.
</paragraph>
<paragraph id="8BC236F6953AB840FF68FD11C97DFC17" blockId="6.[171,776,253,919]" pageId="6" pageNumber="7">
Leg II (
<figureCitation id="13462A73953AB840FECBFD11CBE1FD57" box="[329,411,701,726]" captionStart="Fig" captionStartId="8.[174,209,1923,1943]" captionTargetBox="[175,1451,842,1897]" captionTargetId="figure-1@8.[748,1470,824,1909]" captionTargetPageId="8" captionText="Fig. 6. Zambedania sekhukhunensis sp. n. Female holotype. Leg II (right): dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12956941" httpUri="https://zenodo.org/record/12956941/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
). Tr: barbed 
<emphasis id="B909EAE4953AB840FDC5FD11C821FD57" box="[583,603,701,726]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
&lt;barbed Fe 
<emphasis id="B909EAE4953AB840FF2CFD71CAB1FD77" box="[174,203,733,758]" italics="true" pageId="6" pageNumber="7">v”;</emphasis>
Fe: 
<emphasis id="B909EAE4953AB840FE87FD71CB33FD77" box="[261,329,733,758]" italics="true" pageId="6" pageNumber="7">l’&gt; v”</emphasis>
; 
<emphasis id="B909EAE4953AB840FEDBFD71CB12FD77" box="[345,360,733,758]" italics="true" pageId="6" pageNumber="7">d</emphasis>
longest seta of leg; Ge: 
<emphasis id="B909EAE4953AB840FD14FD71C97DFD77" box="[662,775,733,758]" italics="true" pageId="6" pageNumber="7">l’ ~ l” ~ v’</emphasis>
with strongly sclerotized annulus at its base; Ti: 
<emphasis id="B909EAE4953AB840FF2CFCB1CAC7FCB7" box="[174,189,797,822]" italics="true" pageId="6" pageNumber="7">d</emphasis>
shortest seta on segment, close to slim striated Φ, feathered 
<emphasis id="B909EAE4953AB840FECCFC91CBEAFCD7" box="[334,400,829,854]" italics="true" pageId="6" pageNumber="7">l’ &lt;v’</emphasis>
&lt;barbed 
<emphasis id="B909EAE4953AB840FD97FC91C856FCD7" box="[533,556,829,854]" italics="true" pageId="6" pageNumber="7">v”</emphasis>
; Ta: barbed 
<emphasis id="B909EAE4953AB840FD4CFC91C97CFCD7" box="[718,774,829,854]" italics="true" pageId="6" pageNumber="7">pl”&gt;</emphasis>
barbed 
<emphasis id="B909EAE4953AB840FE90FCF1CB55FCF7" box="[274,303,861,886]" italics="true" pageId="6" pageNumber="7">tc’</emphasis>
; 
<emphasis id="B909EAE4953AB840FEC0FCF1CB25FCF7" box="[322,351,861,886]" italics="true" pageId="6" pageNumber="7">tc’</emphasis>
&gt; smooth 
<emphasis id="B909EAE4953AB840FE6CFCF1C874FCF7" box="[494,526,861,886]" italics="true" pageId="6" pageNumber="7">tc”</emphasis>
; 
<emphasis id="B909EAE4953AB840FDA0FCF1C81AFCF7" box="[546,608,861,886]" italics="true" pageId="6" pageNumber="7">pv’ &lt;</emphasis>
barbed 
<emphasis id="B909EAE4953AB840FD4DFCF1C88CFCF7" box="[719,758,861,886]" italics="true" pageId="6" pageNumber="7">pv”</emphasis>
~ slim, smooth 
<emphasis id="B909EAE4953AB840FED6FCD1CB19FC17" box="[340,355,893,918]" italics="true" pageId="6" pageNumber="7">u</emphasis>
. Solenidion ω stout, club-shaped.
</paragraph>
<paragraph id="8BC236F6953AB840FAAFFF04CFD1FF3E" blockId="6.[1325,1451,168,191]" box="[1325,1451,168,191]" pageId="6" pageNumber="7">page 7 of 21</paragraph>
<paragraph id="8BC236F6953AB840FCD0FF51C9ADFEB7" blockId="6.[848,1452,253,919]" pageId="6" pageNumber="7">Terminal two simple claws with a fan-shaped empodium.</paragraph>
<paragraph id="8BC236F6953AB840FC0DFE91CFDBFD97" blockId="6.[848,1452,253,919]" pageId="6" pageNumber="7">
Leg III (
<figureCitation id="13462A73953AB840FC70FE91CE3AFED7" box="[1010,1088,317,342]" captionStart="Fig" captionStartId="8.[850,885,1923,1943]" captionText="Fig. 7. Zambedania sekhukhunensis sp. n. Female holotype. Leg III (right): dorsal view." pageId="6" pageNumber="7">Fig. 7</figureCitation>
). Tr: one-sidedly barbed 
<emphasis id="B909EAE4953AB840FAFCFE91CFE8FED7" box="[1406,1426,317,342]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
~ Fe 
<emphasis id="B909EAE4953AB840FCF9FEF1C9F4FEF7" box="[891,910,349,374]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
, Fe 
<emphasis id="B909EAE4953AB840FC4BFEF1C9A2FEF7" box="[969,984,349,374]" italics="true" pageId="6" pageNumber="7">d</emphasis>
longest seta on the leg; Ge: 
<emphasis id="B909EAE4953AB840FABFFEF1CF04FEF7" box="[1341,1406,349,374]" italics="true" pageId="6" pageNumber="7">l’&gt; v’</emphasis>
; Ti: Φ stout, club-shaped; slightly barbed 
<emphasis id="B909EAE4953AB840FA8FFED1CF4AFE17" box="[1293,1328,381,406]" italics="true" pageId="6" pageNumber="7">l’ ~</emphasis>
barbed 
<emphasis id="B909EAE4953AB840FA17FED1CFDEFE17" box="[1429,1444,381,406]" italics="true" pageId="6" pageNumber="7">d</emphasis>
; smooth 
<emphasis id="B909EAE4953AB840FC36FE31C9ADFE37" box="[948,983,413,438]" italics="true" pageId="6" pageNumber="7">v’&lt;</emphasis>
barbed 
<emphasis id="B909EAE4953AB840FBBCFE31CE2FFE37" box="[1086,1109,413,438]" italics="true" pageId="6" pageNumber="7">v”</emphasis>
; Ta: barbed 
<emphasis id="B909EAE4953AB840FB73FE31CF75FE37" box="[1265,1295,413,438]" italics="true" pageId="6" pageNumber="7">pl”</emphasis>
longest seta on verticil, barbed 
<emphasis id="B909EAE4953AB840FBB8FE11CE14FE57" box="[1082,1134,445,470]" italics="true" pageId="6" pageNumber="7">tc’ ~</emphasis>
smooth 
<emphasis id="B909EAE4953AB840FB5EFE11CE80FE57" box="[1244,1274,445,470]" italics="true" pageId="6" pageNumber="7">tc”</emphasis>
; smooth 
<emphasis id="B909EAE4953AB840FAF2FE11CFD1FE57" box="[1392,1451,445,470]" italics="true" pageId="6" pageNumber="7">pv’ &lt;</emphasis>
barbed 
<emphasis id="B909EAE4953AB840FC30FE71C99AFE77" box="[946,992,477,502]" italics="true" pageId="6" pageNumber="7">pv”,</emphasis>
distal-most barbed 
<emphasis id="B909EAE4953AB840FB59FE71CE90FE77" box="[1243,1258,477,502]" italics="true" pageId="6" pageNumber="7">u</emphasis>
~ 
<emphasis id="B909EAE4953AB840FA8CFE71CF40FE77" box="[1294,1338,477,502]" italics="true" pageId="6" pageNumber="7">pv’.</emphasis>
Terminal pair of simple claws with a fan-shaped empodium.
</paragraph>
<paragraph id="8BC236F6953AB840FC0DFDB1CE41FC97" blockId="6.[848,1452,253,919]" pageId="6" pageNumber="7">
Leg IV (
<figureCitation id="13462A73953AB840FC71FDB1CE3AFDB7" box="[1011,1088,541,566]" captionStart="Fig" captionStartId="9.[136,170,1949,1969]" captionText="Fig. 8. Zambedania sekhukhunensis sp. n. Female holotype. Leg IV (right): paraxial view." pageId="6" pageNumber="7">Fig. 8</figureCitation>
). Tr: 
<emphasis id="B909EAE4953AB840FB00FDB1CEECFDB7" box="[1154,1174,541,566]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
~ Fe 
<emphasis id="B909EAE4953AB840FB61FDB1CE8DFDB7" box="[1251,1271,541,566]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
, each barbed; Fe: robust, barbed 
<emphasis id="B909EAE4953AB840FBD1FD91CE18FDD7" box="[1107,1122,573,598]" italics="true" pageId="6" pageNumber="7">d</emphasis>
with strongly sclerotized annulus, longest seta on leg; Ge: barbed 
<emphasis id="B909EAE4953AB840FAFFFDF1CFEBFDF7" box="[1405,1425,605,630]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
&lt;Ti 
<emphasis id="B909EAE4953AB840FCF0FDD1C9FCFD17" box="[882,902,637,662]" italics="true" pageId="6" pageNumber="7">v’</emphasis>
; Ti: smooth 
<emphasis id="B909EAE4953AB840FBA1FDD1CEE5FD17" box="[1059,1183,637,662]" italics="true" pageId="6" pageNumber="7">l’ ~ v’ ~ v”</emphasis>
, both barbed; 
<emphasis id="B909EAE4953AB840FAD5FDD1CF1CFD17" box="[1367,1382,637,662]" italics="true" pageId="6" pageNumber="7">d</emphasis>
long, barbed with strongly sclerotized annulus and tiny solenidion Φ in sacculus at its base. Ta: 
<emphasis id="B909EAE4953AB840FAB5FD11CF2FFD57" box="[1335,1365,701,726]" italics="true" pageId="6" pageNumber="7">pl”</emphasis>
pilose; whipped 
<emphasis id="B909EAE4953AB840FC41FD71C98DFD77" box="[963,1015,733,758]" italics="true" pageId="6" pageNumber="7">tc’&gt;</emphasis>
smooth 
<emphasis id="B909EAE4953AB840FBE4FD71CEFEFD77" box="[1126,1156,733,758]" italics="true" pageId="6" pageNumber="7">tc”</emphasis>
, 
<emphasis id="B909EAE4953AB840FB15FD71CEC7FD77" box="[1175,1213,733,758]" italics="true" pageId="6" pageNumber="7">pv”</emphasis>
~ 
<emphasis id="B909EAE4953AB840FB63FD71CF7EFD77" box="[1249,1284,733,758]" italics="true" pageId="6" pageNumber="7">pv’</emphasis>
both barbed; and 
<emphasis id="B909EAE4953AB840FC04FD51C9C4FC97" box="[902,958,765,790]" italics="true" pageId="6" pageNumber="7">pv’&gt;</emphasis>
barbed 
<emphasis id="B909EAE4953AB840FBA2FD51CE4FFC97" box="[1056,1077,765,790]" italics="true" pageId="6" pageNumber="7">u’</emphasis>
.
</paragraph>
<paragraph id="8BC236F6953AB840FC0DFCB1CF65FC17" blockId="6.[848,1452,253,919]" pageId="6" pageNumber="7">
<emphasis id="B909EAE4953AB840FC0DFCB1CE3AFCB7" box="[911,1088,797,822]" italics="true" pageId="6" pageNumber="7">Etymology</emphasis>
: The specific name 
<taxonomicName id="4C7D4D75953AB840FCD0FC91CE66FCD7" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[850,1052,829,854]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sekhukhunensis">
<emphasis id="B909EAE4953AB840FCD0FC91CE66FCD7" box="[850,1052,829,854]" italics="true" pageId="6" pageNumber="7">sekhukhunensis</emphasis>
</taxonomicName>
refers to the collecting locality of the spider hosts, Sekhukhuneland, near Lydenburg, 
<collectingRegion id="49B9F814953AB840FC5CFCD1CE07FC17" box="[990,1149,893,918]" country="South Africa" name="Mpumalanga" pageId="6" pageNumber="7">Mpumalanga</collectingRegion>
, 
<collectingCountry id="F36A7666953AB840FB05FCD1CF66FC17" box="[1159,1308,893,919]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
.
</paragraph>
<paragraph id="8BC236F6953AB840FF64FC4DCB78FC74" blockId="6.[230,258,993,1013]" box="[230,258,993,1013]" pageId="6" pageNumber="7">(A)</paragraph>
<paragraph id="8BC236F6953AB840FCC1FC4DC925FC74" blockId="6.[835,863,993,1013]" box="[835,863,993,1013]" pageId="6" pageNumber="7">(B)</paragraph>
<caption id="DF02667E953AB840FF2CF832CF62F833" ID-DOI="http://doi.org/10.5281/zenodo.12956925" ID-Zenodo-Dep="12956925" box="[174,1304,1950,1970]" httpUri="https://zenodo.org/record/12956925/files/figure.png" pageId="6" pageNumber="7" startId="6.[174,208,1950,1970]" targetBox="[251,1357,1008,1915]" targetPageId="6" targetType="figure">
<paragraph id="8BC236F6953AB840FF2CF832CF62F833" blockId="6.[174,1304,1950,1970]" box="[174,1304,1950,1970]" pageId="6" pageNumber="7">
<emphasis id="B909EAE4953AB840FF2CF832CA91F833" bold="true" box="[174,235,1950,1970]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
<taxonomicName id="4C7D4D75953AB840FF75F832C877F833" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[247,525,1950,1970]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE4953AB840FF75F832C877F833" box="[247,525,1950,1970]" italics="true" pageId="6" pageNumber="7">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F953AB840FD91F832C83DF833" box="[531,583,1950,1970]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
Female holotype. Gnathosoma A- dorsum, B- venter, C- pharyngeal pumps.
</paragraph>
</caption>
<paragraph id="8BC236F6953BB841FFFFFF04CBBEFF3E" blockId="7.[125,452,168,191]" box="[125,452,168,191]" pageId="7" pageNumber="8">
<bibCitation id="5E573E9A953BB841FFFFFF04CBBEFF3E" box="[125,452,168,191]" journal="Zoological Studies" page="11" pageId="7" pageNumber="8" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE4953BB841FFFFFF04CB4DFF3E" box="[125,311,168,191]" italics="true" pageId="7" pageNumber="8">Zoological Studies</emphasis>
<emphasis id="B909EAE4953BB841FEBFFF04CB24FF3E" bold="true" box="[317,350,168,191]" pageId="7" pageNumber="8">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F6953BB841FF47FF51C873FE97" blockId="7.[136,738,253,982]" box="[197,521,253,278]" pageId="7" pageNumber="8">
Male (
<figureCitation id="13462A73953BB841FE8FFF51CBE2FE97" box="[269,408,253,278]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="11.[136,170,1159,1179]" captionStartId-1="11.[136,170,1951,1971]" captionStartId-2="12.[174,209,1925,1945]" captionStartId-3="12.[850,885,1925,1945]" captionStartId-4="13.[136,171,1926,1946]" captionStartId-5="13.[813,848,1926,1946]" captionTargetBox-0="[134,1414,249,1131]" captionTargetBox-1="[137,1416,1266,1881]" captionTargetBox-3="[102,1452,779,1899]" captionTargetBox-4="[138,736,432,1900]" captionTargetBox-5="[814,1413,638,1898]" captionTargetId-0="figure-45@11.[121,1430,242,1139]" captionTargetId-1="figure-51@11.[123,1429,1248,1962]" captionTargetId-3="figure-224@12.[87,907,775,1912]" captionTargetId-4="figure-121@13.[125,740,424,1922]" captionTargetId-5="figure-135@13.[802,1450,617,1912]" captionTargetPageId-0="11" captionTargetPageId-1="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionTargetPageId-5="13" captionText-0="Fig. 10. Zambedania sekhukhunensis sp. n. Male paratype. Idiosoma: A- dorsum, B- venter; gnathosoma: C- dorsum, D- venter; ascalebar for idiosoma, b- scalebar for gnathosoma; sculpture omitted." captionText-1="Fig. 11. Zambedania sekhukhunensis sp. n. Male paratype. Genital capsule: A- dorsal view, B. ventral view." captionText-2="Fig. 12. Zambedania sekhukhunensis sp. n. Male paratype. Leg I (right): dorsal view." captionText-3="Fig. 13. Zambedania sekhukhunensis sp. n. Male paratype. Leg II (right): dorsal view." captionText-4="Fig. 14. Zambedania sekhukhunensis sp. n. Male paratype. Leg III (right): dorsal view." captionText-5="Fig. 15. Zambedania sekhukhunensis sp. n. Male paratype. Leg IV (left): dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.12956957" figureDoi-1="http://doi.org/10.5281/zenodo.12956967" figureDoi-3="http://doi.org/10.5281/zenodo.12956973" figureDoi-4="http://doi.org/10.5281/zenodo.12956981" figureDoi-5="http://doi.org/10.5281/zenodo.12956989" httpUri-0="https://zenodo.org/record/12956957/files/figure.png" httpUri-1="https://zenodo.org/record/12956967/files/figure.png" httpUri-3="https://zenodo.org/record/12956973/files/figure.png" httpUri-4="https://zenodo.org/record/12956981/files/figure.png" httpUri-5="https://zenodo.org/record/12956989/files/figure.png" pageId="7" pageNumber="8">Figs. 10-15</figureCitation>
, 
<tableCitation id="C6FF034D953BB841FE20FF51CB86FE97" box="[418,508,253,278]" captionStart="Table 3" captionStartId="4.[174,242,1568,1593]" captionText="Table 3. Leg chaeto- and solenidiotaxy of the genus Zambedania sekhukhunensis sp. n. males. SEGMENT LEG I LEG II LEG III LEG IV SEGMENT Nr SIGLA Nr SIGLA Nr SIGLA Nr SIGLA TROCHANTER 1 v' 1 v' 1 v' 1 v' FEMUR 3 d,l',v&quot; 3 d,l',v&quot; 2 d,v' 2 d,v&quot; GENU 4 l',l&quot;,v', v&quot; 3 l',l&quot;,v&quot; 2 l',v&quot; 1 v' TIBIA 6 l',l&quot;,v',v&quot;,k,d 4 d,l',v',v&quot; 4 d,l',v',v&quot; 4 d,l',v',v&quot; Solenidia 2 Φ 1, Φ 2 1 Φ 1 Φ 1 Φ TARSUS 7 pl',pl&quot;,pv',pv&quot;,u',u&quot;,s 7 pl&quot;,tc',tc&quot;,pv',pv&quot;,u',u&quot; 7 pl&quot;,tc',tc&quot;,pv',pv&quot;,u',u&quot; 6 pl&quot;,tc',tc&quot;,pv',pv&quot;,u',u&quot; Eupathidia 6 tc', tc&quot;, p', p&quot;, ft', ft&quot; 0 0 0 Solenidia 2 ω1, ω2 1 ω 1 ω 0 Claws 1 2 2 1" pageId="7" pageNumber="8">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8BC236F6953BB841FF47FEB1C856FD57" blockId="7.[136,738,253,982]" pageId="7" pageNumber="8">
Gnathosoma (
<figureCitation id="13462A73953BB841FEFFFEB1C866FEB7" box="[381,540,285,311]" captionStart="Fig" captionStartId="11.[136,170,1159,1179]" captionTargetBox="[134,1414,249,1131]" captionTargetId="figure-45@11.[121,1430,242,1139]" captionTargetPageId="11" captionText="Fig. 10. Zambedania sekhukhunensis sp. n. Male paratype. Idiosoma: A- dorsum, B- venter; gnathosoma: C- dorsum, D- venter; ascalebar for idiosoma, b- scalebar for gnathosoma; sculpture omitted." figureDoi="http://doi.org/10.5281/zenodo.12956957" httpUri="https://zenodo.org/record/12956957/files/figure.png" pageId="7" pageNumber="8">Fig. 10C, D</figureCitation>
). Gnathosomal capsule of the male much smaller than that of female. However, most of the external structures of the female capsule present on the male gnathosoma except that the mouthparts are absent. The stylophore extends anteriorly into a truncated-triangular distal edge bearing two pairs of tiny, sub-equal in size, smooth cheliceral setae: 
<emphasis id="B909EAE4953BB841FF0AFDB1CACEFDB7" box="[136,180,541,566]" italics="true" pageId="7" pageNumber="8">ch1</emphasis>
and 
<emphasis id="B909EAE4953BB841FF7BFDB1CB5FFDB7" box="[249,293,541,566]" italics="true" pageId="7" pageNumber="8">ch2</emphasis>
. Unlike the female, the male lacks palpi, palpal supracoxal setae (
<emphasis id="B909EAE4953BB841FE7EFD91C860FDD7" box="[508,538,573,598]" italics="true" pageId="7" pageNumber="8">pp</emphasis>
) and chelicerae. Subcapitulum with a pair of relatively long slender, simple setae (
<emphasis id="B909EAE4953BB841FEB4FDD1CB28FD17" box="[310,338,637,662]" italics="true" pageId="7" pageNumber="8">su</emphasis>
); anteriad setigenous accessory structures (
<emphasis id="B909EAE4953BB841FE93FD31CB41FD37" box="[273,315,669,694]" italics="true" pageId="7" pageNumber="8">ass</emphasis>
) laterally flanked by relatively long clavate, non-striated solenidia (sol).
</paragraph>
<paragraph id="8BC236F6953BB841FF47FD71C875FD77" blockId="7.[136,738,253,982]" box="[197,527,733,758]" pageId="7" pageNumber="8">Pharyngeal system: absent.</paragraph>
<paragraph id="8BC236F6953BB841FF47FD51CFFFFF3E" blockId="7.[136,738,253,982]" lastBlockId="7.[1288,1413,168,191]" pageId="7" pageNumber="8">
Idiosomal dorsum (
<figureCitation id="13462A73953BB841FE3FFD51C842FC97" box="[445,568,765,790]" captionStart="Fig" captionStartId="11.[136,170,1159,1179]" captionTargetBox="[134,1414,249,1131]" captionTargetId="figure-45@11.[121,1430,242,1139]" captionTargetPageId="11" captionText="Fig. 10. Zambedania sekhukhunensis sp. n. Male paratype. Idiosoma: A- dorsum, B- venter; gnathosoma: C- dorsum, D- venter; ascalebar for idiosoma, b- scalebar for gnathosoma; sculpture omitted." figureDoi="http://doi.org/10.5281/zenodo.12956957" httpUri="https://zenodo.org/record/12956957/files/figure.png" pageId="7" pageNumber="8">Figs 10A</figureCitation>
, 
<figureCitation id="13462A73953BB841FDCAFD51C804FC97" box="[584,638,765,790]" captionStart="Fig" captionStartId="11.[136,170,1951,1971]" captionTargetBox="[137,1416,1266,1881]" captionTargetId="figure-51@11.[123,1429,1248,1962]" captionTargetPageId="11" captionText="Fig. 11. Zambedania sekhukhunensis sp. n. Male paratype. Genital capsule: A- dorsal view, B. ventral view." figureDoi="http://doi.org/10.5281/zenodo.12956967" httpUri="https://zenodo.org/record/12956967/files/figure.png" pageId="7" pageNumber="8">11A</figureCitation>
). Males on average much smaller than females (range of lengths 172-186 vs 270-309) and with well sclerotized, sparsely punctate exoskeleton. Prodorsal shield (PdS) triangular, dome-shaped, without stigmata, tracheal system or bothridia; verticals (
<emphasis id="B909EAE4953BB841FE89FC11CB52FC57" box="[267,296,957,982]" italics="true" pageId="7" pageNumber="8">v2</emphasis>
) antero-mediad of setiform short page 8 of 21
</paragraph>
<paragraph id="8BC236F6953BB841FCAFFF51CEEDFD77" blockId="7.[813,1415,253,982]" pageId="7" pageNumber="8">
scapulars 
<emphasis id="B909EAE4953BB841FC33FF51C9A7FE97" box="[945,989,253,278]" italics="true" pageId="7" pageNumber="8">sc1</emphasis>
and long 
<emphasis id="B909EAE4953BB841FBE1FF51CEF4FE97" box="[1123,1166,253,278]" italics="true" pageId="7" pageNumber="8">sc2</emphasis>
. Opisthosoma with tergites CD fused, EF fused, and dorsodistally partly covers greatly reduced tergite H which coalesced with caudoventral pseudanal shield (Ps) to form HPs, the genital capsule. All dorsal setae barbed, except for smooth setae 
<emphasis id="B909EAE4953BB841FA92FE31CF65FE37" box="[1296,1311,413,438]" italics="true" pageId="7" pageNumber="8">e</emphasis>
, 
<emphasis id="B909EAE4953BB841FAB3FE31CF35FE37" box="[1329,1359,413,438]" italics="true" pageId="7" pageNumber="8">h1</emphasis>
and 
<emphasis id="B909EAE4953BB841FCAFFE11C930FE57" box="[813,842,445,470]" italics="true" pageId="7" pageNumber="8">h2</emphasis>
. Fused shield CD bearing two pairs of very long tapered, setiform barbed setae (
<emphasis id="B909EAE4953BB841FB3AFE71CEAFFE77" box="[1208,1237,477,502]" italics="true" pageId="7" pageNumber="8">c1</emphasis>
, 
<emphasis id="B909EAE4953BB841FB64FE71CF79FE77" box="[1254,1283,477,502]" italics="true" pageId="7" pageNumber="8">c2</emphasis>
) anteriorly and a pair (
<emphasis id="B909EAE4953BB841FC41FE51C9A8FD97" box="[963,978,509,534]" italics="true" pageId="7" pageNumber="8">d</emphasis>
) of similar setae postero-laterally. Shield EF with setae 
<emphasis id="B909EAE4953BB841FBBBFDB1CE32FDB7" box="[1081,1096,541,566]" italics="true" pageId="7" pageNumber="8">e</emphasis>
less than half as long as barbed 
<emphasis id="B909EAE4953BB841FC0EFD91C9E9FDD7" box="[908,915,573,598]" italics="true" pageId="7" pageNumber="8">f</emphasis>
, covering HPs. Shield H (
<figureCitation id="13462A73953BB841FB5BFD91CF39FDD7" box="[1241,1347,573,598]" captionStart="Fig" captionStartId="11.[136,170,1951,1971]" captionTargetBox="[137,1416,1266,1881]" captionTargetId="figure-51@11.[123,1429,1248,1962]" captionTargetPageId="11" captionText="Fig. 11. Zambedania sekhukhunensis sp. n. Male paratype. Genital capsule: A- dorsal view, B. ventral view." figureDoi="http://doi.org/10.5281/zenodo.12956967" httpUri="https://zenodo.org/record/12956967/files/figure.png" pageId="7" pageNumber="8">Fig. 11A</figureCitation>
) with two pairs of setae: anteriormost setae (
<emphasis id="B909EAE4953BB841FB7BFDF1CF6DFDF7" box="[1273,1303,605,630]" italics="true" pageId="7" pageNumber="8">h1</emphasis>
) smooth, short, thorny and pointed, 
<emphasis id="B909EAE4953BB841FB18FDD1CEC0FD17" box="[1178,1210,637,662]" italics="true" pageId="7" pageNumber="8">h2</emphasis>
setae strongly modified into button-like structures in suckershaped rings. Membranous copulatory flanges border the H shield posteriorly.
</paragraph>
<paragraph id="8BC236F6953BB841FCEBFD51CFFFFC57" blockId="7.[813,1415,253,982]" pageId="7" pageNumber="8">
Idiosomal venter (
<figureCitation id="13462A73953BB841FBC3FD51CEC9FC97" box="[1089,1203,765,790]" captionStart="Fig" captionStartId="11.[136,170,1159,1179]" captionTargetBox="[134,1414,249,1131]" captionTargetId="figure-45@11.[121,1430,242,1139]" captionTargetPageId="11" captionText="Fig. 10. Zambedania sekhukhunensis sp. n. Male paratype. Idiosoma: A- dorsum, B- venter; gnathosoma: C- dorsum, D- venter; ascalebar for idiosoma, b- scalebar for gnathosoma; sculpture omitted." figureDoi="http://doi.org/10.5281/zenodo.12956957" httpUri="https://zenodo.org/record/12956957/files/figure.png" pageId="7" pageNumber="8">Figs 10B</figureCitation>
, 
<figureCitation id="13462A73953BB841FB42FD51CE88FC97" box="[1216,1266,765,790]" captionStart="Fig" captionStartId="11.[136,170,1951,1971]" captionTargetBox="[137,1416,1266,1881]" captionTargetId="figure-51@11.[123,1429,1248,1962]" captionTargetPageId="11" captionText="Fig. 11. Zambedania sekhukhunensis sp. n. Male paratype. Genital capsule: A- dorsal view, B. ventral view." figureDoi="http://doi.org/10.5281/zenodo.12956967" httpUri="https://zenodo.org/record/12956967/files/figure.png" pageId="7" pageNumber="8">11B</figureCitation>
). Compared with the long clearly barbed dorsal setae the ventral setae short, smooth or more or less barbed. Coxisternal plates: anterior and posterior edges of ventral shields smooth, not subdivided, and widely curved. The setae are distributed as in the females. Ap1 very short, seemingly a mere widening of the
</paragraph>
<caption id="DF02667E953BB841FF0AF832CF6CF833" ID-DOI="http://doi.org/10.5281/zenodo.12956931" ID-Zenodo-Dep="12956931" box="[136,1302,1950,1970]" httpUri="https://zenodo.org/record/12956931/files/figure.png" pageId="7" pageNumber="8" startId="7.[136,170,1950,1970]" targetBox="[129,1412,1054,1921]" targetPageId="7" targetType="figure">
<paragraph id="8BC236F6953BB841FF0AF832CF6CF833" blockId="7.[136,1302,1950,1970]" box="[136,1302,1950,1970]" pageId="7" pageNumber="8">
<emphasis id="B909EAE4953BB841FF0AF832CABCF833" bold="true" box="[136,198,1950,1970]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
<taxonomicName id="4C7D4D75953BB841FF50F832CB9DF833" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[210,487,1950,1970]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE4953BB841FF50F832CB9DF833" box="[210,487,1950,1970]" italics="true" pageId="7" pageNumber="8">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F953BB841FE6FF832C858F833" box="[493,546,1950,1970]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
Female holotype. Leg I A- (left): antaxial view, B- tibiotarsus (left): paraxial view.
</paragraph>
</caption>
<paragraph id="8BC236F69534B84EFF2CFF04CB8EFF3E" blockId="8.[174,500,168,191]" box="[174,500,168,191]" pageId="8" pageNumber="9">
<bibCitation id="5E573E9A9534B84EFF2CFF04CB8EFF3E" box="[174,500,168,191]" journal="Zoological Studies" page="11" pageId="8" pageNumber="9" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49534B84EFF2CFF04CB1DFF3E" box="[174,359,168,191]" italics="true" pageId="8" pageNumber="9">Zoological Studies</emphasis>
<emphasis id="B909EAE49534B84EFEEFFF04CBF4FF3E" bold="true" box="[365,398,168,191]" pageId="8" pageNumber="9">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69534B84EFF2CFF51C807FD57" blockId="8.[174,775,253,790]" pageId="8" pageNumber="9">
anterior part of the prosternal apodeme (appr). The appr extends uninterruptedly caudad to form a Y-shaped juncture with complete ap2, to continue posteriorly towards the apsej. Ap3, 4 complete, anteriorly arched; ap3 medially concavely and ap4 convexly joined to completely formed poststernal apodeme (appo). Appo meeting posteriorly of setae 
<emphasis id="B909EAE49534B84EFF7EFE71CB61FE77" box="[252,283,477,502]" italics="true" pageId="8" pageNumber="9">4a</emphasis>
with ap5 from the posterior condyle of Tr IV. Setae 
<emphasis id="B909EAE49534B84EFECAFE51CB1CFD97" box="[328,358,509,534]" italics="true" pageId="8" pageNumber="9">4b</emphasis>
proximal-most on coalesced plate HPs. On the genital capsule modified pseudanal setae 
<emphasis id="B909EAE49534B84EFE82FD91CB57FDD7" box="[256,301,573,598]" italics="true" pageId="8" pageNumber="9">ps2</emphasis>
on a slight elevation, medio-laterad of and very close to 
<emphasis id="B909EAE49534B84EFE39FDF1CB92FDF7" box="[443,488,605,630]" italics="true" pageId="8" pageNumber="9">ps3</emphasis>
. Setae 
<emphasis id="B909EAE49534B84EFDD3FDF1C804FDF7" box="[593,638,605,630]" italics="true" pageId="8" pageNumber="9">ps1</emphasis>
(
<figureCitation id="13462A739534B84EFD13FDF1C885FDF7" box="[657,767,605,630]" captionStart="Fig" captionStartId="11.[136,170,1951,1971]" captionTargetBox="[137,1416,1266,1881]" captionTargetId="figure-51@11.[123,1429,1248,1962]" captionTargetPageId="11" captionText="Fig. 11. Zambedania sekhukhunensis sp. n. Male paratype. Genital capsule: A- dorsal view, B. ventral view." figureDoi="http://doi.org/10.5281/zenodo.12956967" httpUri="https://zenodo.org/record/12956967/files/figure.png" pageId="8" pageNumber="9">Fig. 11A</figureCitation>
) probably implanted medio-anteriorly of 
<emphasis id="B909EAE49534B84EFD07FDD1C8CAFD17" box="[645,688,637,662]" italics="true" pageId="8" pageNumber="9">ps2</emphasis>
, 
<emphasis id="B909EAE49534B84EFD40FDD1C8ABFD17" box="[706,721,637,662]" italics="true" pageId="8" pageNumber="9">3</emphasis>
and covered by copulatory flanges, on either side of the folded aedeagus. No cupules seen.
</paragraph>
<paragraph id="8BC236F69534B84EFF68FD71C97CFC97" blockId="8.[174,775,253,790]" pageId="8" pageNumber="9">
Legs. Leg I (
<figureCitation id="13462A739534B84EFE10FD71CB8FFD77" box="[402,501,733,758]" captionStart="Fig" captionStartId="12.[174,209,1925,1945]" captionText="Fig. 12. Zambedania sekhukhunensis sp. n. Male paratype. Leg I (right): dorsal view." pageId="8" pageNumber="9">Fig. 12</figureCitation>
). All tactile leg setae setiform, tapering, distally acute and barbed.
</paragraph>
<paragraph id="8BC236F69534B84EFAAFFF04CFD1FF3E" blockId="8.[1325,1451,168,191]" box="[1325,1451,168,191]" pageId="8" pageNumber="9">page 9 of 21</paragraph>
<paragraph id="8BC236F69534B84EFCD0FF51CF4FFDD7" blockId="8.[848,1452,253,790]" pageId="8" pageNumber="9">
Trochanter length 
<emphasis id="B909EAE49534B84EFBB9FF51CE35FE97" box="[1083,1103,253,278]" italics="true" pageId="8" pageNumber="9">v’</emphasis>
~ Fe 
<emphasis id="B909EAE49534B84EFB22FF51CECDFE97" box="[1184,1207,253,278]" italics="true" pageId="8" pageNumber="9">v”</emphasis>
; Fe 
<emphasis id="B909EAE49534B84EFB77FF51CF7EFE97" box="[1269,1284,253,278]" italics="true" pageId="8" pageNumber="9">d</emphasis>
setiform and longest seta of leg I; 
<emphasis id="B909EAE49534B84EFBC9FEB1CE2DFEB7" box="[1099,1111,285,310]" italics="true" pageId="8" pageNumber="9">l’</emphasis>
&gt; ½ 
<emphasis id="B909EAE49534B84EFB16FEB1CED9FEB7" box="[1172,1187,285,310]" italics="true" pageId="8" pageNumber="9">d</emphasis>
, 
<emphasis id="B909EAE49534B84EFB30FEB1CEB3FEB7" box="[1202,1225,285,310]" italics="true" pageId="8" pageNumber="9">v”</emphasis>
~ ⅔ 
<emphasis id="B909EAE49534B84EFA84FEB1CF68FEB7" box="[1286,1298,285,310]" italics="true" pageId="8" pageNumber="9">l’</emphasis>
. Genu 
<emphasis id="B909EAE49534B84EFAE8FEB1CF07FEB7" box="[1386,1405,285,310]" italics="true" pageId="8" pageNumber="9">v’</emphasis>
~ 
<emphasis id="B909EAE49534B84EFA1EFEB1CFD1FEB7" box="[1436,1451,285,310]" italics="true" pageId="8" pageNumber="9">l”</emphasis>
&gt; 
<emphasis id="B909EAE49534B84EFCEBFE91C9FAFED7" box="[873,896,317,342]" italics="true" pageId="8" pageNumber="9">v”</emphasis>
, 
<emphasis id="B909EAE49534B84EFC0CFE91C9E0FED7" box="[910,922,317,342]" italics="true" pageId="8" pageNumber="9">l’</emphasis>
longest seta of verticil. All tibial setae barbed on one-side, except smooth 
<emphasis id="B909EAE49534B84EFB2AFEF1CECEFEF7" box="[1192,1204,349,374]" italics="true" pageId="8" pageNumber="9">l’</emphasis>
. Setae 
<emphasis id="B909EAE49534B84EFA91FEF1CF2FFEF7" box="[1299,1365,349,374]" italics="true" pageId="8" pageNumber="9">v’ ~ l”</emphasis>
&lt;
<emphasis id="B909EAE49534B84EFAF5FEF1CFFEFEF7" box="[1399,1412,349,374]" italics="true" pageId="8" pageNumber="9">k</emphasis>
; 
<emphasis id="B909EAE49534B84EFA17FEF1CFD6FEF7" box="[1429,1452,349,374]" italics="true" pageId="8" pageNumber="9">v”</emphasis>
longest, 
<emphasis id="B909EAE49534B84EFC3BFED1C9B2FE17" box="[953,968,381,406]" italics="true" pageId="8" pageNumber="9">d</emphasis>
shortest seta on verticil. Solenidion slim Φ 1&gt; Φ2. Tarsal solenidion ω 2 proximal-most on the verticil, close to and much shorter and thinner than robust ω 1. One-sidedly barbed 
<emphasis id="B909EAE49534B84EFA92FE71CF4CFE77" box="[1296,1334,477,502]" italics="true" pageId="8" pageNumber="9">pv”</emphasis>
~ 
<emphasis id="B909EAE49534B84EFADBFE71CF01FE77" box="[1369,1403,477,502]" italics="true" pageId="8" pageNumber="9">pv’</emphasis>
, 
<emphasis id="B909EAE49534B84EFA0FFE71CFD6FE77" box="[1421,1452,477,502]" italics="true" pageId="8" pageNumber="9">pl”</emphasis>
~1½ 
<emphasis id="B909EAE49534B84EFC11FE51C9D5FD97" box="[915,943,509,534]" italics="true" pageId="8" pageNumber="9">pl’</emphasis>
. Seta 
<emphasis id="B909EAE49534B84EFB85FE51CE6EFD97" box="[1031,1044,509,534]" italics="true" pageId="8" pageNumber="9">s</emphasis>
~ ω 1. Setae 
<emphasis id="B909EAE49534B84EFB40FE51CEADFD97" box="[1218,1239,509,534]" italics="true" pageId="8" pageNumber="9">u’</emphasis>
&lt;
<emphasis id="B909EAE49534B84EFB7EFE51CF6EFD97" box="[1276,1300,509,534]" italics="true" pageId="8" pageNumber="9">u”</emphasis>
. Eupathidia (
<emphasis id="B909EAE49534B84EFCD9FDB0C91CFDB4" box="[859,870,540,565]" italics="true" pageId="8" pageNumber="9">ξ</emphasis>
) 
<emphasis id="B909EAE49534B84EFCFBFDB1C9F4FDB7" box="[889,910,541,566]" italics="true" pageId="8" pageNumber="9">ft’</emphasis>
~ 
<emphasis id="B909EAE49534B84EFC2DFDB1C9BDFDB7" box="[943,967,541,566]" italics="true" pageId="8" pageNumber="9">ft”</emphasis>
; 
<emphasis id="B909EAE49534B84EFC5AFDB1C989FDB7" box="[984,1011,541,566]" italics="true" pageId="8" pageNumber="9">tc’</emphasis>
&lt;
<emphasis id="B909EAE49534B84EFB96FDB1CE48FDB7" box="[1044,1074,541,566]" italics="true" pageId="8" pageNumber="9">tc”</emphasis>
; 
<emphasis id="B909EAE49534B84EFBC1FDB1CE22FDB7" box="[1091,1112,541,566]" italics="true" pageId="8" pageNumber="9">p’</emphasis>
&lt;
<emphasis id="B909EAE49534B84EFBFBFDB1CEEBFDB7" box="[1145,1169,541,566]" italics="true" pageId="8" pageNumber="9">p”</emphasis>
. Tectals and prorals on pinnaculum, dorso-laterad of single claw.
</paragraph>
<paragraph id="8BC236F69534B84EFC0DFDF1CFD1FC97" blockId="8.[848,1452,253,790]" pageId="8" pageNumber="9">
Leg II (
<figureCitation id="13462A739534B84EFC65FDF1CE39FDF7" box="[999,1091,605,630]" captionStart="Fig" captionStartId="12.[850,885,1925,1945]" captionTargetBox="[102,1452,779,1899]" captionTargetId="figure-224@12.[87,907,775,1912]" captionTargetPageId="12" captionText="Fig. 13. Zambedania sekhukhunensis sp. n. Male paratype. Leg II (right): dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12956973" httpUri="https://zenodo.org/record/12956973/files/figure.png" pageId="8" pageNumber="9">Fig. 13</figureCitation>
). Trochanter: 
<emphasis id="B909EAE49534B84EFB6EFDF1CF7DFDF7" box="[1260,1287,605,630]" italics="true" pageId="8" pageNumber="9">”v’</emphasis>
&lt;Fe 
<emphasis id="B909EAE49534B84EFAD6FDF1CF01FDF7" box="[1364,1403,605,630]" italics="true" pageId="8" pageNumber="9">v”,”</emphasis>
the latter barbed on one side. Fe 
<emphasis id="B909EAE49534B84EFB2FFDD1CEC3FD17" box="[1197,1209,637,662]" italics="true" pageId="8" pageNumber="9">l’</emphasis>
barbed on one side, lacking on right leg, but present on the left, shortest seta on segment, 
<emphasis id="B909EAE49534B84EFBAFFD11CE46FD57" box="[1069,1084,701,726]" italics="true" pageId="8" pageNumber="9">d</emphasis>
the longest. Genual smooth 
<emphasis id="B909EAE49534B84EFA1FFD11CFD1FD57" box="[1437,1451,701,726]" italics="true" pageId="8" pageNumber="9">l”</emphasis>
~ slightly barbed 
<emphasis id="B909EAE49534B84EFBA4FD71CE48FD77" box="[1062,1074,733,758]" italics="true" pageId="8" pageNumber="9">l’</emphasis>
&lt;one-sidedly barbed 
<emphasis id="B909EAE49534B84EFAC5FD71CF20FD77" box="[1351,1370,733,758]" italics="true" pageId="8" pageNumber="9">v’</emphasis>
. Tibial one-sidedly barbed 
<emphasis id="B909EAE49534B84EFBBBFD51CE3FFC97" box="[1081,1093,765,790]" italics="true" pageId="8" pageNumber="9">l’</emphasis>
longest seta on verticil, 
<emphasis id="B909EAE49534B84EFADFFD51CF0AFC97" box="[1373,1392,765,790]" italics="true" pageId="8" pageNumber="9">v’</emphasis>
&gt; 
<emphasis id="B909EAE49534B84EFA0CFD51CFDEFC97" box="[1422,1444,765,790]" italics="true" pageId="8" pageNumber="9">v”</emphasis>
,
</paragraph>
<caption id="DF02667E9534B84EFF2CF82FCBE4F832" ID-DOI="http://doi.org/10.5281/zenodo.12956941" ID-Zenodo-Dep="12956941" httpUri="https://zenodo.org/record/12956941/files/figure.png" pageId="8" pageNumber="9" startId="8.[174,209,1923,1943]" targetBox="[175,1451,842,1897]" targetPageId="8" targetType="figure">
<paragraph id="8BC236F69534B84EFF2CF82FCBE4F832" blockId="8.[174,777,1923,1971]" pageId="8" pageNumber="9">
<emphasis id="B909EAE49534B84EFF2CF82FCA94F816" bold="true" box="[174,238,1923,1943]" pageId="8" pageNumber="9">Fig. 6.</emphasis>
<taxonomicName id="4C7D4D759534B84EFF7DF82FC863F816" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[255,537,1923,1943]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49534B84EFF7DF82FC863F816" box="[255,537,1923,1943]" italics="true" pageId="8" pageNumber="9">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9534B84EFDA3F82FC822F816" box="[545,600,1923,1943]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
Female holotype. Leg II (right): dorsal view.
</paragraph>
</caption>
<caption id="DF02667E9534B84EFCD0F82FCE32F832" pageId="8" pageNumber="9" startId="8.[850,885,1923,1943]" targetType="figure">
<paragraph id="8BC236F69534B84EFCD0F82FCE32F832" blockId="8.[850,1453,1923,1971]" pageId="8" pageNumber="9">
<emphasis id="B909EAE49534B84EFCD0F82FC9E9F816" bold="true" box="[850,915,1923,1943]" pageId="8" pageNumber="9">Fig. 7.</emphasis>
<taxonomicName id="4C7D4D759534B84EFC21F82FCEC4F816" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[931,1214,1923,1943]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49534B84EFC21F82FCEC4F816" box="[931,1214,1923,1943]" italics="true" pageId="8" pageNumber="9">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9534B84EFB47F82FCE86F816" box="[1221,1276,1923,1943]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
Female holotype. Leg III (right): dorsal view.
</paragraph>
</caption>
<paragraph id="8BC236F69535B84FFFFFFF04CBBEFF3E" blockId="9.[125,452,168,191]" box="[125,452,168,191]" pageId="9" pageNumber="10">
<bibCitation id="5E573E9A9535B84FFFFFFF04CBBEFF3E" box="[125,452,168,191]" journal="Zoological Studies" page="11" pageId="9" pageNumber="10" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49535B84FFFFFFF04CB4DFF3E" box="[125,311,168,191]" italics="true" pageId="9" pageNumber="10">Zoological Studies</emphasis>
<emphasis id="B909EAE49535B84FFEBFFF04CB24FF3E" bold="true" box="[317,350,168,191]" pageId="9" pageNumber="10">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69535B84FFF0AFF51C8FEFE37" blockId="9.[136,737,253,662]" pageId="9" pageNumber="10">
<emphasis id="B909EAE49535B84FFF0AFF51CAEDFE97" box="[136,151,253,278]" italics="true" pageId="9" pageNumber="10">d</emphasis>
shortest seta, closest to solenidion. Tarsus with robust solenidion ω proximal-most on segment; 
<emphasis id="B909EAE49535B84FFF0AFE91CADDFED7" box="[136,167,317,342]" italics="true" pageId="9" pageNumber="10">pl”</emphasis>
longest seta, on protuberance; barbed 
<emphasis id="B909EAE49535B84FFD28FE91C8BCFED7" box="[682,710,317,342]" italics="true" pageId="9" pageNumber="10">tc’</emphasis>
&lt;smooth 
<emphasis id="B909EAE49535B84FFF65FEF1CB7FFEF7" box="[231,261,349,374]" italics="true" pageId="9" pageNumber="10">tc”</emphasis>
, 
<emphasis id="B909EAE49535B84FFE91FEF1CB4FFEF7" box="[275,309,349,374]" italics="true" pageId="9" pageNumber="10">pv’</emphasis>
&lt;one-sidedly barbed 
<emphasis id="B909EAE49535B84FFDBFFEF1C819FEF7" box="[573,611,349,374]" italics="true" pageId="9" pageNumber="10">pv”</emphasis>
, 
<emphasis id="B909EAE49535B84FFDF3FEF1C8FCFEF7" box="[625,646,349,374]" italics="true" pageId="9" pageNumber="10">u’</emphasis>
&lt;to 
<emphasis id="B909EAE49535B84FFD40FEF1C8A0FEF7" box="[706,730,349,374]" italics="true" pageId="9" pageNumber="10">u”</emphasis>
, both one-sidedly barbed. Cordate empodium with medial fold between a pair of simple claws,
</paragraph>
<paragraph id="8BC236F69535B84FFF46FE11C89BFD17" blockId="9.[136,737,253,662]" pageId="9" pageNumber="10">
Leg III (
<figureCitation id="13462A739535B84FFEA0FE11CB01FE57" box="[290,379,445,470]" captionStart="Fig" captionStartId="13.[136,171,1926,1946]" captionTargetBox="[138,736,432,1900]" captionTargetId="figure-121@13.[125,740,424,1922]" captionTargetPageId="13" captionText="Fig. 14. Zambedania sekhukhunensis sp. n. Male paratype. Leg III (right): dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12956981" httpUri="https://zenodo.org/record/12956981/files/figure.png" pageId="9" pageNumber="10">Fig. 14</figureCitation>
). Third leg very much like the second leg; trochanteral setae barbed 
<emphasis id="B909EAE49535B84FFDD8FE71C817FE77" box="[602,621,477,502]" italics="true" pageId="9" pageNumber="10">v’</emphasis>
&gt; barbed Fe 
<emphasis id="B909EAE49535B84FFF30FE51CABCFD97" box="[178,198,509,534]" italics="true" pageId="9" pageNumber="10">v’</emphasis>
. Femur with longest, robust, barbed seta 
<emphasis id="B909EAE49535B84FFD50FE51C89BFD97" box="[722,737,509,534]" italics="true" pageId="9" pageNumber="10">d</emphasis>
extending to tarsus. Genu with barbed setae 
<emphasis id="B909EAE49535B84FFD36FDB1C8BDFDB7" box="[692,711,541,566]" italics="true" pageId="9" pageNumber="10">v’</emphasis>
~ 
<emphasis id="B909EAE49535B84FFF0AFD91CAE1FDD7" box="[136,155,573,598]" italics="true" pageId="9" pageNumber="10">l’.</emphasis>
Tibial 
<emphasis id="B909EAE49535B84FFF6BFD91CA82FDD7" box="[233,248,573,598]" italics="true" pageId="9" pageNumber="10">d</emphasis>
closest to solenidion Φ and &lt;one-sidedly barbed 
<emphasis id="B909EAE49535B84FFF66FDF1CA8DFDF7" box="[228,247,605,630]" italics="true" pageId="9" pageNumber="10">l’.</emphasis>
Seta 
<emphasis id="B909EAE49535B84FFEC2FDF1CB37FDF7" box="[320,333,605,630]" italics="true" pageId="9" pageNumber="10">v</emphasis>
’ ~ 
<emphasis id="B909EAE49535B84FFEF1FDF1CBF0FDF7" box="[371,394,605,630]" italics="true" pageId="9" pageNumber="10">v”</emphasis>
, both slightly barbed; similar to leg II, leg III bears a robust tarsal solenidion ω;
</paragraph>
<paragraph id="8BC236F69535B84FFB79FF04CFFFFF3E" blockId="9.[1275,1413,168,191]" box="[1275,1413,168,191]" pageId="9" pageNumber="10">page 10 of 21</paragraph>
<paragraph id="8BC236F69535B84FFCAFFF51CE93FE17" blockId="9.[813,1414,253,663]" pageId="9" pageNumber="10">
one-sidedly barbed 
<emphasis id="B909EAE49535B84FFBA1FF51CE3BFE97" box="[1059,1089,253,278]" italics="true" pageId="9" pageNumber="10">pl”</emphasis>
longest seta on segment, slightly barbed 
<emphasis id="B909EAE49535B84FFC61FEB1C984FEB7" box="[995,1022,285,310]" italics="true" pageId="9" pageNumber="10">tc’</emphasis>
&gt; smooth 
<emphasis id="B909EAE49535B84FFB02FEB1CEE4FEB7" box="[1152,1182,285,310]" italics="true" pageId="9" pageNumber="10">tc”</emphasis>
, slightly barbed 
<emphasis id="B909EAE49535B84FFAE7FEB1CFFCFEB7" box="[1381,1414,285,310]" italics="true" pageId="9" pageNumber="10">pv’</emphasis>
~ one-sidedly barbed 
<emphasis id="B909EAE49535B84FFBBAFE91CE27FED7" box="[1080,1117,317,342]" italics="true" pageId="9" pageNumber="10">pv”</emphasis>
; barbed 
<emphasis id="B909EAE49535B84FFB4FFE91CE80FED7" box="[1229,1274,317,342]" italics="true" pageId="9" pageNumber="10">u’ &lt;</emphasis>
smooth 
<emphasis id="B909EAE49535B84FFAE4FE91CF04FED7" box="[1382,1406,317,342]" italics="true" pageId="9" pageNumber="10">u”</emphasis>
; as in leg II terminating in a pair of simple claws with a cordate empodium in-between.
</paragraph>
<paragraph id="8BC236F69535B84FFCEBFE31CFFFFD17" blockId="9.[813,1414,253,663]" pageId="9" pageNumber="10">
Leg IV (
<figureCitation id="13462A739535B84FFC47FE31CE67FE37" box="[965,1053,413,438]" captionStart="Fig" captionStartId="13.[813,848,1926,1946]" captionTargetBox="[814,1413,638,1898]" captionTargetId="figure-135@13.[802,1450,617,1912]" captionTargetPageId="13" captionText="Fig. 15. Zambedania sekhukhunensis sp. n. Male paratype. Leg IV (left): dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12956989" httpUri="https://zenodo.org/record/12956989/files/figure.png" pageId="9" pageNumber="10">Fig. 15</figureCitation>
). Leg IV modified; trochanteral seta barbed 
<emphasis id="B909EAE49535B84FFC55FE11CE7CFE57" box="[983,1030,445,470]" italics="true" pageId="9" pageNumber="10">v’&gt;</emphasis>
Fe 
<emphasis id="B909EAE49535B84FFBB7FE11CE30FE57" box="[1077,1098,445,470]" italics="true" pageId="9" pageNumber="10">v’</emphasis>
; Fe 
<emphasis id="B909EAE49535B84FFB0EFE11CEE1FE57" box="[1164,1179,445,470]" italics="true" pageId="9" pageNumber="10">d</emphasis>
slightly barbed ~ barbed 
<emphasis id="B909EAE49535B84FFC0AFE71C9E1FE77" box="[904,923,477,502]" italics="true" pageId="9" pageNumber="10">v’</emphasis>
. Genu: barbed 
<emphasis id="B909EAE49535B84FFBD5FE71CE10FE77" box="[1111,1130,477,502]" italics="true" pageId="9" pageNumber="10">v’</emphasis>
&gt; Fe 
<emphasis id="B909EAE49535B84FFB33FE71CEC4FE77" box="[1201,1214,477,502]" italics="true" pageId="9" pageNumber="10">v</emphasis>
’; 
<emphasis id="B909EAE49535B84FFB56FE71CE9AFE77" box="[1236,1248,477,502]" italics="true" pageId="9" pageNumber="10">l’</emphasis>
absent. Tibia: short Φ close to smooth 
<emphasis id="B909EAE49535B84FFBE9FE51CE00FD97" box="[1131,1146,509,534]" italics="true" pageId="9" pageNumber="10">d</emphasis>
; smooth 
<emphasis id="B909EAE49535B84FFB74FE51CF79FD97" box="[1270,1283,509,534]" italics="true" pageId="9" pageNumber="10">l’</emphasis>
&lt;
<emphasis id="B909EAE49535B84FFAA8FE51CF43FD97" box="[1322,1337,509,534]" italics="true" pageId="9" pageNumber="10">d</emphasis>
; one-sidedly barbed 
<emphasis id="B909EAE49535B84FFC69FDB1C985FDB7" box="[1003,1023,541,566]" italics="true" pageId="9" pageNumber="10">v’</emphasis>
longest on segment, smooth 
<emphasis id="B909EAE49535B84FFAEDFDB1CFFCFDB7" box="[1391,1414,541,566]" italics="true" pageId="9" pageNumber="10">v”</emphasis>
the shortest. Tarsus very short with modified claw that seems to fit in a hooked claw-like structure 
<emphasis id="B909EAE49535B84FFAEBFDF1CFFCFDF7" box="[1385,1414,605,630]" italics="true" pageId="9" pageNumber="10">u’.</emphasis>
Setae 
<emphasis id="B909EAE49535B84FFCF9FDD1C9B4FD17" box="[891,974,637,662]" italics="true" pageId="9" pageNumber="10">pl”, pv”</emphasis>
and 
<emphasis id="B909EAE49535B84FFB8EFDD1CE54FD17" box="[1036,1070,637,662]" italics="true" pageId="9" pageNumber="10">pv’</emphasis>
short and smooth; very long
</paragraph>
<caption id="DF02667E9535B84FFF0AF831C9A0F830" box="[136,986,1949,1969]" pageId="9" pageNumber="10" startId="9.[136,170,1949,1969]" targetType="figure">
<paragraph id="8BC236F69535B84FFF0AF831C9A0F830" blockId="9.[136,986,1949,1969]" box="[136,986,1949,1969]" pageId="9" pageNumber="10">
<emphasis id="B909EAE49535B84FFF0AF831CABCF830" bold="true" box="[136,198,1949,1969]" pageId="9" pageNumber="10">Fig. 8.</emphasis>
<taxonomicName id="4C7D4D759535B84FFF50F831CB9DF830" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[210,487,1949,1969]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49535B84FFF50F831CB9DF830" box="[210,487,1949,1969]" italics="true" pageId="9" pageNumber="10">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9535B84FFE6FF831C858F830" box="[493,546,1949,1969]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
Female holotype. Leg IV (right): paraxial view.
</paragraph>
</caption>
<paragraph id="8BC236F69536B84CFF2CFF04CB8EFF3E" blockId="10.[174,500,168,191]" box="[174,500,168,191]" pageId="10" pageNumber="11">
<bibCitation id="5E573E9A9536B84CFF2CFF04CB8EFF3E" box="[174,500,168,191]" journal="Zoological Studies" page="11" pageId="10" pageNumber="11" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49536B84CFF2CFF04CB1DFF3E" box="[174,359,168,191]" italics="true" pageId="10" pageNumber="11">Zoological Studies</emphasis>
<emphasis id="B909EAE49536B84CFEEFFF04CBF4FF3E" bold="true" box="[365,398,168,191]" pageId="10" pageNumber="11">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69536B84CFAA0FF04CFD1FF3E" blockId="10.[1314,1451,168,191]" box="[1314,1451,168,191]" pageId="10" pageNumber="11">page 11 of 21</paragraph>
<caption id="DF02667E9536B84CFF2CFB5CCA81FA0E" ID-DOI="http://doi.org/10.5281/zenodo.12956951" ID-Zenodo-Dep="12956951" httpUri="https://zenodo.org/record/12956951/files/figure.png" pageId="10" pageNumber="11" startId="10.[174,208,1264,1284]" targetBox="[176,1449,252,1233]" targetPageId="10" targetType="figure">
<paragraph id="8BC236F69536B84CFF2CFB5CCA81FA0E" blockId="10.[174,1453,1264,1423]" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFF2CFB5CCA97FA85" bold="true" box="[174,237,1264,1284]" pageId="10" pageNumber="11">Fig. 9.</emphasis>
<taxonomicName id="4C7D4D759536B84CFF79FB5CC868FA85" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[251,530,1264,1284]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49536B84CFF79FB5CC868FA85" box="[251,530,1264,1284]" italics="true" pageId="10" pageNumber="11">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9536B84CFD9BFB5CC835FA85" box="[537,591,1264,1284]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
Female. A- dorsal view; B- caput of bothridial seta, dorsal view; in front “pilose” setae; C- Leg I (right). Note TiTa in dorsal view. At distal end “turret” carrying claw with at its base dorsal pinnaculum of the “locking system”, flanked by part of counterpiece. Eupathidia dorsad on pinnaculi. The cluster 
<emphasis id="B909EAE49536B84CFC01FA84C9F5FABD" box="[899,911,1320,1340]" italics="true" pageId="10" pageNumber="11">d</emphasis>
and 
<emphasis id="B909EAE49536B84CFC3DFA84C9B0FABD" box="[959,970,1320,1340]" italics="true" pageId="10" pageNumber="11">k</emphasis>
associated with the three solenidia. Ventro-laterad near to counter piece seta 
<emphasis id="B909EAE49536B84CFE32FAE8CBC1FAD9" box="[432,443,1348,1368]" italics="true" pageId="10" pageNumber="11">s</emphasis>
followed by feathered 
<emphasis id="B909EAE49536B84CFD14FAE8C8D7FAD9" box="[662,685,1348,1368]" italics="true" pageId="10" pageNumber="11">pv</emphasis>
and 
<emphasis id="B909EAE49536B84CFD5CFAE8C894FAD9" box="[734,750,1348,1368]" italics="true" pageId="10" pageNumber="11">v.</emphasis>
D- detail distal end of left TiTa I in lateral view, claw and butterfly-shaped counter piece made up of setae 
<emphasis id="B909EAE49536B84CFE64FACCC864FAF5" box="[486,542,1376,1396]" italics="true" pageId="10" pageNumber="11">u’ - u”</emphasis>
, below seta 
<emphasis id="B909EAE49536B84CFD18FACCC8D0FAF5" box="[666,682,1376,1396]" italics="true" pageId="10" pageNumber="11">s,</emphasis>
followed by “feathered” primiventrals 
<emphasis id="B909EAE49536B84CFB97FACCCE55FAF5" box="[1045,1071,1376,1396]" italics="true" pageId="10" pageNumber="11">pv’</emphasis>
and 
<emphasis id="B909EAE49536B84CFBE3FACCCE04FAF5" box="[1121,1150,1376,1396]" italics="true" pageId="10" pageNumber="11">pv”</emphasis>
(photos by G. Bauchan and R. Ochoa).
</paragraph>
</caption>
<caption id="DF02667E9536B84CFF2CFA6BCF7DFA61" box="[174,1287,1479,1504]" pageId="10" pageNumber="11" startId="10.[174,242,1479,1504]" targetType="table">
<paragraph id="8BC236F69536B84CFF2CFA6BCF7DFA61" blockId="10.[174,1287,1479,1504]" box="[174,1287,1479,1504]" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFF2CFA6BCB75FA61" bold="true" box="[174,271,1479,1504]" pageId="10" pageNumber="11">Table 4.</emphasis>
Leg chaeto- and solenidiotaxy of the genus 
<taxonomicName id="4C7D4D759536B84CFCA0FA6BCE07FA61" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[802,1149,1479,1504]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49536B84CFCA0FA6BCE07FA61" box="[802,1149,1479,1504]" italics="true" pageId="10" pageNumber="11">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9536B84CFB06FA6BCEBFFA61" box="[1156,1221,1479,1504]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
larva
</paragraph>
</caption>
<paragraph id="8BC236F69536B84CFE38F9A4CE22F8D8" pageId="10" pageNumber="11">
<table id="F97DC456953647B9FF30F9A4CF52F8D8" box="[178,1320,1544,1881]" gridcols="7" gridrows="11" pageId="10" pageNumber="11">
<tr id="354D34B4953647B9FF30F9A4CF52F99C" box="[178,1320,1544,1565]" gridrow="0" pageId="10" pageNumber="11" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<th id="769C5DC8953647B9FE3AF9A4C8BEF99C" box="[440,708,1544,1565]" gridcol="2" gridrow="0" pageId="10" pageNumber="11">LEG I</th>
<th id="769C5DC8953647B9FC99F9A4C998F99C" box="[795,994,1544,1565]" gridcol="4" gridrow="0" pageId="10" pageNumber="11">LEG II</th>
<th id="769C5DC8953647B9FBFDF9A4CF52F99C" box="[1151,1320,1544,1565]" gridcol="6" gridrow="0" pageId="10" pageNumber="11">LEG III</th>
</tr>
<tr id="354D34B4953647B9FF30F9ECCF52F9D4" box="[178,1320,1600,1621]" gridrow="1" pageId="10" pageNumber="11">
<th id="769C5DC8953647B9FF30F9ECCB3CF9D4" box="[178,326,1600,1621]" gridcol="0" gridrow="1" pageId="10" pageNumber="11">SEGMENT</th>
<td id="769C5DC8953647B9FE07F9ECCBE6F9D4" box="[389,412,1600,1621]" gridcol="1" gridrow="1" pageId="10" pageNumber="11">Nr</td>
<td id="769C5DC8953647B9FE3AF9ECC8BEF9D4" box="[440,708,1600,1621]" gridcol="2" gridrow="1" pageId="10" pageNumber="11">SIGLA</td>
<td id="769C5DC8953647B9FD6AF9ECC97AF9D4" box="[744,768,1600,1621]" gridcol="3" gridrow="1" pageId="10" pageNumber="11">Nr</td>
<td id="769C5DC8953647B9FC99F9ECC998F9D4" box="[795,994,1600,1621]" gridcol="4" gridrow="1" pageId="10" pageNumber="11">SIGLA</td>
<td id="769C5DC8953647B9FBCEF9ECCE19F9D4" box="[1100,1123,1600,1621]" gridcol="5" gridrow="1" pageId="10" pageNumber="11">Nr</td>
<td id="769C5DC8953647B9FBFDF9ECCF52F9D4" box="[1151,1320,1600,1621]" gridcol="6" gridrow="1" pageId="10" pageNumber="11">SIGLA</td>
</tr>
<tr id="354D34B4953647B9FF30F9F1CF52F9F0" box="[178,1320,1629,1649]" gridrow="2" pageId="10" pageNumber="11" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<th id="769C5DC8953647B9FF30F9F1CB3CF9F0" box="[178,326,1629,1649]" gridcol="0" gridrow="2" pageId="10" pageNumber="11">TROCHANTER</th>
<td id="769C5DC8953647B9FE07F9F1CBE6F9F0" box="[389,412,1629,1649]" gridcol="1" gridrow="2" pageId="10" pageNumber="11">0</td>
<td id="769C5DC8953647B9FD6AF9F1C97AF9F0" box="[744,768,1629,1649]" gridcol="3" gridrow="2" pageId="10" pageNumber="11">0</td>
<td id="769C5DC8953647B9FBCEF9F1CE19F9F0" box="[1100,1123,1629,1649]" gridcol="5" gridrow="2" pageId="10" pageNumber="11">0</td>
</tr>
<tr id="354D34B4953647B9FF30F9D6CF52F90F" box="[178,1320,1658,1678]" gridrow="3" pageId="10" pageNumber="11">
<th id="769C5DC8953647B9FF30F9D6CB3CF90F" box="[178,326,1658,1678]" gridcol="0" gridrow="3" pageId="10" pageNumber="11">FEMUR</th>
<td id="769C5DC8953647B9FE07F9D6CBE6F90F" box="[389,412,1658,1678]" gridcol="1" gridrow="3" pageId="10" pageNumber="11">3</td>
<td id="769C5DC8953647B9FE3AF9D6C8BEF90F" box="[440,708,1658,1678]" gridcol="2" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFE38F9D6CBBCF90F" box="[442,454,1658,1678]" italics="true" pageId="10" pageNumber="11">d</emphasis>
, 
<emphasis id="B909EAE49536B84CFE4EF9D6CBAFF90F" box="[460,469,1658,1678]" italics="true" pageId="10" pageNumber="11">l'</emphasis>
, 
<emphasis id="B909EAE49536B84CFE59F9D6CB97F90F" box="[475,493,1658,1678]" italics="true" pageId="10" pageNumber="11">v&quot;</emphasis>
</td>
<td id="769C5DC8953647B9FD6AF9D6C97AF90F" box="[744,768,1658,1678]" gridcol="3" gridrow="3" pageId="10" pageNumber="11">3</td>
<td id="769C5DC8953647B9FC99F9D6C998F90F" box="[795,994,1658,1678]" gridcol="4" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFC9FF9D6C953F90F" box="[797,809,1658,1678]" italics="true" pageId="10" pageNumber="11">d</emphasis>
, 
<emphasis id="B909EAE49536B84CFCADF9D6C942F90F" box="[815,824,1658,1678]" italics="true" pageId="10" pageNumber="11">l'</emphasis>
, 
<emphasis id="B909EAE49536B84CFCBCF9D6C92BF90F" box="[830,849,1658,1678]" italics="true" pageId="10" pageNumber="11">v&quot;</emphasis>
</td>
<td id="769C5DC8953647B9FBCEF9D6CE19F90F" box="[1100,1123,1658,1678]" gridcol="5" gridrow="3" pageId="10" pageNumber="11">2</td>
<td id="769C5DC8953647B9FBFDF9D6CF52F90F" box="[1151,1320,1658,1678]" gridcol="6" gridrow="3" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFB03F9D6CEF7F90F" box="[1153,1165,1658,1678]" italics="true" pageId="10" pageNumber="11">d</emphasis>
, 
<emphasis id="B909EAE49536B84CFB11F9D6CED8F90F" box="[1171,1186,1658,1678]" italics="true" pageId="10" pageNumber="11">v'</emphasis>
</td>
</tr>
<tr id="354D34B4953647B9FF30F93BCF52F92A" box="[178,1320,1687,1707]" gridrow="4" pageId="10" pageNumber="11">
<th id="769C5DC8953647B9FF30F93BCB3CF92A" box="[178,326,1687,1707]" gridcol="0" gridrow="4" pageId="10" pageNumber="11">GENU</th>
<td id="769C5DC8953647B9FE07F93BCBE6F92A" box="[389,412,1687,1707]" gridcol="1" gridrow="4" pageId="10" pageNumber="11">4</td>
<td id="769C5DC8953647B9FE3AF93BC8BEF92A" box="[440,708,1687,1707]" gridcol="2" gridrow="4" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFE38F93BCBB9F92A" box="[442,451,1687,1707]" italics="true" pageId="10" pageNumber="11">l'</emphasis>
, 
<emphasis id="B909EAE49536B84CFE4BF93BCBACF92A" box="[457,470,1687,1707]" italics="true" pageId="10" pageNumber="11">l&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFE59F93BCB90F92A" box="[475,490,1687,1707]" italics="true" pageId="10" pageNumber="11">v'</emphasis>
, 
<emphasis id="B909EAE49536B84CFE74F93BC872F92A" box="[502,520,1687,1707]" italics="true" pageId="10" pageNumber="11">v&quot;</emphasis>
</td>
<td id="769C5DC8953647B9FD6AF93BC97AF92A" box="[744,768,1687,1707]" gridcol="3" gridrow="4" pageId="10" pageNumber="11">3</td>
<td id="769C5DC8953647B9FC99F93BC998F92A" box="[795,994,1687,1707]" gridcol="4" gridrow="4" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFC9FF93BC95CF92A" box="[797,806,1687,1707]" italics="true" pageId="10" pageNumber="11">l'</emphasis>
, 
<emphasis id="B909EAE49536B84CFCAEF93BC945F92A" box="[812,831,1687,1707]" italics="true" pageId="10" pageNumber="11">v&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFCC6F93BC929F92A" box="[836,851,1687,1707]" italics="true" pageId="10" pageNumber="11">v'</emphasis>
</td>
<td id="769C5DC8953647B9FBCEF93BCE19F92A" box="[1100,1123,1687,1707]" gridcol="5" gridrow="4" pageId="10" pageNumber="11">2</td>
<td id="769C5DC8953647B9FBFDF93BCF52F92A" box="[1151,1320,1687,1707]" gridcol="6" gridrow="4" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFB03F93BCEF0F92A" box="[1153,1162,1687,1707]" italics="true" pageId="10" pageNumber="11">l'</emphasis>
, 
<emphasis id="B909EAE49536B84CFB0DF93BCEE4F92A" box="[1167,1182,1687,1707]" italics="true" pageId="10" pageNumber="11">v'</emphasis>
</td>
</tr>
<tr id="354D34B4953647B9FF30F918CF52F949" box="[178,1320,1716,1736]" gridrow="5" pageId="10" pageNumber="11">
<th id="769C5DC8953647B9FF30F918CB3CF949" box="[178,326,1716,1736]" gridcol="0" gridrow="5" pageId="10" pageNumber="11">TIBIA</th>
<td id="769C5DC8953647B9FE07F918CBE6F949" box="[389,412,1716,1736]" gridcol="1" gridrow="5" pageId="10" pageNumber="11">6</td>
<td id="769C5DC8953647B9FE3AF918C8BEF949" box="[440,708,1716,1736]" gridcol="2" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFE38F918CBB9F949" box="[442,451,1716,1736]" italics="true" pageId="10" pageNumber="11">l'</emphasis>
, 
<emphasis id="B909EAE49536B84CFE4BF918CBACF949" box="[457,470,1716,1736]" italics="true" pageId="10" pageNumber="11">l&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFE59F918CB90F949" box="[475,490,1716,1736]" italics="true" pageId="10" pageNumber="11">v'</emphasis>
, 
<emphasis id="B909EAE49536B84CFE72F918C878F949" box="[496,514,1716,1736]" italics="true" pageId="10" pageNumber="11">v&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFD8AF918C869F949" box="[520,531,1716,1736]" italics="true" pageId="10" pageNumber="11">k</emphasis>
, 
<emphasis id="B909EAE49536B84CFD9BF918C85FF949" box="[537,549,1716,1736]" italics="true" pageId="10" pageNumber="11">d</emphasis>
</td>
<td id="769C5DC8953647B9FD6AF918C97AF949" box="[744,768,1716,1736]" gridcol="3" gridrow="5" pageId="10" pageNumber="11">4</td>
<td id="769C5DC8953647B9FC99F918C998F949" box="[795,994,1716,1736]" gridcol="4" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFC9FF918C953F949" box="[797,809,1716,1736]" italics="true" pageId="10" pageNumber="11">d</emphasis>
, 
<emphasis id="B909EAE49536B84CFCADF918C942F949" box="[815,824,1716,1736]" italics="true" pageId="10" pageNumber="11">l'</emphasis>
, 
<emphasis id="B909EAE49536B84CFCBCF918C937F949" box="[830,845,1716,1736]" italics="true" pageId="10" pageNumber="11">v'</emphasis>
, 
<emphasis id="B909EAE49536B84CFCD1F918C91FF949" box="[851,869,1716,1736]" italics="true" pageId="10" pageNumber="11">v&quot;</emphasis>
</td>
<td id="769C5DC8953647B9FBCEF918CE19F949" box="[1100,1123,1716,1736]" gridcol="5" gridrow="5" pageId="10" pageNumber="11">4</td>
<td id="769C5DC8953647B9FBFDF918CF52F949" box="[1151,1320,1716,1736]" gridcol="6" gridrow="5" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFB03F918CEF7F949" box="[1153,1165,1716,1736]" italics="true" pageId="10" pageNumber="11">d</emphasis>
, 
<emphasis id="B909EAE49536B84CFB10F918CEE1F949" box="[1170,1179,1716,1736]" italics="true" pageId="10" pageNumber="11">l'</emphasis>
, 
<emphasis id="B909EAE49536B84CFB23F918CECAF949" box="[1185,1200,1716,1736]" italics="true" pageId="10" pageNumber="11">v'</emphasis>
, 
<emphasis id="B909EAE49536B84CFB34F918CEB2F949" box="[1206,1224,1716,1736]" italics="true" pageId="10" pageNumber="11">v&quot;</emphasis>
</td>
</tr>
<tr id="354D34B4953647B9FF30F97DCF52F964" box="[178,1320,1745,1765]" gridrow="6" pageId="10" pageNumber="11">
<th id="769C5DC8953647B9FF30F97DCB3CF964" box="[178,326,1745,1765]" gridcol="0" gridrow="6" pageId="10" pageNumber="11">Solenidia</th>
<td id="769C5DC8953647B9FE07F97DCBE6F964" box="[389,412,1745,1765]" gridcol="1" gridrow="6" pageId="10" pageNumber="11">1</td>
<td id="769C5DC8953647B9FE3AF97DC8BEF964" box="[440,708,1745,1765]" gridcol="2" gridrow="6" pageId="10" pageNumber="11">
Φ 
<emphasis id="B909EAE49536B84CFE47F97DCBABF964" box="[453,465,1745,1765]" italics="true" pageId="10" pageNumber="11">1</emphasis>
</td>
<td id="769C5DC8953647B9FD6AF97DC97AF964" box="[744,768,1745,1765]" gridcol="3" gridrow="6" pageId="10" pageNumber="11">1</td>
<td id="769C5DC8953647B9FC99F97DC998F964" box="[795,994,1745,1765]" gridcol="4" gridrow="6" pageId="10" pageNumber="11">Φ</td>
<td id="769C5DC8953647B9FBCEF97DCE19F964" box="[1100,1123,1745,1765]" gridcol="5" gridrow="6" pageId="10" pageNumber="11">1</td>
<td id="769C5DC8953647B9FBFDF97DCF52F964" box="[1151,1320,1745,1765]" gridcol="6" gridrow="6" pageId="10" pageNumber="11">Φ</td>
</tr>
<tr id="354D34B4953647B9FF30F942CF52F883" box="[178,1320,1774,1794]" gridrow="7" pageId="10" pageNumber="11">
<th id="769C5DC8953647B9FF30F942CB3CF883" box="[178,326,1774,1794]" gridcol="0" gridrow="7" pageId="10" pageNumber="11">TARSUS</th>
<td id="769C5DC8953647B9FE07F942CBE6F883" box="[389,412,1774,1794]" gridcol="1" gridrow="7" pageId="10" pageNumber="11">11</td>
<td id="769C5DC8953647B9FE3AF942C8BEF883" box="[440,708,1774,1794]" gridcol="2" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFE38F942CBB5F883" box="[442,463,1774,1794]" italics="true" pageId="10" pageNumber="11">pl'</emphasis>
, 
<emphasis id="B909EAE49536B84CFE57F942CB97F883" box="[469,493,1774,1794]" italics="true" pageId="10" pageNumber="11">pl&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFE71F942C874F883" box="[499,526,1774,1794]" italics="true" pageId="10" pageNumber="11">pv'</emphasis>
, 
<emphasis id="B909EAE49536B84CFD91F942C848F883" box="[531,562,1774,1794]" italics="true" pageId="10" pageNumber="11">pv&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFDB5F942C83DF883" box="[567,583,1774,1794]" italics="true" pageId="10" pageNumber="11">u'</emphasis>
, 
<emphasis id="B909EAE49536B84CFDCFF942C81BF883" box="[589,609,1774,1794]" italics="true" pageId="10" pageNumber="11">u&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFDE5F942C808F883" box="[615,626,1774,1794]" italics="true" pageId="10" pageNumber="11">s</emphasis>
, 
<emphasis id="B909EAE49536B84CFDF5F942C8F6F883" box="[631,652,1774,1794]" italics="true" pageId="10" pageNumber="11">tc'</emphasis>
, 
<emphasis id="B909EAE49536B84CFD10F942C8D0F883" box="[658,682,1774,1794]" italics="true" pageId="10" pageNumber="11">tc&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFD32F942C8BEF883" box="[688,708,1774,1794]" italics="true" pageId="10" pageNumber="11">ft&quot;</emphasis>
</td>
<td id="769C5DC8953647B9FD6AF942C97AF883" box="[744,768,1774,1794]" gridcol="3" gridrow="7" pageId="10" pageNumber="11">7</td>
<td id="769C5DC8953647B9FC99F942C998F883" box="[795,994,1774,1794]" gridcol="4" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFC9FF942C94CF883" box="[797,822,1774,1794]" italics="true" pageId="10" pageNumber="11">pl&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFCBEF942C92BF883" box="[828,849,1774,1794]" italics="true" pageId="10" pageNumber="11">tc'</emphasis>
, 
<emphasis id="B909EAE49536B84CFCD4F942C915F883" box="[854,879,1774,1794]" italics="true" pageId="10" pageNumber="11">tc&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFCF6F942C9F5F883" box="[884,911,1774,1794]" italics="true" pageId="10" pageNumber="11">pv'</emphasis>
, 
<emphasis id="B909EAE49536B84CFC17F942C9C9F883" box="[917,947,1774,1794]" italics="true" pageId="10" pageNumber="11">pv&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFC3BF942C9B3F883" box="[953,969,1774,1794]" italics="true" pageId="10" pageNumber="11">u'</emphasis>
, 
<emphasis id="B909EAE49536B84CFC4DF942C998F883" box="[975,994,1774,1794]" italics="true" pageId="10" pageNumber="11">u&quot;</emphasis>
</td>
<td id="769C5DC8953647B9FBCEF942CE19F883" box="[1100,1123,1774,1794]" gridcol="5" gridrow="7" pageId="10" pageNumber="11">7</td>
<td id="769C5DC8953647B9FBFDF942CF52F883" box="[1151,1320,1774,1794]" gridcol="6" gridrow="7" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFB03F942CEE3F883" box="[1153,1177,1774,1794]" italics="true" pageId="10" pageNumber="11">pl&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFB1DF942CECEF883" box="[1183,1204,1774,1794]" italics="true" pageId="10" pageNumber="11">tc'</emphasis>
, 
<emphasis id="B909EAE49536B84CFB3BF942CEA8F883" box="[1209,1234,1774,1794]" italics="true" pageId="10" pageNumber="11">tc&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFB5AF942CE89F883" box="[1240,1267,1774,1794]" italics="true" pageId="10" pageNumber="11">pv'</emphasis>
, 
<emphasis id="B909EAE49536B84CFB7AF942CF6CF883" box="[1272,1302,1774,1794]" italics="true" pageId="10" pageNumber="11">pv&quot;</emphasis>
, 
<emphasis id="B909EAE49536B84CFA9EF942CF52F883" box="[1308,1320,1774,1794]" italics="true" pageId="10" pageNumber="11">u</emphasis>
</td>
</tr>
<tr id="354D34B4953647B9FF30F8A7CF52F89E" box="[178,1320,1803,1823]" gridrow="8" pageId="10" pageNumber="11" rowspan-1="1" rowspan-4="1" rowspan-6="1">
<th id="769C5DC8953647B9FF30F8A7CB3CF89E" box="[178,326,1803,1823]" gridcol="0" gridrow="8" pageId="10" pageNumber="11">Eupathidia</th>
<td id="769C5DC8953647B9FE3AF8A7C8BEF89E" box="[440,708,1803,1823]" gridcol="2" gridrow="8" pageId="10" pageNumber="11">
<emphasis id="B909EAE49536B84CFE38F8A7CBB0F89E" box="[442,458,1803,1823]" italics="true" pageId="10" pageNumber="11">ft'</emphasis>
</td>
<td id="769C5DC8953647B9FD6AF8A7C97AF89E" box="[744,768,1803,1823]" gridcol="3" gridrow="8" pageId="10" pageNumber="11">0</td>
<td id="769C5DC8953647B9FBCEF8A7CE19F89E" box="[1100,1123,1803,1823]" gridcol="5" gridrow="8" pageId="10" pageNumber="11">0</td>
</tr>
<tr id="354D34B4953647B9FF30F88BCF52F8BD" box="[178,1320,1831,1852]" gridrow="9" pageId="10" pageNumber="11" rowspan-6="1">
<th id="769C5DC8953647B9FF30F88BCB3CF8BD" box="[178,326,1831,1852]" gridcol="0" gridrow="9" pageId="10" pageNumber="11">Solenidia</th>
<td id="769C5DC8953647B9FE07F88BCBE6F8BD" box="[389,412,1831,1852]" gridcol="1" gridrow="9" pageId="10" pageNumber="11">1</td>
<td id="769C5DC8953647B9FE3AF88BC8BEF8BD" box="[440,708,1831,1852]" gridcol="2" gridrow="9" pageId="10" pageNumber="11">
ω 
<emphasis id="B909EAE49536B84CFE4BF884CBAFF8BD" box="[457,469,1832,1852]" italics="true" pageId="10" pageNumber="11">1</emphasis>
</td>
<td id="769C5DC8953647B9FD6AF88BC97AF8BD" box="[744,768,1831,1852]" gridcol="3" gridrow="9" pageId="10" pageNumber="11">1</td>
<td id="769C5DC8953647B9FC99F88BC998F8BD" box="[795,994,1831,1852]" gridcol="4" gridrow="9" pageId="10" pageNumber="11">ω</td>
<td id="769C5DC8953647B9FBCEF88BCE19F8BD" box="[1100,1123,1831,1852]" gridcol="5" gridrow="9" pageId="10" pageNumber="11">0</td>
</tr>
<tr id="354D34B4953647B9FF30F8E8CF52F8D8" box="[178,1320,1860,1881]" gridrow="10" pageId="10" pageNumber="11" rowspan-2="1" rowspan-4="1" rowspan-6="1">
<th id="769C5DC8953647B9FF30F8E8CB3CF8D8" box="[178,326,1860,1881]" gridcol="0" gridrow="10" pageId="10" pageNumber="11">Claws</th>
<td id="769C5DC8953647B9FE07F8E8CBE6F8D8" box="[389,412,1860,1881]" gridcol="1" gridrow="10" pageId="10" pageNumber="11">2</td>
<td id="769C5DC8953647B9FD6AF8E8C97AF8D8" box="[744,768,1860,1881]" gridcol="3" gridrow="10" pageId="10" pageNumber="11">2</td>
<td id="769C5DC8953647B9FBCEF8E8CE19F8D8" box="[1100,1123,1860,1881]" gridcol="5" gridrow="10" pageId="10" pageNumber="11">2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC236F69537B84DFFFFFF04CBBEFF3E" blockId="11.[125,452,168,191]" box="[125,452,168,191]" pageId="11" pageNumber="12">
<bibCitation id="5E573E9A9537B84DFFFFFF04CBBEFF3E" box="[125,452,168,191]" journal="Zoological Studies" page="11" pageId="11" pageNumber="12" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49537B84DFFFFFF04CB4DFF3E" box="[125,311,168,191]" italics="true" pageId="11" pageNumber="12">Zoological Studies</emphasis>
<emphasis id="B909EAE49537B84DFEBFFF04CB24FF3E" bold="true" box="[317,350,168,191]" pageId="11" pageNumber="12">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69537B84DFB79FF04CFFFFF3E" blockId="11.[1275,1413,168,191]" box="[1275,1413,168,191]" pageId="11" pageNumber="12">page 12 of 21</paragraph>
<caption id="DF02667E9537B84DFF0AFB2BC962FB36" ID-DOI="http://doi.org/10.5281/zenodo.12956957" ID-Zenodo-Dep="12956957" httpUri="https://zenodo.org/record/12956957/files/figure.png" pageId="11" pageNumber="12" startId="11.[136,170,1159,1179]" targetBox="[134,1414,249,1131]" targetPageId="11" targetType="figure">
<paragraph id="8BC236F69537B84DFF0AFB2BC962FB36" blockId="11.[136,1414,1159,1207]" pageId="11" pageNumber="12">
<emphasis id="B909EAE49537B84DFF0AFB2BCAAEFB1A" bold="true" box="[136,212,1159,1179]" pageId="11" pageNumber="12">Fig. 10.</emphasis>
<taxonomicName id="4C7D4D759537B84DFF61FB2BCB81FB1A" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[227,507,1159,1179]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49537B84DFF61FB2BCB81FB1A" box="[227,507,1159,1179]" italics="true" pageId="11" pageNumber="12">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9537B84DFD81FB2BC840FB1A" box="[515,570,1159,1179]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
Male paratype. Idiosoma: A- dorsum, B- venter; gnathosoma: C- dorsum, D- venter; ascalebar for idiosoma, b- scalebar for gnathosoma; sculpture omitted.
</paragraph>
</caption>
<footnote id="E8662AF89537B84DFE17F8DECBC8F807" box="[405,434,1906,1926]" pageId="11" pageNumber="12">
<paragraph id="8BC236F69537B84DFE17F8DECBC8F807" blockId="11.[405,434,1906,1926]" box="[405,434,1906,1926]" pageId="11" pageNumber="12">(A)</paragraph>
</footnote>
<footnote id="E8662AF89537B84DFBBBF8DECE2FF807" box="[1081,1109,1906,1926]" pageId="11" pageNumber="12">
<paragraph id="8BC236F69537B84DFBBBF8DECE2FF807" blockId="11.[1081,1109,1906,1926]" box="[1081,1109,1906,1926]" pageId="11" pageNumber="12">(B)</paragraph>
</footnote>
<caption id="DF02667E9537B84DFF0AF833CEFEF832" ID-DOI="http://doi.org/10.5281/zenodo.12956967" ID-Zenodo-Dep="12956967" box="[136,1156,1951,1971]" httpUri="https://zenodo.org/record/12956967/files/figure.png" pageId="11" pageNumber="12" startId="11.[136,170,1951,1971]" targetBox="[137,1416,1266,1881]" targetPageId="11" targetType="figure">
<paragraph id="8BC236F69537B84DFF0AF833CEFEF832" blockId="11.[136,1156,1951,1971]" box="[136,1156,1951,1971]" pageId="11" pageNumber="12">
<emphasis id="B909EAE49537B84DFF0AF833CAAAF832" bold="true" box="[136,208,1951,1971]" pageId="11" pageNumber="12">Fig. 11.</emphasis>
<taxonomicName id="4C7D4D759537B84DFF5EF833CB88F832" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[220,498,1951,1971]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49537B84DFF5EF833CB88F832" box="[220,498,1951,1971]" italics="true" pageId="11" pageNumber="12">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9537B84DFE7AF833C856F832" box="[504,556,1951,1971]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
Male paratype. Genital capsule: A- dorsal view, B. ventral view.
</paragraph>
</caption>
<paragraph id="8BC236F69530B84AFF2CFF04CB8EFF3E" blockId="12.[174,500,168,191]" box="[174,500,168,191]" pageId="12" pageNumber="13">
<bibCitation id="5E573E9A9530B84AFF2CFF04CB8EFF3E" box="[174,500,168,191]" journal="Zoological Studies" page="11" pageId="12" pageNumber="13" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49530B84AFF2CFF04CB1DFF3E" box="[174,359,168,191]" italics="true" pageId="12" pageNumber="13">Zoological Studies</emphasis>
<emphasis id="B909EAE49530B84AFEEFFF04CBF4FF3E" bold="true" box="[365,398,168,191]" pageId="12" pageNumber="13">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69530B84AFF2CFF51CB1BFEB7" blockId="12.[174,775,253,726]" pageId="12" pageNumber="13">
<emphasis id="B909EAE49530B84AFF2CFF51CAB3FE97" box="[174,201,253,278]" italics="true" pageId="12" pageNumber="13">tc’</emphasis>
~ 1½ 
<emphasis id="B909EAE49530B84AFE9CFF51CB46FE97" box="[286,316,253,278]" italics="true" pageId="12" pageNumber="13">tc”</emphasis>
, both setae barbed. Seta 
<emphasis id="B909EAE49530B84AFD01FF51C8E1FE97" box="[643,667,253,278]" italics="true" pageId="12" pageNumber="13">u”</emphasis>
inserted laterad of claw.
</paragraph>
<paragraph id="8BC236F69530B84AFF68FE91C84AFED7" blockId="12.[174,775,253,726]" box="[234,560,317,342]" pageId="12" pageNumber="13">
Larva (
<figureCitation id="13462A739530B84AFEB9FE91CBC5FED7" box="[315,447,317,342]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="14.[174,208,1953,1973]" captionStartId-1="15.[137,171,1953,1973]" captionStartId-2="16.[174,208,1951,1971]" captionStartId-3="17.[137,171,1950,1970]" captionStartId-4="18.[174,208,1953,1973]" captionTargetBox-0="[171,1451,705,1917]" captionTargetBox-1="[159,1402,1038,1928]" captionTargetBox-2="[292,1330,709,1916]" captionTargetId-0="figure-227@14.[159,1461,689,1927]" captionTargetId-1="figure-401@15.[136,1442,1029,1949]" captionTargetId-2="figure-149@16.[281,1344,690,1927]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="Fig. 16. Zambedania sekhukhunensis sp. n. Larva paratype. Idiosoma: A- dorsum, B- venter." captionText-1="Fig. 17. Zambedania sekhukhunensis sp. n. Larva paratype. Gnathosoma: A- dorsal view, B. ventral view, C- pharyngeal pump system." captionText-2="Fig. 18. Zambedania sekhukhunensis sp. n. Larva paratype. Leg I (right): dorsal view." captionText-3="Fig. 19. Zambedania sekhukhunensis sp. n. Larva paratype. Leg II (right): dorsal view." captionText-4="Fig. 20. Zambedania sekhukhunensis sp. n. Larva paratype. Leg III (right): dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.12956993" figureDoi-1="http://doi.org/10.5281/zenodo.12956997" figureDoi-2="http://doi.org/10.5281/zenodo.12957009" httpUri-0="https://zenodo.org/record/12956993/files/figure.png" httpUri-1="https://zenodo.org/record/12956997/files/figure.png" httpUri-2="https://zenodo.org/record/12957009/files/figure.png" pageId="12" pageNumber="13">Figs 16-20</figureCitation>
, 
<tableCitation id="C6FF034D9530B84AFE48FE91C85EFED7" box="[458,548,317,342]" captionStart="Table 4" captionStartId="10.[174,242,1479,1504]" captionText="Table 4. Leg chaeto- and solenidiotaxy of the genus Zambedania sekhukhunensis sp. n. larva" pageId="12" pageNumber="13">Table 4</tableCitation>
).
</paragraph>
<paragraph id="8BC236F69530B84AFF68FEF1CFD1FF3E" blockId="12.[174,775,253,726]" lastBlockId="12.[1313,1451,168,191]" pageId="12" pageNumber="13">
Gnathosoma (
<figureCitation id="13462A739530B84AFE10FEF1C850FEF7" box="[402,554,349,374]" captionStart="Fig" captionStartId="15.[137,171,1953,1973]" captionTargetBox="[159,1402,1038,1928]" captionTargetId="figure-401@15.[136,1442,1029,1949]" captionTargetPageId="15" captionText="Fig. 17. Zambedania sekhukhunensis sp. n. Larva paratype. Gnathosoma: A- dorsal view, B. ventral view, C- pharyngeal pump system." figureDoi="http://doi.org/10.5281/zenodo.12956997" httpUri="https://zenodo.org/record/12956997/files/figure.png" pageId="12" pageNumber="13">Figs. 17A, B</figureCitation>
). The gnathosoma about the same size and shape as that of the adult female. Dorsally the stylophore terminates in a trapezoid antero-distal edge. The pedipalps situated antero-laterad and subdivided into four segments, palptrochanter, fused palpfemoro-genu with smooth setae 
<emphasis id="B909EAE49530B84AFD8BFDB1C841FDB7" box="[521,571,541,566]" italics="true" pageId="12" pageNumber="13">dFe</emphasis>
and 
<emphasis id="B909EAE49530B84AFD0AFDB1C8C5FDB7" box="[648,703,541,566]" italics="true" pageId="12" pageNumber="13">dGe</emphasis>
, and palptibiotarsus. The palptrochanter fused with the stylophore. The palptrochanter and palpfemoro-genu connected by membranes as are the palpfemoro-genu and the palptibiotarsus. The palptibiotarsus, laterad of the stylophore clearly page 13 of 21
</paragraph>
<paragraph id="8BC236F69530B84AFCD0FF51CFE6FDB7" blockId="12.[850,1452,253,726]" pageId="12" pageNumber="13">
subdivided into three parts. The distal claw medially with thorn-like structures. Stylophore with very short, blunt peglike 
<emphasis id="B909EAE49530B84AFB14FE91CEC7FED4" box="[1174,1213,317,342]" italics="true" pageId="12" pageNumber="13">
ch 
<subScript id="17F934B39530B84AFB36FEEACEC7FED4" attach="left" box="[1204,1213,326,341]" fontSize="6" pageId="12" pageNumber="13">1</subScript>
</emphasis>
setae and slightly postero-laterad thereof smooth setiform 
<emphasis id="B909EAE49530B84AFAB0FEF1CF2DFEF4" box="[1330,1367,349,374]" italics="true" pageId="12" pageNumber="13">
ch 
<subScript id="17F934B39530B84AFACCFECACF2DFEF4" attach="left" box="[1358,1367,358,373]" fontSize="6" pageId="12" pageNumber="13">2</subScript>
</emphasis>
, which is about three and a half times the length of 
<emphasis id="B909EAE49530B84AFAFCFED1CFD1FE17" box="[1406,1451,381,406]" italics="true" pageId="12" pageNumber="13">
ch 
<subScript id="17F934B39530B84AFA19FE2ACFDEFE14" attach="right" box="[1435,1444,390,405]" fontSize="6" pageId="12" pageNumber="13">1</subScript>
.
</emphasis>
Palpal supracoxal setae 
<emphasis id="B909EAE49530B84AFBFDFE31CEE7FE37" box="[1151,1181,413,438]" italics="true" pageId="12" pageNumber="13">pp</emphasis>
absent. Subcapitulum with a pair of slender, simple, smooth subcapitular setae (
<emphasis id="B909EAE49530B84AFC2EFE71C9B3FE77" box="[940,969,477,502]" italics="true" pageId="12" pageNumber="13">su</emphasis>
); antero-mediad a mushroom-shaped setigenous accessory structure (
<emphasis id="B909EAE49530B84AFB61FE51CF77FD97" box="[1251,1293,509,534]" italics="true" pageId="12" pageNumber="13">ass</emphasis>
) and laterad at its base a clavate, non-striated solenidion (
<emphasis id="B909EAE49530B84AFAE8FDB1CFF6FDB7" box="[1386,1420,541,566]" italics="true" pageId="12" pageNumber="13">sol</emphasis>
).
</paragraph>
<paragraph id="8BC236F69530B84AFC0DFD91C9C5FD57" blockId="12.[850,1452,253,726]" pageId="12" pageNumber="13">
Pharyngeal system (
<figureCitation id="13462A739530B84AFB0AFD91CE8EFDD6" box="[1160,1268,573,599]" captionStart="Fig" captionStartId="15.[137,171,1953,1973]" captionTargetBox="[159,1402,1038,1928]" captionTargetId="figure-401@15.[136,1442,1029,1949]" captionTargetPageId="15" captionText="Fig. 17. Zambedania sekhukhunensis sp. n. Larva paratype. Gnathosoma: A- dorsal view, B. ventral view, C- pharyngeal pump system." figureDoi="http://doi.org/10.5281/zenodo.12956997" httpUri="https://zenodo.org/record/12956997/files/figure.png" pageId="12" pageNumber="13">Fig. 17C</figureCitation>
) similar to that of the adult female situated in the propodosomal area. The pharyngeal pumps shaped similarly to those of the adult female, but more compact and very thin.
</paragraph>
<caption id="DF02667E9530B84AFF2CF829CBE2F834" pageId="12" pageNumber="13" startId="12.[174,209,1925,1945]" targetType="figure">
<paragraph id="8BC236F69530B84AFF2CF829CBE2F834" blockId="12.[174,776,1925,1973]" pageId="12" pageNumber="13">
<emphasis id="B909EAE49530B84AFF2CF829CA86F818" bold="true" box="[174,252,1925,1945]" pageId="12" pageNumber="13">Fig. 12.</emphasis>
<taxonomicName id="4C7D4D759530B84AFE8EF829C851F818" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[268,555,1925,1945]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49530B84AFE8EF829C851F818" box="[268,555,1925,1945]" italics="true" pageId="12" pageNumber="13">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9530B84AFDB6F829C816F818" box="[564,620,1925,1945]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
Male paratype. Leg I (right): dorsal view.
</paragraph>
</caption>
<caption id="DF02667E9530B84AFCD0F829CE38F834" ID-DOI="http://doi.org/10.5281/zenodo.12956973" ID-Zenodo-Dep="12956973" httpUri="https://zenodo.org/record/12956973/files/figure.png" pageId="12" pageNumber="13" startId="12.[850,885,1925,1945]" targetBox="[102,1452,779,1899]" targetPageId="12" targetType="figure">
<paragraph id="8BC236F69530B84AFCD0F829CE38F834" blockId="12.[850,1453,1925,1973]" pageId="12" pageNumber="13">
<emphasis id="B909EAE49530B84AFCD0F829C9DAF818" bold="true" box="[850,928,1925,1945]" pageId="12" pageNumber="13">Fig. 13.</emphasis>
<taxonomicName id="4C7D4D759530B84AFC33F829CEAAF818" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[945,1232,1925,1945]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49530B84AFC33F829CEAAF818" box="[945,1232,1925,1945]" italics="true" pageId="12" pageNumber="13">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9530B84AFB5AF829CF6AF818" box="[1240,1296,1925,1945]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
Male paratype. Leg II (right): dorsal view.
</paragraph>
</caption>
<paragraph id="8BC236F69531B84BFFFFFF04CBBEFF3E" blockId="13.[125,452,168,191]" box="[125,452,168,191]" pageId="13" pageNumber="14">
<bibCitation id="5E573E9A9531B84BFFFFFF04CBBEFF3E" box="[125,452,168,191]" journal="Zoological Studies" page="11" pageId="13" pageNumber="14" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49531B84BFFFFFF04CB4DFF3E" box="[125,311,168,191]" italics="true" pageId="13" pageNumber="14">Zoological Studies</emphasis>
<emphasis id="B909EAE49531B84BFEBFFF04CB24FF3E" bold="true" box="[317,350,168,191]" pageId="13" pageNumber="14">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69531B84BFF47FF51CFFFFF3E" blockId="13.[136,736,253,374]" lastBlockId="13.[1275,1413,168,191]" pageId="13" pageNumber="14">
Idiosomal dorsum (
<figureCitation id="13462A739531B84BFE45FF51C838FE97" box="[455,578,253,278]" captionStart="Fig" captionStartId="14.[174,208,1953,1973]" captionTargetBox="[171,1451,705,1917]" captionTargetId="figure-227@14.[159,1461,689,1927]" captionTargetPageId="14" captionText="Fig. 16. Zambedania sekhukhunensis sp. n. Larva paratype. Idiosoma: A- dorsum, B- venter." figureDoi="http://doi.org/10.5281/zenodo.12956993" httpUri="https://zenodo.org/record/12956993/files/figure.png" pageId="13" pageNumber="14">Fig. 16A</figureCitation>
). The larval exoskeleton very lightly sclerotized and without clearly defined tergites and sculpture. The PdS with three pairs of tapered setiform setae: one page 14 of 21
</paragraph>
<paragraph id="8BC236F69531B84BFCAFFF51CFFFFDB7" blockId="13.[813,1414,253,567]" pageId="13" pageNumber="14">
pair of short, smooth verticals (
<emphasis id="B909EAE49531B84BFB3FFF51CEB0FE97" box="[1213,1226,253,278]" italics="true" pageId="13" pageNumber="14">v</emphasis>
) anterolaterad of barbed scapulars 
<emphasis id="B909EAE49531B84BFBB2FEB1CE21FEB7" box="[1072,1115,285,310]" italics="true" pageId="13" pageNumber="14">sc1</emphasis>
, followed posterolateral by a very long pair 
<emphasis id="B909EAE49531B84BFBADFE91CE20FED7" box="[1071,1114,317,342]" italics="true" pageId="13" pageNumber="14">sc2</emphasis>
. Bothridia, stigmata or podocephalic canals absent. Opisthosomal tergites fused in pairs: CD (partly), EF, and dorsoventrocaudal HPs. All dorsal opisthosomal setae very long and barbed. Segment C is subdivided into three shields, the median one coalesced with plate D bearing setae 
<emphasis id="B909EAE49531B84BFBB7FE51CE2BFD97" box="[1077,1105,509,534]" italics="true" pageId="13" pageNumber="14">c1</emphasis>
and two separate laterals with 
<emphasis id="B909EAE49531B84BFCEAFDB1C9F6FDB7" box="[872,908,541,566]" italics="true" pageId="13" pageNumber="14">c2.</emphasis>
Setae 
<emphasis id="B909EAE49531B84BFC6AFDB1CE7FFDB7" box="[1000,1029,541,566]" italics="true" pageId="13" pageNumber="14">c1</emphasis>
are situated mediad of 
<emphasis id="B909EAE49531B84BFAB3FDB1CF34FDB7" box="[1329,1358,541,566]" italics="true" pageId="13" pageNumber="14">c2</emphasis>
and
</paragraph>
<caption id="DF02667E9531B84BFF0AF82ACB04F837" ID-DOI="http://doi.org/10.5281/zenodo.12956981" ID-Zenodo-Dep="12956981" httpUri="https://zenodo.org/record/12956981/files/figure.png" pageId="13" pageNumber="14" startId="13.[136,171,1926,1946]" targetBox="[138,736,432,1900]" targetPageId="13" targetType="figure">
<paragraph id="8BC236F69531B84BFF0AF82ACB04F837" blockId="13.[136,738,1926,1974]" pageId="13" pageNumber="14">
<emphasis id="B909EAE49531B84BFF0AF82ACAACF81B" bold="true" box="[136,214,1926,1946]" pageId="13" pageNumber="14">Fig. 14.</emphasis>
<taxonomicName id="4C7D4D759531B84BFF64F82AC87FF81B" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[230,517,1926,1946]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49531B84BFF64F82AC87FF81B" box="[230,517,1926,1946]" italics="true" pageId="13" pageNumber="14">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9531B84BFD8CF82AC83CF81B" box="[526,582,1926,1946]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
Male paratype. Leg III (right): dorsal view.
</paragraph>
</caption>
<caption id="DF02667E9531B84BFCAFF82ACE62F837" ID-DOI="http://doi.org/10.5281/zenodo.12956989" ID-Zenodo-Dep="12956989" httpUri="https://zenodo.org/record/12956989/files/figure.png" pageId="13" pageNumber="14" startId="13.[813,848,1926,1946]" targetBox="[814,1413,638,1898]" targetPageId="13" targetType="figure">
<paragraph id="8BC236F69531B84BFCAFF82ACE62F837" blockId="13.[813,1415,1926,1974]" pageId="13" pageNumber="14">
<emphasis id="B909EAE49531B84BFCAFF82AC900F81B" bold="true" box="[813,890,1926,1946]" pageId="13" pageNumber="14">Fig. 15.</emphasis>
<taxonomicName id="4C7D4D759531B84BFC09F82ACED0F81B" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[907,1194,1926,1946]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49531B84BFC09F82ACED0F81B" box="[907,1194,1926,1946]" italics="true" pageId="13" pageNumber="14">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9531B84BFB30F82ACE90F81B" box="[1202,1258,1926,1946]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
Male paratype. Leg IV (left): dorsal view.
</paragraph>
</caption>
<paragraph id="8BC236F69532B848FF2CFF04CB8EFF3E" blockId="14.[174,500,168,191]" box="[174,500,168,191]" pageId="14" pageNumber="15">
<bibCitation id="5E573E9A9532B848FF2CFF04CB8EFF3E" box="[174,500,168,191]" journal="Zoological Studies" page="11" pageId="14" pageNumber="15" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49532B848FF2CFF04CB1DFF3E" box="[174,359,168,191]" italics="true" pageId="14" pageNumber="15">Zoological Studies</emphasis>
<emphasis id="B909EAE49532B848FEEFFF04CBF4FF3E" bold="true" box="[365,398,168,191]" pageId="14" pageNumber="15">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69532B848FF2CFF51CB74FE37" blockId="14.[174,775,253,630]" pageId="14" pageNumber="15">
anteriad of 
<emphasis id="B909EAE49532B848FEBDFF51CB34FE97" box="[319,334,253,278]" italics="true" pageId="14" pageNumber="15">d</emphasis>
. Setae 
<emphasis id="B909EAE49532B848FE36FF51CBB9FE97" box="[436,451,253,278]" italics="true" pageId="14" pageNumber="15">d</emphasis>
on CD and 
<emphasis id="B909EAE49532B848FDE3FF51C812FE97" box="[609,616,253,278]" italics="true" pageId="14" pageNumber="15">f</emphasis>
on plate EF implanted on protrusions. Setae 
<emphasis id="B909EAE49532B848FDBFFEB1C836FEB7" box="[573,588,285,310]" italics="true" pageId="14" pageNumber="15">e</emphasis>
laterad of seta 
<emphasis id="B909EAE49532B848FF2CFE91CACFFED7" box="[174,181,317,342]" italics="true" pageId="14" pageNumber="15">f</emphasis>
on segment EF. On segment HPs setal pair 
<emphasis id="B909EAE49532B848FD6BFE91C97DFED7" box="[745,775,317,342]" italics="true" pageId="14" pageNumber="15">h1</emphasis>
positioned slightly antero-mediad of 
<emphasis id="B909EAE49532B848FD08FEF1C8D3FEF7" box="[650,681,349,374]" italics="true" pageId="14" pageNumber="15">h2</emphasis>
on the dorsal side, and the pseudanal 
<emphasis id="B909EAE49532B848FDBBFED1C81EFE17" box="[569,612,381,406]" italics="true" pageId="14" pageNumber="15">ps1</emphasis>
, 
<emphasis id="B909EAE49532B848FDF5FED1C8FCFE17" box="[631,646,381,406]" italics="true" pageId="14" pageNumber="15">2</emphasis>
, 
<emphasis id="B909EAE49532B848FD1AFED1C8DDFE17" box="[664,679,381,406]" italics="true" pageId="14" pageNumber="15">3</emphasis>
ventrocaudad.
</paragraph>
<paragraph id="8BC236F69532B848FF68FE11C97DFDF7" blockId="14.[174,775,253,630]" pageId="14" pageNumber="15">
Idiosomal venter (
<figureCitation id="13462A739532B848FE5CFE11C822FE57" box="[478,600,445,470]" captionStart="Fig" captionStartId="14.[174,208,1953,1973]" captionTargetBox="[171,1451,705,1917]" captionTargetId="figure-227@14.[159,1461,689,1927]" captionTargetPageId="14" captionText="Fig. 16. Zambedania sekhukhunensis sp. n. Larva paratype. Idiosoma: A- dorsum, B- venter." figureDoi="http://doi.org/10.5281/zenodo.12956993" httpUri="https://zenodo.org/record/12956993/files/figure.png" pageId="14" pageNumber="15">Fig. 16B</figureCitation>
). Coxisternal plates I and II lightly sclerotized bearing two pairs of setae on each plate (
<emphasis id="B909EAE49532B848FDA7FE51C839FD97" box="[549,579,509,534]" italics="true" pageId="14" pageNumber="15">1a</emphasis>
, 
<emphasis id="B909EAE49532B848FDD4FE51C80EFD97" box="[598,628,509,534]" italics="true" pageId="14" pageNumber="15">1b</emphasis>
and 
<emphasis id="B909EAE49532B848FD3AFE51C8ACFD97" box="[696,726,509,534]" italics="true" pageId="14" pageNumber="15">2a</emphasis>
, 
<emphasis id="B909EAE49532B848FD6BFE51C97CFD97" box="[745,774,509,534]" italics="true" pageId="14" pageNumber="15">2b</emphasis>
respectively). Anterior edges of the plates form apodemes (ap) 1 and 2, medially separated from each other and do not form the prosternal (appr)
</paragraph>
<paragraph id="8BC236F69532B848FAA3FF04CFD1FF3E" blockId="14.[1313,1451,168,191]" box="[1313,1451,168,191]" pageId="14" pageNumber="15">page 15 of 21</paragraph>
<paragraph id="8BC236F69532B848FCD0FF51CF2DFE17" blockId="14.[850,1452,253,631]" pageId="14" pageNumber="15">
apodeme. Sejugal apodeme (apsej) absent. CX III with two pairs of setae (
<emphasis id="B909EAE49532B848FB5AFEB1CE8CFEB7" box="[1240,1270,285,310]" italics="true" pageId="14" pageNumber="15">3a</emphasis>
, 
<emphasis id="B909EAE49532B848FA8BFEB1CF5DFEB7" box="[1289,1319,285,310]" italics="true" pageId="14" pageNumber="15">3b</emphasis>
). Ap3 and poststernal apodeme (appo) absent. In place of appo, some creases between CX III plates present. All ventral setae short, tapered and smooth.
</paragraph>
<paragraph id="8BC236F69532B848FC0DFE31CFD1FDF7" blockId="14.[850,1452,253,631]" pageId="14" pageNumber="15">
Legs. Leg I (
<figureCitation id="13462A739532B848FBA3FE31CE02FE37" box="[1057,1144,413,438]" captionStart="Fig" captionStartId="16.[174,208,1951,1971]" captionTargetBox="[292,1330,709,1916]" captionTargetId="figure-149@16.[281,1344,690,1927]" captionTargetPageId="16" captionText="Fig. 18. Zambedania sekhukhunensis sp. n. Larva paratype. Leg I (right): dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12957009" httpUri="https://zenodo.org/record/12957009/files/figure.png" pageId="14" pageNumber="15">Fig. 18</figureCitation>
). Femur: barbed 
<emphasis id="B909EAE49532B848FAC2FE31CF35FE37" box="[1344,1359,413,438]" italics="true" pageId="14" pageNumber="15">d</emphasis>
longest seta on segment, slightly barbed 
<emphasis id="B909EAE49532B848FB75FE11CF67FE57" box="[1271,1309,445,470]" italics="true" pageId="14" pageNumber="15">l’&gt;</emphasis>
smooth 
<emphasis id="B909EAE49532B848FA0FFE11CFDEFE57" box="[1421,1444,445,470]" italics="true" pageId="14" pageNumber="15">v”</emphasis>
, both on pinnaculum. Genu: all setae barbed, except smooth 
<emphasis id="B909EAE49532B848FB8DFE51CE5CFD97" box="[1039,1062,509,534]" italics="true" pageId="14" pageNumber="15">v”</emphasis>
&lt;
<emphasis id="B909EAE49532B848FBC8FE51CE27FD97" box="[1098,1117,509,534]" italics="true" pageId="14" pageNumber="15">v’</emphasis>
; 
<emphasis id="B909EAE49532B848FBF2FE51CE06FD97" box="[1136,1148,509,534]" italics="true" pageId="14" pageNumber="15">l’</emphasis>
(on pinnaculum) longest seta and 
<emphasis id="B909EAE49532B848FC43FDB1C9B5FDB7" box="[961,975,541,566]" italics="true" pageId="14" pageNumber="15">l”</emphasis>
the shortest; all tibial setae on pinnaculi and smooth except feathered 
<emphasis id="B909EAE49532B848FB40FD91CEAFFDD7" box="[1218,1237,573,598]" italics="true" pageId="14" pageNumber="15">v’</emphasis>
: 
<emphasis id="B909EAE49532B848FB65FD91CFE8FDD7" box="[1255,1426,573,598]" italics="true" pageId="14" pageNumber="15">v”&gt; l’&gt; l”&gt; d</emphasis>
~ 
<emphasis id="B909EAE49532B848FCD0FDF1C925FDF7" box="[850,863,605,630]" italics="true" pageId="14" pageNumber="15">k</emphasis>
. Striation of solenidion Φ wider than of ω (both
</paragraph>
<caption id="DF02667E9532B848FF2CF80DCE65F834" ID-DOI="http://doi.org/10.5281/zenodo.12956993" ID-Zenodo-Dep="12956993" box="[174,1055,1953,1973]" httpUri="https://zenodo.org/record/12956993/files/figure.png" pageId="14" pageNumber="15" startId="14.[174,208,1953,1973]" targetBox="[171,1451,705,1917]" targetPageId="14" targetType="figure">
<paragraph id="8BC236F69532B848FF2CF80DCE65F834" blockId="14.[174,1055,1953,1973]" box="[174,1055,1953,1973]" pageId="14" pageNumber="15">
<emphasis id="B909EAE49532B848FF2CF80DCA82F834" bold="true" box="[174,248,1953,1973]" pageId="14" pageNumber="15">Fig. 16.</emphasis>
<taxonomicName id="4C7D4D759532B848FE81F80DC863F834" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[259,537,1953,1973]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49532B848FE81F80DC863F834" box="[259,537,1953,1973]" italics="true" pageId="14" pageNumber="15">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9532B848FD9DF80DC829F834" box="[543,595,1953,1973]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
Larva paratype. Idiosoma: A- dorsum, B- venter.
</paragraph>
</caption>
<paragraph id="8BC236F69533B849FFFFFF04CBBEFF3E" blockId="15.[125,452,168,191]" box="[125,452,168,191]" pageId="15" pageNumber="16">
<bibCitation id="5E573E9A9533B849FFFFFF04CBBEFF3E" box="[125,452,168,191]" journal="Zoological Studies" page="11" pageId="15" pageNumber="16" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE49533B849FFFFFF04CB4DFF3E" box="[125,311,168,191]" italics="true" pageId="15" pageNumber="16">Zoological Studies</emphasis>
<emphasis id="B909EAE49533B849FEBFFF04CB24FF3E" bold="true" box="[317,350,168,191]" pageId="15" pageNumber="16">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F69533B849FF0AFF51CAA9FE37" blockId="15.[133,738,253,950]" pageId="15" pageNumber="16">
on pinnaculi). Tarsal setae all smooth except little barbed 
<emphasis id="B909EAE49533B849FF61FEB1CA84FEB7" box="[227,254,285,310]" italics="true" pageId="15" pageNumber="16">pl’</emphasis>
; 
<emphasis id="B909EAE49533B849FE8EFEB1CB18FEB7" box="[268,354,285,310]" italics="true" pageId="15" pageNumber="16">pl’ ~ pl”</emphasis>
; 
<emphasis id="B909EAE49533B849FEF3FEB1CBEDFEB7" box="[369,407,285,310]" italics="true" pageId="15" pageNumber="16">pv”</emphasis>
&gt; 
<emphasis id="B909EAE49533B849FE37FEB1CBADFEB7" box="[437,471,285,310]" italics="true" pageId="15" pageNumber="16">pv’</emphasis>
; 
<emphasis id="B909EAE49533B849FE64FEB1C8C5FEB7" box="[486,703,285,310]" italics="true" pageId="15" pageNumber="16">ft’~ ft”&gt; s&gt; u’&gt; u”</emphasis>
; 
<emphasis id="B909EAE49533B849FD4CFEB1CA8FFED7" italics="true" pageId="15" pageNumber="16">ft’ &lt;tc’ &lt;tc”</emphasis>
. Setae 
<emphasis id="B909EAE49533B849FED2FE91CB08FED7" box="[336,370,317,342]" italics="true" pageId="15" pageNumber="16">pv’</emphasis>
, 
<emphasis id="B909EAE49533B849FE03FE91CBE5FED7" box="[385,415,317,342]" italics="true" pageId="15" pageNumber="16">pl”</emphasis>
, 
<emphasis id="B909EAE49533B849FE2CFE91CBB9FED7" box="[430,451,317,342]" italics="true" pageId="15" pageNumber="16">ft’</emphasis>
, 
<emphasis id="B909EAE49533B849FE50FE91CB91FED7" box="[466,491,317,342]" italics="true" pageId="15" pageNumber="16">tc’</emphasis>
and 
<emphasis id="B909EAE49533B849FDA4FE91C83EFED7" box="[550,580,317,342]" italics="true" pageId="15" pageNumber="16">tc”</emphasis>
on pinnaculi. On tarsus two distal claws present (only one on the right leg- artefact); no empodium between the claws.
</paragraph>
<paragraph id="8BC236F69533B849FF47FE11C808FC97" blockId="15.[133,738,253,950]" pageId="15" pageNumber="16">
Leg II (
<figureCitation id="13462A739533B849FEA7FE11CBF2FE57" box="[293,392,445,470]" captionStart="Fig" captionStartId="17.[137,171,1950,1970]" captionText="Fig. 19. Zambedania sekhukhunensis sp. n. Larva paratype. Leg II (right): dorsal view." pageId="15" pageNumber="16">Fig. 19</figureCitation>
). Femur: slightly barbed 
<emphasis id="B909EAE49533B849FD57FE11CAE2FE77" italics="true" pageId="15" pageNumber="16">l’&gt;</emphasis>
smooth 
<emphasis id="B909EAE49533B849FE87FE71CB66FE77" box="[261,284,477,502]" italics="true" pageId="15" pageNumber="16">v”</emphasis>
(both on pinnaculi), 
<emphasis id="B909EAE49533B849FDB8FE71C833FE77" box="[570,585,477,502]" italics="true" pageId="15" pageNumber="16">d</emphasis>
longest on segment and barbed. Genu: 
<emphasis id="B909EAE49533B849FD84FE51C835FD97" box="[518,591,509,534]" italics="true" pageId="15" pageNumber="16">v’ ~ l’</emphasis>
, both one-sidedly barbed; slightly barbed 
<emphasis id="B909EAE49533B849FD98FDB1C850FDB7" box="[538,554,541,566]" italics="true" pageId="15" pageNumber="16">l”</emphasis>
shortest seta. Tibia: one-sidedly barbed 
<emphasis id="B909EAE49533B849FE57FD91CB81FDD7" box="[469,507,573,598]" italics="true" pageId="15" pageNumber="16">l’&gt;</emphasis>
slightly barbed 
<emphasis id="B909EAE49533B849FD48FD91C8A3FDD7" box="[714,729,573,598]" italics="true" pageId="15" pageNumber="16">d</emphasis>
; pilose 
<emphasis id="B909EAE49533B849FF54FDF1CA93FDF7" box="[214,233,605,630]" italics="true" pageId="15" pageNumber="16">v’</emphasis>
&lt;one-sidedly barbed 
<emphasis id="B909EAE49533B849FE75FDF1C874FDF7" box="[503,526,605,630]" italics="true" pageId="15" pageNumber="16">v”</emphasis>
; solenidion 
<emphasis id="B909EAE49533B849FD1CFDF1C8DEFDF7" box="[670,676,605,630]" italics="true" pageId="15" pageNumber="16">j</emphasis>
&lt;Ta ω. Tarsus: smooth 
<emphasis id="B909EAE49533B849FEF0FDD1CBD1FD17" box="[370,427,637,662]" italics="true" pageId="15" pageNumber="16">pl”&gt;</emphasis>
smooth 
<emphasis id="B909EAE49533B849FD9BFDD1C837FD17" box="[537,589,637,662]" italics="true" pageId="15" pageNumber="16">tc’&gt;</emphasis>
smooth 
<emphasis id="B909EAE49533B849FD39FDD1C8A3FD17" box="[699,729,637,662]" italics="true" pageId="15" pageNumber="16">tc”</emphasis>
; slightly barbed 
<emphasis id="B909EAE49533B849FECAFD31CBFEFD37" box="[328,388,669,694]" italics="true" pageId="15" pageNumber="16">pv’&gt;</emphasis>
smooth 
<emphasis id="B909EAE49533B849FE71FD31C863FD37" box="[499,537,669,694]" italics="true" pageId="15" pageNumber="16">pv”</emphasis>
; slightly barbed 
<emphasis id="B909EAE49533B849FF0AFD11CAA4FD57" box="[136,222,701,726]" italics="true" pageId="15" pageNumber="16">u’&gt; u”</emphasis>
. Solenidion ω large. Two claws with a spatulate empodium in between. Tibial setae 
<emphasis id="B909EAE49533B849FD25FD71C8CCFD77" box="[679,694,733,758]" italics="true" pageId="15" pageNumber="16">d</emphasis>
, 
<emphasis id="B909EAE49533B849FD44FD71C8A3FD77" box="[710,729,733,758]" italics="true" pageId="15" pageNumber="16">v’</emphasis>
, 
<emphasis id="B909EAE49533B849FF0AFD51CAE5FC97" box="[136,159,765,790]" italics="true" pageId="15" pageNumber="16">v”</emphasis>
, Φ and tarsal 
<emphasis id="B909EAE49533B849FEBDFD51CB27FC97" box="[319,349,765,790]" italics="true" pageId="15" pageNumber="16">pl”</emphasis>
, 
<emphasis id="B909EAE49533B849FEE9FD51CBFCFC97" box="[363,390,765,790]" italics="true" pageId="15" pageNumber="16">tc’</emphasis>
and ω on pinnaculi.
</paragraph>
<paragraph id="8BC236F69533B849FF47FCB1CFFFFF3E" blockId="15.[133,738,253,950]" lastBlockId="15.[1275,1413,168,191]" pageId="15" pageNumber="16">
Leg III (
<figureCitation id="13462A739533B849FEAEFCB1CBF4FCB7" box="[300,398,797,822]" captionStart="Fig" captionStartId="18.[174,208,1953,1973]" captionText="Fig. 20. Zambedania sekhukhunensis sp. n. Larva paratype. Leg III (right): dorsal view." pageId="15" pageNumber="16">Fig. 20</figureCitation>
). Femur: barbed 
<emphasis id="B909EAE49533B849FDF3FCB1C8FFFCB7" box="[625,645,797,822]" italics="true" pageId="15" pageNumber="16">v’</emphasis>
&lt;long one-sidedly feathered 
<emphasis id="B909EAE49533B849FE23FC91CBCAFCD7" box="[417,432,829,854]" italics="true" pageId="15" pageNumber="16">d</emphasis>
(on pinnaculum). Genu: smooth 
<emphasis id="B909EAE49533B849FF68FCF1CB6CFCF7" box="[234,278,861,886]" italics="true" pageId="15" pageNumber="16">v’ &lt;</emphasis>
one-sidedly barbed 
<emphasis id="B909EAE49533B849FD90FCF1C864FCF7" box="[530,542,861,886]" italics="true" pageId="15" pageNumber="16">l’</emphasis>
. Tibia: all setae one-sidedly barbed: 
<emphasis id="B909EAE49533B849FEF9FCD1CBC3FC17" box="[379,441,893,918]" italics="true" pageId="15" pageNumber="16">l’ &lt;v’</emphasis>
&lt;
<emphasis id="B909EAE49533B849FE5BFCD1CB8AFC17" box="[473,496,893,918]" italics="true" pageId="15" pageNumber="16">v”</emphasis>
&lt;
<emphasis id="B909EAE49533B849FD8DFCD1C864FC17" box="[527,542,893,918]" italics="true" pageId="15" pageNumber="16">d</emphasis>
which is closest to solenidion 
<emphasis id="B909EAE49533B849FEA7FC31CB51FC37" box="[293,299,925,950]" italics="true" pageId="15" pageNumber="16">j</emphasis>
. Tarsus: smooth 
<emphasis id="B909EAE49533B849FE77FC31C850FC37" box="[501,554,925,950]" italics="true" pageId="15" pageNumber="16">pl” &lt;</emphasis>
smooth, robust page 16 of 21
</paragraph>
<paragraph id="8BC236F69533B849FCAFFF51CE40FED7" blockId="15.[813,1415,253,950]" pageId="15" pageNumber="16">
<emphasis id="B909EAE49533B849FCAFFF51C91AFE97" box="[813,864,253,278]" italics="true" pageId="15" pageNumber="16">tc’&gt;</emphasis>
smooth 
<emphasis id="B909EAE49533B849FC4FFF51C991FE97" box="[973,1003,253,278]" italics="true" pageId="15" pageNumber="16">tc”</emphasis>
; slightly barbed 
<emphasis id="B909EAE49533B849FB3AFF51CE8EFE97" box="[1208,1268,253,278]" italics="true" pageId="15" pageNumber="16">pv’&gt;</emphasis>
smooth 
<emphasis id="B909EAE49533B849FAE2FF51CFFCFE97" box="[1376,1414,253,278]" italics="true" pageId="15" pageNumber="16">pv”</emphasis>
&gt; smooth 
<emphasis id="B909EAE49533B849FC2CFEB1CE72FEB7" box="[942,1032,285,310]" italics="true" pageId="15" pageNumber="16">u’ ~ u”.</emphasis>
Two claws with a fan-shaped empodium in between.
</paragraph>
<paragraph id="8BC236F69533B849FCEBFEF1C90FFDD7" blockId="15.[813,1415,253,950]" pageId="15" pageNumber="16">
<emphasis id="B909EAE49533B849FCEBFEF1CE3EFEF7" box="[873,1092,349,374]" italics="true" pageId="15" pageNumber="16">Material examined</emphasis>
: 
<typeStatus id="54C688549533B849FBD6FEF1CEC7FEF7" box="[1108,1213,349,374]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
adult female, 
<specimenCount id="9D7BFD7F9533B849FAEAFEF1C9AEFE17" count="28" pageId="15" pageNumber="16" type="adult" typeStatus="paratype">28 paratype adult</specimenCount>
females, 
<specimenCount id="9D7BFD7F9533B849FBC5FED1CF7EFE17" box="[1095,1284,381,406]" count="5" pageId="15" pageNumber="16" type="adult" typeStatus="paratype">5 paratype adult</specimenCount>
males and 
<specimenCount id="9D7BFD7F9533B849FCAFFE31C98FFE37" box="[813,1013,413,438]" count="1" pageId="15" pageNumber="16" type="larva" typeStatus="paratype">1 paratype larva</specimenCount>
. Females primarily aggregated in and around the fovea (
<figureCitation id="13462A739533B849FBBEFE11CEF8FE57" box="[1084,1154,445,470]" captionStart="Fig" captionStartId="2.[174,208,1924,1944]" captionTargetBox="[191,1435,1067,1896]" captionTargetId="figure-360@2.[188,1436,1064,1898]" captionTargetPageId="2" captionText="Fig. 2. Zambedania sekhukhunensis sp. n phoretic female mites aggregated in and around the fovea of the cephalothorax and around the coxae of Harpactirella overdijki Gallon, 2010 (Aranaeae: Theraphosidae) (photo by I. Engelbrecht)." figureDoi="http://doi.org/10.5281/zenodo.12956911" httpUri="https://zenodo.org/record/12956911/files/figure.png" pageId="15" pageNumber="16">Fig. 2</figureCitation>
) of the cephalothorax and around the coxae, near the spinnerets and anus of spiders; in addition, females found with associated males and larva in and around spider’s nests.
</paragraph>
<paragraph id="8BC236F69533B849FCEBFDF1CFFFFC37" blockId="15.[813,1415,253,950]" pageId="15" pageNumber="16">
<emphasis id="B909EAE49533B849FCEBFDF1C9D9FDF7" box="[873,931,605,630]" italics="true" pageId="15" pageNumber="16">Host</emphasis>
: 
<taxonomicName id="4C7D4D759533B849FC3AFDF1CFFFFDF7" authority="Gallon, 2010" authorityName="Gallon" authorityYear="2010" box="[952,1413,605,631]" class="Arachnida" family="Theraphosidae" genus="Harpactirella" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="overdijki">
<emphasis id="B909EAE49533B849FC3AFDF1CEA9FDF7" box="[952,1235,605,630]" italics="true" pageId="15" pageNumber="16">Harpactirella overdijki</emphasis>
<bibRefCitation id="EFEC4B079533B849FB5DFDF1CFFFFDF7" author="Gallon RC" box="[1247,1413,605,631]" pageId="15" pageNumber="16" pagination="29 - 48" refId="ref9345" refString="Gallon RC. 2010. On some Southern African Harpactirinae, with notes on the eumenophorines Pelinobius muticus Karsch, 1885 and Monoen tropella sp. Strand, 1907 (Araneae, Theraphosidae). Arachnology 15: 29 - 48." type="journal article" year="2010">Gallon, 2010</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C7D4D759533B849FCB1FDD1C9E5FD17" box="[819,927,637,662]" class="Pilidiophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Nemertea" rank="order">Araneae</taxonomicName>
: 
<taxonomicName id="4C7D4D759533B849FC28FDD1CE19FD17" box="[938,1123,637,662]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Theraphosidae</taxonomicName>
). Specimens were found on 
<specimenCount id="9D7BFD7F9533B849FCD7FD31C9B7FD37" box="[853,973,669,694]" count="4" pageId="15" pageNumber="16" type="female">4 females</specimenCount>
and 
<specimenCount id="9D7BFD7F9533B849FB8CFD31CEE2FD37" box="[1038,1176,669,694]" count="2" pageId="15" pageNumber="16" type="immature">2 immature</specimenCount>
spiders. Site: most spiders were collected under a large tree (
<figureCitation id="13462A739533B849FAACFD11CF09FD57" box="[1326,1395,701,726]" captionStart="Fig" captionStartId="1.[136,170,1925,1945]" captionTargetBox="[139,1414,942,1899]" captionTargetId="figure-0@1.[134,1417,937,1904]" captionTargetPageId="1" captionText="Fig. 1. Soon to be exploited North Open Pit (NOP) area at the Two Rivers Platinum Mine at Leydenburg, South Africa (photo by P. Hawkes)." figureDoi="http://doi.org/10.5281/zenodo.12956897" httpUri="https://zenodo.org/record/12956897/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
); 
<quantity id="4C859B139533B849FCAFFD71C90BFD77" box="[813,881,733,758]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="m" value="200.0">200m</quantity>
transect; midpoint of transect: 
<geoCoordinate id="EE4950319533B849FB7FFD71CFFBFD76" box="[1277,1409,733,759]" degrees="24.9086" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="5" value="-24.9086">24.9086 S</geoCoordinate>
, 
<geoCoordinate id="EE4950319533B849FCAFFD51C9C1FC97" box="[813,955,765,790]" degrees="30.09971" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="30.09971">30.09971 E</geoCoordinate>
; altitude 
<quantity id="4C859B139533B849FBA8FD51CE0CFC97" box="[1066,1142,765,790]" metricMagnitude="2" metricUnit="m" metricValue="9.45" pageId="15" pageNumber="16" unit="m" value="945.0">945 m</quantity>
(range 938-950 along transect). Locality: Two Rivers Platinum Mine, 
<quantity id="4C859B139533B849FAEAFCB1C92BFCD7" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="16" unit="km" value="38.0">38 km</quantity>
SE of Burgersfort, 
<quantity id="4C859B139533B849FBBFFC91CEF2FCD7" box="[1085,1160,829,854]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="km" value="40.0">40 km</quantity>
WNW of Lydenburg, Sekhukhuneland, 
<collectingRegion id="49B9F8149533B849FB86FCF1CE9AFCF7" box="[1028,1248,861,886]" country="South Africa" name="Limpopo" pageId="15" pageNumber="16">Limpopo Province</collectingRegion>
, 
<collectingCountry id="F36A76669533B849FB6EFCF1CFF9FCF7" box="[1260,1411,861,887]" name="South Africa" pageId="15" pageNumber="16">South Africa</collectingCountry>
. Dates: 25 March - 25 April 2008, 
<date id="FFC310369533B849FB56FCD1C90BFC37" pageId="15" pageNumber="16" value="2008-11-03" valueMax="2008-11-07" valueMin="2008-11-03">3-7 November 2008</date>
, 
<date id="FFC310369533B849FC03FC31CE4FFC37" box="[897,1077,925,950]" pageId="15" pageNumber="16" value="2010-04-19">19 April 2010</date>
, 
<date id="FFC310369533B849FBC7FC31CF30FC37" box="[1093,1354,925,950]" pageId="15" pageNumber="16" value="2011-01-12" valueMax="2011-01-13" valueMin="2011-01-12">12-13 January 2011</date>
and
</paragraph>
<caption id="DF02667E9533B849FF0BF80DCFFDF834" ID-DOI="http://doi.org/10.5281/zenodo.12956997" ID-Zenodo-Dep="12956997" box="[137,1415,1953,1973]" httpUri="https://zenodo.org/record/12956997/files/figure.png" pageId="15" pageNumber="16" startId="15.[137,171,1953,1973]" targetBox="[159,1402,1038,1928]" targetPageId="15" targetType="figure">
<paragraph id="8BC236F69533B849FF0BF80DCFFDF834" blockId="15.[137,1415,1953,1973]" box="[137,1415,1953,1973]" pageId="15" pageNumber="16">
<emphasis id="B909EAE49533B849FF0BF80DCAA9F834" bold="true" box="[137,211,1953,1973]" pageId="15" pageNumber="16">Fig. 17.</emphasis>
<taxonomicName id="4C7D4D759533B849FF5FF80DCB88F834" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[221,498,1953,1973]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE49533B849FF5FF80DCB88F834" box="[221,498,1953,1973]" italics="true" pageId="15" pageNumber="16">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F9533B849FE7AF80DC857F834" box="[504,557,1953,1973]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
Larva paratype. Gnathosoma: A- dorsal view, B. ventral view, C- pharyngeal pump system.
</paragraph>
</caption>
<paragraph id="8BC236F6952CB856FF2CFF04CB8EFF3E" blockId="16.[174,500,168,191]" box="[174,500,168,191]" pageId="16" pageNumber="17">
<bibCitation id="5E573E9A952CB856FF2CFF04CB8EFF3E" box="[174,500,168,191]" journal="Zoological Studies" page="11" pageId="16" pageNumber="17" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE4952CB856FF2CFF04CB1DFF3E" box="[174,359,168,191]" italics="true" pageId="16" pageNumber="17">Zoological Studies</emphasis>
<emphasis id="B909EAE4952CB856FEEFFF04CBF4FF3E" bold="true" box="[365,398,168,191]" pageId="16" pageNumber="17">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
<paragraph id="8BC236F6952CB856FF2CFF51CB23FE17" blockId="16.[174,775,253,599]" pageId="16" pageNumber="17">
<date id="FFC31036952CB856FF2CFF51CBD6FE97" box="[174,428,253,278]" pageId="16" pageNumber="17" value="2011-02-07" valueMax="2011-02-11" valueMin="2011-02-07">7-11 February 2011</date>
. Collectors: Peter Hawkes, Jonathan Fisher and assistants: Faith Kgafane, Katarina Jovanovic, Maphilisi Zulu, Matimba Baloyi, Michael Pierce, Nkosinathi Babu and Philippa Wing.
</paragraph>
<paragraph id="8BC236F6952CB856FF68FE31C97DFDD7" blockId="16.[174,775,253,599]" pageId="16" pageNumber="17">
<emphasis id="B909EAE4952CB856FF68FE31CB8BFE37" box="[234,497,413,438]" italics="true" pageId="16" pageNumber="17">Deposition of types</emphasis>
: 
<materialsCitation id="3B153CAB952CB856FD85FE31C97DFDD7" collectionCode="USNM" country="Poland" location="Beltsville" municipality="University Adam Mickiewicz University in Poznan" pageId="16" pageNumber="17" specimenCount="13" specimenCount-female="10" specimenCount-larva="1" specimenCount-male="2" stateProvince="Maryland" typeStatus="holotype">
<typeStatus id="54C68854952CB856FD85FE31C806FE37" box="[519,636,413,438]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
female, 
<specimenCount id="9D7BFD7F952CB856FD7AFE31CBFAFE57" count="4" pageId="16" pageNumber="17" type="female" typeStatus="paratype">4 paratype females</specimenCount>
, 
<specimenCount id="9D7BFD7F952CB856FE09FE11C836FE57" box="[395,588,445,470]" count="1" pageId="16" pageNumber="17" type="male" typeStatus="paratype">1 paratype male</specimenCount>
and 
<specimenCount id="9D7BFD7F952CB856FD0AFE11CA88FE77" count="1" pageId="16" pageNumber="17" type="larva" typeStatus="paratype">1 paratype larva</specimenCount>
, deposited in the 
<collectionCode id="ED6CAE33952CB856FE60FE71C84EFE76" box="[482,564,477,503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="17" type="Museum">USNM</collectionCode>
mite collection, 
<location id="8EA2602D952CB856FF2CFE51CB66FD97" LSID="urn:lsid:plazi:treatment:03D487E09538B857FF43FD2BCBBEFF3E:8EA2602D952CB856FF2CFE51CB66FD97" box="[174,284,509,534]" country="Poland" municipality="University Adam Mickiewicz University in Poznan" name="Beltsville" pageId="16" pageNumber="17" stateProvince="Maryland">Beltsville</location>
, 
<collectingRegion id="49B9F814952CB856FEA5FE51CBE3FD97" box="[295,409,509,534]" country="United States of America" name="Maryland" pageId="16" pageNumber="17">Maryland</collectingRegion>
, USA; 
<specimenCount id="9D7BFD7F952CB856FE6EFE51C8ABFD97" box="[492,721,509,534]" count="6" pageId="16" pageNumber="17" type="female" typeStatus="paratype">6 paratype females</specimenCount>
and 
<specimenCount id="9D7BFD7F952CB856FF2CFDB1CB08FDB7" box="[174,370,541,566]" count="1" pageId="16" pageNumber="17" type="male" typeStatus="paratype">1 paratype male</specimenCount>
in the collection of the 
<collectingMunicipality id="6BA6AC8C952CB856FD13FDB1C8E3FDD6" pageId="16" pageNumber="17">University Adam Mickiewicz University in Poznań</collectingMunicipality>
, 
<collectingCountry id="F36A7666952CB856FD2AFD91C978FDD7" box="[680,770,573,598]" name="Poland" pageId="16" pageNumber="17">Poland</collectingCountry>
;
</materialsCitation>
</paragraph>
<paragraph id="8BC236F6952CB856FAA3FF04CFD1FF3E" blockId="16.[1313,1451,168,191]" box="[1313,1451,168,191]" pageId="16" pageNumber="17">page 17 of 21</paragraph>
<paragraph id="8BC236F6952CB856FCD0FF51CE2DFDB7" blockId="16.[850,1452,253,567]" pageId="16" pageNumber="17">
<specimenCount id="9D7BFD7F952CB856FCD0FF51CE3DFE97" box="[850,1095,253,278]" count="4" pageId="16" pageNumber="17" type="female" typeStatus="paratype">4 paratype females</specimenCount>
and 
<specimenCount id="9D7BFD7F952CB856FB09FF51CF23FE97" box="[1163,1369,253,278]" count="1" pageId="16" pageNumber="17" type="male" typeStatus="paratype">1 paratype male</specimenCount>
in the collection of the Zoologisches Institut und Museum of Universität 
<collectingRegion id="49B9F814952CB856FB8EFE91CEFCFED7" box="[1036,1158,317,342]" country="Germany" name="Hamburg" pageId="16" pageNumber="17">Hamburg</collectingRegion>
, 
<collectingCountry id="F36A7666952CB856FB14FE91CF6AFED6" box="[1174,1296,317,343]" name="Germany" pageId="16" pageNumber="17">Germany</collectingCountry>
; 
<specimenCount id="9D7BFD7F952CB856FAA2FE91C9C9FEF7" count="4" pageId="16" pageNumber="17" type="female" typeStatus="paratype">4 paratype females</specimenCount>
and 
<specimenCount id="9D7BFD7F952CB856FC7AFEF1CEBFFEF7" box="[1016,1221,349,374]" count="1" pageId="16" pageNumber="17" type="male" typeStatus="paratype">1 paratype male</specimenCount>
in the Muséum d’ Histoire naturelle, Département des Arthropodes et d’ Entomologie, 
<collectingRegion id="49B9F814952CB856FB96FE31CE0CFE36" box="[1044,1142,413,439]" country="Switzerland" name="Geneve" pageId="16" pageNumber="17">Genève</collectingRegion>
, 
<collectingCountry id="F36A7666952CB856FB00FE31CF6AFE36" box="[1154,1296,413,439]" name="Switzerland" pageId="16" pageNumber="17">Switzerland</collectingCountry>
; 
<specimenCount id="9D7BFD7F952CB856FA9FFE31C9C8FE57" count="10" pageId="16" pageNumber="17" type="female" typeStatus="paratype">10 paratype females</specimenCount>
and several non-type female specimens and 
<specimenCount id="9D7BFD7F952CB856FC08FE71CE2EFE77" box="[906,1108,477,502]" count="1" pageId="16" pageNumber="17" type="male" typeStatus="paratype">1 paratype male</specimenCount>
in the 
<taxonomicName id="4C7D4D75952CB856FB2FFE71CF52FE77" box="[1197,1320,477,502]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Arthropoda" rank="class">Arachnida</taxonomicName>
Collection of the ARC Plant Protection Research Institute, Pretoria, 
<collectingCountry id="F36A7666952CB856FC3DFDB1CE29FDB7" box="[959,1107,541,567]" name="South Africa" pageId="16" pageNumber="17">South Africa</collectingCountry>
.
</paragraph>
<caption id="DF02667E952CB856FF2CF833C9A7F832" ID-DOI="http://doi.org/10.5281/zenodo.12957009" ID-Zenodo-Dep="12957009" box="[174,989,1951,1971]" httpUri="https://zenodo.org/record/12957009/files/figure.png" pageId="16" pageNumber="17" startId="16.[174,208,1951,1971]" targetBox="[292,1330,709,1916]" targetPageId="16" targetType="figure">
<paragraph id="8BC236F6952CB856FF2CF833C9A7F832" blockId="16.[174,989,1951,1971]" box="[174,989,1951,1971]" pageId="16" pageNumber="17">
<emphasis id="B909EAE4952CB856FF2CF833CA82F832" bold="true" box="[174,248,1951,1971]" pageId="16" pageNumber="17">Fig. 18.</emphasis>
<taxonomicName id="4C7D4D75952CB856FE81F833C863F832" authorityName="Camerik &amp; Magowski &amp; Hawkes &amp; Ueckermann &amp; Ochoa &amp; Bauchan" authorityYear="2016" box="[259,537,1951,1971]" class="Arachnida" family="Pygmephoridae" genus="Zambedania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sekhukhunensis" status="sp. nov.">
<emphasis id="B909EAE4952CB856FE81F833C863F832" box="[259,537,1951,1971]" italics="true" pageId="16" pageNumber="17">Zambedania sekhukhunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A579F952CB856FD9DF833C829F832" box="[543,595,1951,1971]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
Larva paratype. Leg I (right): dorsal view.
</paragraph>
</caption>
<paragraph id="8BC236F6952DB857FFFFFF04CBBEFF3E" blockId="17.[125,452,168,191]" box="[125,452,168,191]" pageId="17" pageNumber="18">
<bibCitation id="5E573E9A952DB857FFFFFF04CBBEFF3E" box="[125,452,168,191]" journal="Zoological Studies" page="11" pageId="17" pageNumber="18" score="5.2987992369807255" volume="55" year="2016">
<emphasis id="B909EAE4952DB857FFFFFF04CB4DFF3E" box="[125,311,168,191]" italics="true" pageId="17" pageNumber="18">Zoological Studies</emphasis>
<emphasis id="B909EAE4952DB857FEBFFF04CB24FF3E" bold="true" box="[317,350,168,191]" pageId="17" pageNumber="18">55:</emphasis>
11 (2016)
</bibCitation>
</paragraph>
</subSubSection>
</treatment>
</document>