treatments-xml/data/03/B0/87/03B087BAFF94FFC0F5F517E3473EFE7D.xml

<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3887.2.1" ID-GBIF-Dataset="2c9e69f1-6daf-4139-b5aa-9363b2c83ad4" ID-ISSN="1175-5326" ID-Zenodo-Dep="228058" ID-ZooBank="D8477203-4DDE-4893-81FA-5F62CA142B03" checkinTime="1460975027897" checkinUser="plazi" docAuthor="Gardini, Giulio" docDate="2014" docId="03B087BAFF94FFC0F5F517E3473EFE7D" docLanguage="en" docName="zt03887p137.pdf" docOrigin="Zootaxa 3887 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chthonius (Chthonius) pusillus Beier 1947" docType="treatment" docVersion="9" lastPageNumber="124" masterDocId="FF89FFC2FF81FFD7F56217744641FFD0" masterDocTitle="The species of the Chthonius heterodactylus group (Arachnida, Pseudoscorpiones, Chthoniidae) from the eastern Alps and the Carpathians" masterLastPageNumber="137" masterPageNumber="101" pageNumber="122" updateTime="1676314267194" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The species of the Chthonius heterodactylus group (Arachnida, Pseudoscorpiones, Chthoniidae) from the eastern Alps and the Carpathians</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gardini, Giulio</mods:namePart>
<mods:affiliation>c / o DISTAV, Università degli Studi, corso Europa 26, I- 16132 Genova, Italy.</mods:affiliation>
<mods:nameIdentifier type="email">giuliogardini@libero.it</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-11-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3887</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>101</mods:start>
<mods:end>137</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3887.2.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">2c9e69f1-6daf-4139-b5aa-9363b2c83ad4</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">228058</mods:identifier>
<mods:identifier type="ZooBank">D8477203-4DDE-4893-81FA-5F62CA142B03</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6128244" ID-GBIF-Taxon="119594110" ID-Zenodo-Dep="6128244" LSID="urn:lsid:plazi:treatment:03B087BAFF94FFC0F5F517E3473EFE7D" httpUri="http://treatment.plazi.org/id/03B087BAFF94FFC0F5F517E3473EFE7D" lastPageId="23" lastPageNumber="124" pageId="21" pageNumber="122">
<subSubSection box="[151,578,151,178]" pageId="21" pageNumber="122" type="nomenclature">
<paragraph blockId="21.[151,578,151,211]" box="[151,578,151,178]" pageId="21" pageNumber="122">
<heading bold="true" box="[151,578,151,178]" fontSize="11" level="1" pageId="21" pageNumber="122" reason="1">
<taxonomicName ID-CoL="V3FS" authority="Beier, 1947" authorityName="Beier" authorityYear="1947" box="[151,578,151,178]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="122" phylum="Arthropoda" rank="species" species="pusillus" subGenus="Chthonius">
<emphasis bold="true" box="[151,578,151,178]" pageId="21" pageNumber="122">
<emphasis bold="true" box="[151,433,151,177]" italics="true" pageId="21" pageNumber="122">Chthonius (C.) pusillus</emphasis>
<bibRefCitation author="Beier" box="[441,578,152,178]" firstAuthor="Beier" pageId="21" pageNumber="122" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
Beier, 
<number box="[519,578,152,178]" pageId="21" pageNumber="122" value="1947.0">1947</number>
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,337,186,211]" pageId="21" pageNumber="122" type="description">
<paragraph blockId="21.[151,578,151,211]" box="[151,337,186,211]" pageId="21" pageNumber="122">
(
<figureCitation box="[159,288,186,211]" captionStart="FIGURES 50 – 58" captionStartId="22.[151,264,1628,1651]" captionTargetBox="[255,1332,407,1626]" captionTargetId="figure@22.[226,1360,378,1638]" captionTargetPageId="22" captionText="FIGURES 50 – 58. Chthonius (C.) pusillus Beier, 1947. 50 – 52, Anterior margin of carapace of ♂ (Austria) (50), ♀ (Austria) (51) and ♂ (Slovenia) (52). 53, Anterolateral portion of carapace of ♂ (Austria). 54 – 55, Right chelicera of ♂ (54) and ♀ (55) (Austria). 56 – 58, Right chela, lateral view, of ♂ (Austria) (56), ♀ (Austria) (57) and ♂ (Slovenia) (58). Figures 50 – 51, 53 – 57: adults from Austria, Weizklamm; 52, 58: male from Slovenia, Šturmova graba." httpUri="https://zenodo.org/record/228063/files/figure.png" pageId="21" pageNumber="122">
Figs 
<number box="[215,244,186,211]" pageId="21" pageNumber="122" value="50.0">50</number>
–58
</figureCitation>
, 
<figureCitation box="[299,328,187,211]" captionStart="FIGURE 90" captionStartId="33.[153,252,1653,1675]" captionTargetBox="[167,1422,866,1638]" captionTargetId="figure@33.[165,1424,863,1640]" captionTargetPageId="33" captionText="FIGURE 90. Distribution map for Chthonius (C.) cavernarum Ellingsen, 1909 (▲), C. (C.) heterodactylus Tömösváry, 1882 (Δ), C. (C.) hungaricus Mahnert, 1981 (■), C. (C.) lessiniensis Schawaller, 1982 (□), C. (C.) pusillus Beier, 1947 (●), C. (C.) raridentatus Hadži, 1930 (○)." httpUri="https://zenodo.org/record/228068/files/figure.png" pageId="21" pageNumber="122">90</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[151,628,258,280]" pageId="21" pageNumber="122" type="reference_group">
<paragraph blockId="21.[151,628,258,280]" box="[151,628,258,280]" pageId="21" pageNumber="122">
<treatmentCitationGroup box="[151,628,258,280]" pageId="21" pageNumber="122">
<taxonomicName ID-CoL="V3FS" authority="Beier 1947: 298" authorityName="Beier" authorityPageNumber="298" authorityYear="1947" box="[151,559,258,280]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="122" phylum="Arthropoda" rank="species" species="pusillus" subGenus="Chthonius">
<emphasis box="[151,387,258,280]" italics="true" pageId="21" pageNumber="122">Chthonius (C.) pusillus</emphasis>
<treatmentCitation author="Beier" box="[393,559,258,280]" page="298" pageId="21" pageNumber="122" year="1947">
<bibRefCitation author="Beier" box="[393,559,258,280]" firstAuthor="Beier" page="298" pageId="21" pageNumber="122" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
Beier 
<number box="[454,508,258,280]" pageId="21" pageNumber="122" value="1947.0">1947</number>
: 
<number box="[518,559,258,280]" pageId="21" pageNumber="122" value="298.0">298</number>
</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 
<number box="[609,624,258,280]" pageId="21" pageNumber="122" value="2.0">2</number>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection box="[151,1232,325,350]" pageId="21" pageNumber="122" type="materials_examined">
<paragraph blockId="21.[151,1437,325,2010]" box="[151,1232,325,350]" pageId="21" pageNumber="122">
<emphasis bold="true" box="[151,303,325,350]" pageId="21" pageNumber="122">
<typeStatus box="[151,210,325,350]" pageId="21" pageNumber="122">Type</typeStatus>
locality
</emphasis>
: 
<collectingCountry box="[318,403,325,350]" name="Austria" pageId="21" pageNumber="122">Austria</collectingCountry>
, Styria, Mitterdorf im Mürztal towards Veitsch (
<number box="[944,973,325,350]" pageId="21" pageNumber="122" value="47.0">47</number>
° 
<number box="[982,1011,325,350]" pageId="21" pageNumber="122" value="32.0">32</number>
′ 
<number box="[1016,1044,325,350]" pageId="21" pageNumber="122" value="43.0">43</number>
″ 
<geoCoordinate box="[1057,1160,325,350]" direction="north" orientation="latitude" pageId="21" pageNumber="122" precision="925" value="15.5">
N 
<number box="[1086,1113,325,350]" pageId="21" pageNumber="122" value="15.0">15</number>
° 
<number box="[1122,1153,325,350]" pageId="21" pageNumber="122" value="30.0">30</number>
′
</geoCoordinate>
<number box="[1160,1188,325,350]" pageId="21" pageNumber="122" value="17.0">17</number>
″E).
</paragraph>
</subSubSection>
<subSubSection box="[199,561,361,386]" pageId="21" pageNumber="122" type="distribution">
<paragraph blockId="21.[151,1437,325,2010]" box="[199,561,361,386]" pageId="21" pageNumber="122">
<emphasis bold="true" box="[199,352,361,386]" pageId="21" pageNumber="122">Distribution.</emphasis>
<collectingCountry box="[359,444,361,386]" name="Austria" pageId="21" pageNumber="122">Austria</collectingCountry>
, 
<collectingCountry box="[455,557,361,386]" name="Slovenia" pageId="21" pageNumber="122">Slovenia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="122" type="diagnosis">
<paragraph blockId="21.[151,1437,325,2010]" pageId="21" pageNumber="122">
<emphasis bold="true" box="[199,331,397,422]" pageId="21" pageNumber="122">Diagnosis (</emphasis>
♂♀
<emphasis bold="true" box="[373,389,397,422]" pageId="21" pageNumber="122">).</emphasis>
An eyed epigean 
<emphasis box="[597,854,397,422]" italics="true" pageId="21" pageNumber="122">
<taxonomicName box="[597,712,397,422]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="122" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(
<taxonomicName box="[729,846,397,422]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="122" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
)
</emphasis>
from the eastern Alps of 
<collectingCountry box="[1148,1231,397,422]" name="Austria" pageId="21" pageNumber="122">Austria</collectingCountry>
and 
<collectingCountry box="[1288,1386,397,422]" name="Slovenia" pageId="21" pageNumber="122">Slovenia</collectingCountry>
that differs from other species of the 
<taxonomicName box="[514,678,433,458]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="122" phylum="Arthropoda" rank="species" species="heterodactylus">
<emphasis box="[514,678,433,458]" italics="true" pageId="21" pageNumber="122">heterodactylus</emphasis>
</taxonomicName>
group in the following combination of characters: carapace without epistome, chaetotaxy m 
<quantity box="[420,460,469,494]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="122" unit="m" value="4.0">4m</quantity>
: 
<number box="[464,478,469,494]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[482,497,469,494]" pageId="21" pageNumber="122" value="4.0">4</number>
: 
<number box="[501,516,469,494]" pageId="21" pageNumber="122" value="2.0">2</number>
:m 
<quantity box="[544,584,469,494]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="122" unit="m" value="2.0">2m</quantity>
(
<number box="[600,632,470,494]" pageId="21" pageNumber="122" value="22.0">22</number>
); chaetotaxy tergites I–IV: 
<number box="[937,954,470,494]" pageId="21" pageNumber="122" value="4.0">4</number>
: 
<number box="[959,976,470,494]" pageId="21" pageNumber="122" value="4.0">4</number>
: 
<number box="[980,997,470,494]" pageId="21" pageNumber="122" value="4.0">4</number>
: 
<number box="[1002,1019,470,494]" pageId="21" pageNumber="122" value="4.0">4</number>
; genital opening of males flanked by aligned setae on each side; fixed chelal finger with 
<number box="[757,827,505,530]" pageId="21" pageNumber="122" value="17.0" valueMax="18.0" valueMin="16.0">16–18</number>
teeth with dental canals, usually without intercalary microdenticles, and with distal paraxial seta apically sinuous and weakly enlarged; movable chelal finger with 
<number box="[151,221,577,601]" pageId="21" pageNumber="122" value="15.0" valueMax="18.0" valueMin="12.0">12–18</number>
low, reclined teeth increasingly reduced towards finger base, reaching back almost to 
<emphasis box="[1184,1209,577,602]" italics="true" pageId="21" pageNumber="122">sb</emphasis>
; coupled sensilla 
<emphasis box="[1409,1435,578,601]" italics="true" pageId="21" pageNumber="122">pc</emphasis>
of movable chelal finger between trichobothria 
<emphasis box="[697,722,613,638]" italics="true" pageId="21" pageNumber="122">sb</emphasis>
and 
<emphasis box="[780,794,613,638]" italics="true" pageId="21" pageNumber="122">b</emphasis>
; chela length 
<number box="[959,1072,613,638]" pageId="21" pageNumber="122" value="0.74" valueMax="0.81" valueMin="0.67">0.67–0.81</number>
(♂) or 
<number box="[1162,1274,613,638]" pageId="21" pageNumber="122" value="0.77" valueMax="0.79" valueMin="0.75">0.75–0.79</number>
(♀); movable finger length 
<number box="[302,414,649,674]" pageId="21" pageNumber="122" value="0.45999999999999996" valueMax="0.5" valueMin="0.42">0.42–0.50</number>
(♂) or 
<number box="[498,610,649,674]" pageId="21" pageNumber="122" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48</number>
(♀).
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="122" type="materials_examined">
<paragraph blockId="21.[151,1437,325,2010]" pageId="21" pageNumber="122">
<emphasis bold="true" box="[199,433,685,710]" pageId="21" pageNumber="122">Material examined.</emphasis>
AUSTRIA— 
<emphasis box="[589,787,685,710]" italics="true" pageId="21" pageNumber="122">Niederösterreich:</emphasis>
<specimenCount box="[795,837,686,710]" pageId="21" pageNumber="122" type="generic">
<number box="[795,809,686,710]" pageId="21" pageNumber="122" value="2.0">2</number>
♂
</specimenCount>
(
<taxonomicName authority="M. Beier" authorityName="M. Beier" box="[854,1165,685,710]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="122" phylum="Arthropoda" rank="species" species="heterodactylus">
<emphasis box="[854,1051,685,710]" italics="true" pageId="21" pageNumber="122">C. heterodactylus</emphasis>
, M. Beier
</taxonomicName>
det.), Harzberg bei Bad Vöslau, Lindenwald, 
<date box="[401,519,721,746]" pageId="21" pageNumber="122" value="1960-10-22">
<number box="[401,431,721,746]" pageId="21" pageNumber="122" value="22.0">22</number>
.X. 
<number box="[460,519,721,746]" pageId="21" pageNumber="122" value="1960.0">1960</number>
</date>
(collector not indicated) (
<collectionCode box="[828,926,722,746]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="122">NHMW</collectionCode>
); 
<specimenCount box="[951,999,722,746]" pageId="21" pageNumber="122" type="generic">
<number box="[951,965,722,746]" pageId="21" pageNumber="122" value="1.0">1</number>
♂
</specimenCount>
, 
<number box="[1016,1030,722,746]" pageId="21" pageNumber="122" value="3.0">3</number>
♀ (V. Mahnert det. 
<number box="[1278,1337,722,746]" pageId="21" pageNumber="122" value="1978.0">1978</number>
), Bezirk Scheibbs, Gaming, Kienberg-S-Hang, 
<date box="[594,724,758,782]" pageId="21" pageNumber="122" value="1978-11-13">
<number box="[594,622,758,782]" pageId="21" pageNumber="122" value="13.0">13</number>
.XI. 
<number box="[665,724,758,782]" pageId="21" pageNumber="122" value="1978.0">1978</number>
</date>
, F. Ressl leg., leaf litter (
<collectionCode box="[1025,1117,758,782]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="21" pageNumber="122">MHNG</collectionCode>
). 
<emphasis box="[1139,1270,757,782]" italics="true" pageId="21" pageNumber="122">Steiermark:</emphasis>
<specimenCount box="[1281,1329,758,782]" pageId="21" pageNumber="122" type="generic">
<number box="[1281,1295,758,782]" pageId="21" pageNumber="122" value="2.0">2</number>
♂
</specimenCount>
, 
<number box="[1344,1358,758,782]" pageId="21" pageNumber="122" value="1.0">1</number>
♀ (
<taxonomicName authority="M. Beier" authorityName="M. Beier" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="122" phylum="Arthropoda" rank="species" species="heterodactylus">
<emphasis italics="true" pageId="21" pageNumber="122">C. heterodactylus</emphasis>
, M. Beier
</taxonomicName>
det.), Weberkogel zw. Birkfeld und Fischbach, 
<date box="[1035,1136,793,818]" pageId="21" pageNumber="122" value="1967-05-06">
<number box="[1035,1049,793,818]" pageId="21" pageNumber="122" value="6.0">6</number>
.V. 
<number box="[1078,1136,793,818]" pageId="21" pageNumber="122" value="1967.0">1967</number>
</date>
(collector not indicated) (
<collectionCode box="[158,256,830,854]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="122">NHMW</collectionCode>
); 
<specimenCount box="[278,339,830,854]" pageId="21" pageNumber="122" type="generic">
<number box="[278,306,830,854]" pageId="21" pageNumber="122" value="10.0">10</number>
♂
</specimenCount>
, 
<number box="[353,367,830,854]" pageId="21" pageNumber="122" value="2.0">2</number>
♀ (V. Mahnert det. 
<number box="[604,663,830,854]" pageId="21" pageNumber="122" value="1993.0">1993</number>
), Bezirk Weiz, Raabklamm, 
<date box="[994,1131,830,854]" pageId="21" pageNumber="122" value="1979-12-11">
<number box="[994,1020,830,854]" pageId="21" pageNumber="122" value="11.0">11</number>
.XII. 
<number box="[1071,1131,830,854]" pageId="21" pageNumber="122" value="1979.0">1979</number>
</date>
– 
<date box="[1148,1273,830,854]" pageId="21" pageNumber="122" value="1980-03-31">
<number box="[1148,1176,830,854]" pageId="21" pageNumber="122" value="31.0">31</number>
.III. 
<number box="[1213,1273,830,854]" pageId="21" pageNumber="122" value="1980.0">1980</number>
</date>
, P. Horak leg. (
<collectionCode box="[159,251,866,890]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="21" pageNumber="122">MHNG</collectionCode>
); 
<specimenCount box="[270,315,866,890]" pageId="21" pageNumber="122" type="generic">
<number box="[270,284,866,890]" pageId="21" pageNumber="122" value="4.0">4</number>
♂
</specimenCount>
, 
<number box="[328,342,866,890]" pageId="21" pageNumber="122" value="3.0">3</number>
♀ (V. Mahnert det. 
<number box="[570,629,866,890]" pageId="21" pageNumber="122" value="1993.0">1993</number>
), Bezirk Weiz, Weizklamm, 
<number box="[953,994,865,890]" pageId="21" pageNumber="122" value="635.0">635</number>
/ 
<quantity box="[1003,1074,865,890]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="21" pageNumber="122" unit="m" value="660.0">660 m</quantity>
a.s.l., 
<date box="[1148,1263,865,890]" pageId="21" pageNumber="122" value="1980-11-08">
<number box="[1148,1161,865,890]" pageId="21" pageNumber="122" value="8.0">8</number>
.XI. 
<number box="[1202,1263,865,890]" pageId="21" pageNumber="122" value="1980.0">1980</number>
</date>
– 
<date box="[1279,1404,865,890]" pageId="21" pageNumber="122" value="1981-03-25">
<number box="[1279,1309,865,890]" pageId="21" pageNumber="122" value="25.0">25</number>
.III. 
<number box="[1346,1404,865,890]" pageId="21" pageNumber="122" value="1981.0">1981</number>
</date>
, P. Horak leg., 
<taxonomicName box="[283,498,901,926]" class="Magnoliopsida" family="Betulaceae" genus="Ostrya" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="122" phylum="Tracheophyta" rank="species" species="carpinifolia">
<emphasis box="[283,498,901,926]" italics="true" pageId="21" pageNumber="122">Ostrya carpinifolia</emphasis>
</taxonomicName>
wood (
<collectionCode box="[583,675,902,926]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="21" pageNumber="122">MHNG</collectionCode>
).
</paragraph>
<paragraph blockId="21.[151,1437,325,2010]" pageId="21" pageNumber="122">
<collectingCountry box="[199,339,938,962]" name="Slovenia" pageId="21" pageNumber="122">SLOVENIA</collectingCountry>
<emphasis bold="true" box="[339,367,937,962]" pageId="21" pageNumber="122">—</emphasis>
<emphasis box="[367,570,937,962]" italics="true" pageId="21" pageNumber="122">Podravska regija:</emphasis>
<specimenCount box="[579,624,938,962]" pageId="21" pageNumber="122" type="generic">
<number box="[579,593,938,962]" pageId="21" pageNumber="122" value="1.0">1</number>
♂
</specimenCount>
, Maribor, Selnica ob Dravi, Šturmova graba Valley, artificial gallery in Šturmova graba, no. U3, 
<date box="[433,553,974,998]" pageId="21" pageNumber="122" value="1977-10-21">
21.X. 
<number box="[492,553,974,998]" pageId="21" pageNumber="122" value="1977.0">1977</number>
</date>
, V. Kuštor &amp; T. Novak leg., trap (
<collectionCode box="[932,1012,974,998]" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="21" pageNumber="122">PMSL</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="123" pageId="21" pageNumber="122" type="description">
<paragraph blockId="21.[151,1437,325,2010]" lastBlockId="22.[151,1436,151,359]" lastPageId="22" lastPageNumber="123" pageId="21" pageNumber="122">
<emphasis bold="true" box="[199,471,1009,1034]" pageId="21" pageNumber="122">Description of adults (</emphasis>
♂♀
<emphasis bold="true" box="[513,522,1009,1034]" pageId="21" pageNumber="122">)</emphasis>
. Carapace, tergites, chelicerae and pedipalps pale brown; hispid granulation on lateral surfaces of carapace, on cheliceral palm and on base of cheliceral movable finger. Carapace 
<number box="[1257,1369,1045,1070]" pageId="21" pageNumber="122" value="1.0" valueMax="1.05" valueMin="0.95">0.95–1.05</number>
times longer than broad, subquadrate, slightly constricted posteriorly; anterior margin (
<figureCitation box="[1104,1237,1081,1106]" captionStart="FIGURES 50 – 58" captionStartId="22.[151,264,1628,1651]" captionTargetBox="[255,1332,407,1626]" captionTargetId="figure@22.[226,1360,378,1638]" captionTargetPageId="22" captionText="FIGURES 50 – 58. Chthonius (C.) pusillus Beier, 1947. 50 – 52, Anterior margin of carapace of ♂ (Austria) (50), ♀ (Austria) (51) and ♂ (Slovenia) (52). 53, Anterolateral portion of carapace of ♂ (Austria). 54 – 55, Right chelicera of ♂ (54) and ♀ (55) (Austria). 56 – 58, Right chela, lateral view, of ♂ (Austria) (56), ♀ (Austria) (57) and ♂ (Slovenia) (58). Figures 50 – 51, 53 – 57: adults from Austria, Weizklamm; 52, 58: male from Slovenia, Šturmova graba." httpUri="https://zenodo.org/record/228063/files/figure.png" pageId="21" pageNumber="122">
Figs 
<number box="[1166,1237,1081,1106]" pageId="21" pageNumber="122" value="51.0" valueMax="52.0" valueMin="50.0">50–52</number>
</figureCitation>
) usually without epistome, with thin, sharp denticles between median macrosetae; ocular area as in 
<figureCitation box="[1125,1211,1117,1142]" captionStart="FIGURES 50 – 58" captionStartId="22.[151,264,1628,1651]" captionTargetBox="[255,1332,407,1626]" captionTargetId="figure@22.[226,1360,378,1638]" captionTargetPageId="22" captionText="FIGURES 50 – 58. Chthonius (C.) pusillus Beier, 1947. 50 – 52, Anterior margin of carapace of ♂ (Austria) (50), ♀ (Austria) (51) and ♂ (Slovenia) (52). 53, Anterolateral portion of carapace of ♂ (Austria). 54 – 55, Right chelicera of ♂ (54) and ♀ (55) (Austria). 56 – 58, Right chela, lateral view, of ♂ (Austria) (56), ♀ (Austria) (57) and ♂ (Slovenia) (58). Figures 50 – 51, 53 – 57: adults from Austria, Weizklamm; 52, 58: male from Slovenia, Šturmova graba." httpUri="https://zenodo.org/record/228063/files/figure.png" pageId="21" pageNumber="122">
Fig. 
<number box="[1181,1211,1117,1142]" pageId="21" pageNumber="122" value="53.0">53</number>
</figureCitation>
, anterior eyes with convex lens (diameter 
<quantity box="[403,598,1153,1178]" metricMagnitude="-5" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="21" pageNumber="122" unit="mm" value="0.0525" valueMax="0.055" valueMin="0.05">0.050–0.055 mm</quantity>
), posterior eyes with flat lens, both pairs with tapeta; distance from anterior eyes to anterior margin of carapace 
<quantity box="[554,749,1190,1214]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="21" pageNumber="122" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.030–0.040 mm</quantity>
; chaetotaxy m 
<quantity box="[914,954,1189,1214]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="122" unit="m" value="4.0">4m</quantity>
: 
<number box="[958,972,1189,1214]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[976,991,1189,1214]" pageId="21" pageNumber="122" value="4.0">4</number>
: 
<number box="[994,1009,1189,1214]" pageId="21" pageNumber="122" value="2.0">2</number>
:m 
<quantity box="[1038,1077,1189,1214]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="122" unit="m" value="2.0">2m</quantity>
(
<number box="[1093,1125,1190,1214]" pageId="21" pageNumber="122" value="22.0">22</number>
), macrosetae thick, rarely 
<number box="[1422,1436,1190,1214]" pageId="21" pageNumber="122" value="2.0">2</number>
preocular or 
<number box="[293,307,1226,1250]" pageId="21" pageNumber="122" value="2.0">2</number>
posterior microsetae on one side; posterior lateral microsetae very short, rarely half length of median macrosetae (
<specimenCount box="[294,334,1262,1286]" pageId="21" pageNumber="122" type="generic">
<number box="[294,306,1262,1286]" pageId="21" pageNumber="122" value="1.0">1</number>
♂
</specimenCount>
from 
<collectingCountry box="[402,488,1261,1286]" name="Austria" pageId="21" pageNumber="122">Austria</collectingCountry>
, Raabklamm, 
<number box="[646,660,1262,1286]" pageId="21" pageNumber="122" value="1.0">1</number>
♀ from Weizklamm; 
<specimenCount box="[905,946,1262,1286]" pageId="21" pageNumber="122" type="generic">
<number box="[905,919,1262,1286]" pageId="21" pageNumber="122" value="1.0">1</number>
♂
</specimenCount>
from 
<collectingCountry box="[1013,1114,1261,1286]" name="Slovenia" pageId="21" pageNumber="122">Slovenia</collectingCountry>
, Selnica ob Dravi); length of anteromedian and anterolateral macrosetae respectively 
<number box="[811,950,1297,1322]" pageId="21" pageNumber="122" value="0.10500000000000001" valueMax="0.115" valueMin="0.095">0.095–0.115</number>
and 
<quantity box="[1017,1216,1297,1322]" metricMagnitude="-5" metricUnit="m" metricValue="8.25" metricValueMax="9.0" metricValueMin="7.5" pageId="21" pageNumber="122" unit="mm" value="0.08249999999999999" valueMax="0.09" valueMin="0.075">0.075–0.090 mm</quantity>
, length of median macrosetae of ocular row 
<quantity box="[439,632,1333,1358]" metricMagnitude="-5" metricUnit="m" metricValue="6.75" metricValueMax="8.0" metricValueMin="5.5" pageId="21" pageNumber="122" unit="mm" value="0.0675" valueMax="0.08" valueMin="0.055">0.055–0.080 mm</quantity>
, length of posterior macrosetae 
<quantity box="[987,1181,1334,1358]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7" pageId="21" pageNumber="122" unit="mm" value="0.075" valueMax="0.08" valueMin="0.07">0.070–0.080 mm</quantity>
. Chaetotaxy of tergites 
<date box="[151,226,1369,1394]" pageId="21" pageNumber="122">
I–X 
<number box="[208,226,1369,1394]" pageId="21" pageNumber="122" value="4.0">4</number>
</date>
: 
<number box="[231,249,1369,1394]" pageId="21" pageNumber="122" value="4.0">4</number>
: 
<number box="[253,271,1369,1394]" pageId="21" pageNumber="122" value="4.0">4</number>
: 
<number box="[275,293,1369,1394]" pageId="21" pageNumber="122" value="4.0">4</number>
: 
<number box="[297,314,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[318,335,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[340,357,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[361,378,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[382,399,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[403,421,1369,1394]" pageId="21" pageNumber="122" value="4.0">4</number>
. Chaetotaxy of sternites 
<date box="[726,825,1369,1394]" pageId="21" pageNumber="122" value="2010-02" valueMax="2010-10-31" valueMin="2010-02-10">
II–X 
<number box="[793,825,1369,1394]" pageId="21" pageNumber="122" value="10.0">10</number>
</date>
:(
<number box="[838,852,1369,1394]" pageId="21" pageNumber="122" value="3.0">3</number>
) 
<number box="[862,926,1369,1394]" pageId="21" pageNumber="122" value="9.0" valueMax="10.0" valueMin="8.0">8–10</number>
(
<number box="[935,949,1369,1394]" pageId="21" pageNumber="122" value="3.0">3</number>
):(
<number box="[972,988,1369,1394]" pageId="21" pageNumber="122" value="2.0">2</number>
) 
<number box="[997,1013,1369,1394]" pageId="21" pageNumber="122" value="7.0">7</number>
(
<number box="[1022,1038,1369,1394]" pageId="21" pageNumber="122" value="2.0">2</number>
): 
<number box="[1052,1100,1369,1394]" pageId="21" pageNumber="122" value="8.5" valueMax="9.0" valueMin="8.0">8–9</number>
: 
<number box="[1104,1120,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[1124,1140,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[1144,1160,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[1163,1179,1369,1394]" pageId="21" pageNumber="122" value="6.0">6</number>
: 
<number box="[1183,1199,1369,1394]" pageId="21" pageNumber="122" value="7.0">7</number>
; genital opening of males flanked by 
<number box="[356,398,1405,1430]" pageId="21" pageNumber="122" value="6.0" valueMax="7.0" valueMin="5.0">5–7</number>
(usually 
<number box="[507,522,1405,1430]" pageId="21" pageNumber="122" value="6.0">6</number>
) setae on each side [
<bibRefCitation author="Beier" box="[765,1309,1405,1430]" pageId="21" pageNumber="122" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
<number box="[765,780,1406,1430]" pageId="21" pageNumber="122" value="8.0">8</number>
setae on each side according to Beier (
<number box="[1237,1300,1406,1430]" pageId="21" pageNumber="122" value="1947.0">1947</number>
)
</bibRefCitation>
]. Chelicera (
<figureCitation box="[159,289,1441,1466]" captionStart="FIGURES 50 – 58" captionStartId="22.[151,264,1628,1651]" captionTargetBox="[255,1332,407,1626]" captionTargetId="figure@22.[226,1360,378,1638]" captionTargetPageId="22" captionText="FIGURES 50 – 58. Chthonius (C.) pusillus Beier, 1947. 50 – 52, Anterior margin of carapace of ♂ (Austria) (50), ♀ (Austria) (51) and ♂ (Slovenia) (52). 53, Anterolateral portion of carapace of ♂ (Austria). 54 – 55, Right chelicera of ♂ (54) and ♀ (55) (Austria). 56 – 58, Right chela, lateral view, of ♂ (Austria) (56), ♀ (Austria) (57) and ♂ (Slovenia) (58). Figures 50 – 51, 53 – 57: adults from Austria, Weizklamm; 52, 58: male from Slovenia, Šturmova graba." httpUri="https://zenodo.org/record/228063/files/figure.png" pageId="21" pageNumber="122">
Figs 
<number box="[217,289,1441,1466]" pageId="21" pageNumber="122" value="54.5" valueMax="55.0" valueMin="54.0">54–55</number>
</figureCitation>
) 
<number box="[306,390,1442,1466]" pageId="21" pageNumber="122" value="2.1500000000000004" valueMax="2.2" valueMin="2.1">2.1–2.2</number>
(♂) or 
<number box="[480,577,1441,1466]" pageId="21" pageNumber="122" value="2.075" valueMax="2.1" valueMin="2.05">2.05–2.1</number>
(♀) times as long as broad, palm with 
<number box="[1032,1046,1441,1466]" pageId="21" pageNumber="122" value="6.0">6</number>
setae and one microseta laterally; fixed finger with 
<number box="[351,407,1478,1502]" pageId="21" pageNumber="122" value="11.0" valueMax="13.0" valueMin="9.0">9–13</number>
teeth, the two distal teeth larger; movable finger with a large isolated subapical tooth (
<emphasis box="[1405,1427,1477,1502]" italics="true" pageId="21" pageNumber="122">di</emphasis>
) and 
<number box="[200,241,1513,1538]" pageId="21" pageNumber="122" value="7.0" valueMax="9.0" valueMin="5.0">5–9</number>
(usually 
<number box="[349,364,1514,1538]" pageId="21" pageNumber="122" value="7.0">7</number>
) teeth decreasing in size proximally; 
<emphasis box="[792,814,1513,1538]" italics="true" pageId="21" pageNumber="122">gl</emphasis>
ratio 
<number box="[882,998,1513,1538]" pageId="21" pageNumber="122" value="0.56" valueMax="0.6" valueMin="0.52">0.52–0.60</number>
; spinneret weakly prominent in males, prominent and apically rounded in females; rallum with 
<number box="[790,817,1550,1574]" pageId="21" pageNumber="122" value="11.0">11</number>
blades; serrulae interior and exterior respectively with 
<number box="[151,179,1586,1610]" pageId="21" pageNumber="122" value="12.0">12</number>
and 
<number box="[232,302,1585,1610]" pageId="21" pageNumber="122" value="15.0" valueMax="16.0" valueMin="14.0">14–16</number>
blades. Coxal setae: pedipalp 
<number box="[641,655,1585,1610]" pageId="21" pageNumber="122" value="5.0">5</number>
(including 
<number box="[783,797,1586,1610]" pageId="21" pageNumber="122" value="2.0">2</number>
on manducatory process), 
<date box="[1098,1128,1586,1610]" pageId="21" pageNumber="122">
I 
<number box="[1114,1128,1586,1610]" pageId="21" pageNumber="122" value="4.0">4</number>
</date>
+ 
<number box="[1156,1170,1586,1610]" pageId="21" pageNumber="122" value="3.0">3</number>
marginal microsetae, 
<date pageId="21" pageNumber="122">
II 
<number box="[151,167,1622,1646]" pageId="21" pageNumber="122" value="4.0">4</number>
</date>
, 
<date box="[179,228,1621,1646]" pageId="21" pageNumber="122">
III 
<number box="[214,228,1621,1646]" pageId="21" pageNumber="122" value="5.0">5</number>
</date>
(rarely 
<number box="[317,333,1622,1646]" pageId="21" pageNumber="122" value="4.0">4</number>
), 
<date box="[355,408,1621,1646]" pageId="21" pageNumber="122">
IV 
<number box="[392,408,1621,1646]" pageId="21" pageNumber="122" value="6.0">6</number>
</date>
; coxa II with 
<number box="[564,619,1621,1646]" pageId="21" pageNumber="122" value="8.5" valueMax="11.0" valueMin="6.0">6–11</number>
(usually 
<number box="[723,794,1622,1646]" pageId="21" pageNumber="122" value="10.5" valueMax="11.0" valueMin="10.0">10–11</number>
) coxal spines, coxa III with 
<number box="[1113,1155,1621,1646]" pageId="21" pageNumber="122" value="5.0" valueMax="6.0" valueMin="4.0">4–6</number>
(usually 
<number box="[1260,1275,1622,1646]" pageId="21" pageNumber="122" value="4.0">4</number>
) coxal spines; intercoxal tubercle bisetose. Pedipalp: femur 
<number box="[658,741,1657,1682]" pageId="21" pageNumber="122" value="5.0" valueMax="5.4" valueMin="4.6">4.6–5.4</number>
(♂) or 
<number box="[825,909,1657,1682]" pageId="21" pageNumber="122" value="4.949999999999999" valueMax="5.1" valueMin="4.8">4.8–5.1</number>
(♀) times as long as broad; chela (
<figureCitation box="[1300,1428,1657,1682]" captionStart="FIGURES 50 – 58" captionStartId="22.[151,264,1628,1651]" captionTargetBox="[255,1332,407,1626]" captionTargetId="figure@22.[226,1360,378,1638]" captionTargetPageId="22" captionText="FIGURES 50 – 58. Chthonius (C.) pusillus Beier, 1947. 50 – 52, Anterior margin of carapace of ♂ (Austria) (50), ♀ (Austria) (51) and ♂ (Slovenia) (52). 53, Anterolateral portion of carapace of ♂ (Austria). 54 – 55, Right chelicera of ♂ (54) and ♀ (55) (Austria). 56 – 58, Right chela, lateral view, of ♂ (Austria) (56), ♀ (Austria) (57) and ♂ (Slovenia) (58). Figures 50 – 51, 53 – 57: adults from Austria, Weizklamm; 52, 58: male from Slovenia, Šturmova graba." httpUri="https://zenodo.org/record/228063/files/figure.png" pageId="21" pageNumber="122">
Figs 
<number box="[1357,1428,1657,1682]" pageId="21" pageNumber="122" value="57.0" valueMax="58.0" valueMin="56.0">56–58</number>
</figureCitation>
) 
<number box="[151,235,1693,1718]" pageId="21" pageNumber="122" value="5.0" valueMax="5.2" valueMin="4.8">4.8–5.2</number>
(♂) or 
<number box="[325,409,1693,1718]" pageId="21" pageNumber="122" value="4.65" valueMax="4.8" valueMin="4.5">4.5–4.8</number>
(♀) times as long as deep [
<number box="[728,816,1694,1718]" pageId="21" pageNumber="122" value="4.35" valueMax="4.4" valueMin="4.3">4.3–4.4</number>
as long as deep (♂♀) in 
<bibRefCitation author="Beier" box="[1110,1260,1693,1718]" firstAuthor="Beier" pageId="21" pageNumber="122" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
Beier (
<number box="[1189,1251,1694,1718]" pageId="21" pageNumber="122" value="1947.0">1947</number>
)
</bibRefCitation>
]; hand of chela 
<number box="[151,235,1729,1754]" pageId="21" pageNumber="122" value="1.7000000000000002" valueMax="1.8" valueMin="1.6">1.6–1.8</number>
(♂) or 
<number box="[329,441,1729,1754]" pageId="21" pageNumber="122" value="1.6" valueMax="1.65" valueMin="1.55">1.55–1.65</number>
(♀) times as long as deep [
<number box="[770,807,1729,1754]" pageId="21" pageNumber="122" value="1.5">1.5</number>
times as long as deep (♂♀) in 
<bibRefCitation author="Beier" box="[1183,1333,1729,1754]" firstAuthor="Beier" pageId="21" pageNumber="122" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
Beier (
<number box="[1263,1325,1730,1754]" pageId="21" pageNumber="122" value="1947.0">1947</number>
)
</bibRefCitation>
], weakly depressed behind 
<emphasis box="[352,415,1765,1790]" italics="true" pageId="21" pageNumber="122">ib/isb</emphasis>
; fixed chelal finger sigmoid in lateral view, with 
<number box="[972,1042,1765,1790]" pageId="21" pageNumber="122" value="17.0" valueMax="18.0" valueMin="16.0">16–18</number>
teeth [
<bibRefCitation author="Beier" box="[1117,1371,1765,1790]" pageId="21" pageNumber="122" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
<quantity box="[1117,1220,1765,1790]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" pageId="21" pageNumber="122" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">20–22 in</quantity>
Beier (
<number box="[1303,1363,1766,1790]" pageId="21" pageNumber="122" value="1947.0">1947</number>
)
</bibRefCitation>
] with dental canals, usually without intercalary microdenticles (spaces between successive distal teeth undulated); reduced distal tooth at level of trichobothria 
<emphasis box="[667,721,1837,1862]" italics="true" pageId="21" pageNumber="122">dx/et</emphasis>
; 
<number box="[738,808,1838,1862]" pageId="21" pageNumber="122" value="12.0" valueMax="14.0" valueMin="10.0">10–14</number>
large, sharp, reclined teeth increasing in size as far as halfway along the finger, then reduced in size towards the base; 
<number box="[869,912,1874,1898]" pageId="21" pageNumber="122" value="5.5" valueMax="7.0" valueMin="4.0">4–7</number>
proximal teeth low [
<quantity box="[1144,1231,1873,1898]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="21" pageNumber="122" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8–10 in</quantity>
<bibRefCitation author="Beier" box="[1238,1382,1873,1898]" firstAuthor="Beier" pageId="21" pageNumber="122" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
Beier (
<number box="[1314,1374,1874,1898]" pageId="21" pageNumber="122" value="1947.0">1947</number>
)
</bibRefCitation>
, 
<quantity metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.286" metricValueMin="2.032" pageId="21" pageNumber="122" unit="in" value="8.5" valueMax="9.0" valueMin="8.0">8–9 in</quantity>
<bibRefCitation author="Beier" box="[180,326,1909,1934]" firstAuthor="Beier" pageId="21" pageNumber="122" refString="Beier, M. (1963) Ordnung Pseudoscorpionidea (Afterskorpione). In: d'Aguilar, J., Beier, M., Franz, H. &amp; Raw, F. (Eds.), Bestimmungsbucher zur Bodenfauna Europas. Vol. 1. Akademie-Verlag, Berlin, VI + 313 pp." type="book" year="1963">
Beier (
<number box="[256,318,1909,1934]" pageId="21" pageNumber="122" value="1963.0">1963</number>
)
</bibRefCitation>
], with rounded tips; base of fixed finger with 
<number box="[838,880,1909,1934]" pageId="21" pageNumber="122" value="4.5" valueMax="5.0" valueMin="4.0">4–5</number>
microtubercles; fixed finger at level of 
<emphasis box="[1325,1379,1910,1933]" italics="true" pageId="21" pageNumber="122">it/est</emphasis>
with 
<number box="[151,192,1946,1970]" pageId="21" pageNumber="122" value="3.5" valueMax="4.0" valueMin="3.0">3–4</number>
teeth occupying 
<quantity box="[386,472,1946,1970]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="122" unit="mm" value="0.1">0.1 mm</quantity>
(distance between successive apices 
<quantity box="[890,1086,1945,1970]" metricMagnitude="-5" metricUnit="m" metricValue="3.1" metricValueMax="3.5999999999999996" metricValueMin="2.6" pageId="21" pageNumber="122" unit="mm" value="0.031" valueMax="0.036" valueMin="0.026">0.026–0.036 mm</quantity>
); tip of fixed finger with apical sensilla 
<emphasis box="[242,264,1982,2006]" italics="true" pageId="21" pageNumber="122">af</emphasis>
<emphasis box="[264,272,1996,2010]" italics="true" pageId="21" pageNumber="122">
<number box="[264,272,1996,2010]" pageId="21" pageNumber="122" value="1.0">1</number>
</emphasis>
‾ 
<emphasis box="[287,295,1996,2010]" italics="true" pageId="21" pageNumber="122">
<number box="[287,295,1996,2010]" pageId="21" pageNumber="122" value="2.0">2</number>
</emphasis>
, distal paraxial seta apically sinuous and weakly thickened; movable finger strongly curved in lateral view, with 
<number box="[286,355,152,176]" pageId="22" pageNumber="123" value="15.0" valueMax="18.0" valueMin="12.0">12–18</number>
low, reclined teeth, increasingly reduced towards finger base, reaching back almost to 
<emphasis box="[1403,1428,151,176]" italics="true" pageId="22" pageNumber="123">sb</emphasis>
; proximal teeth between 
<emphasis box="[424,449,188,213]" italics="true" pageId="22" pageNumber="123">sb</emphasis>
and 
<emphasis box="[505,519,188,213]" italics="true" pageId="22" pageNumber="123">b</emphasis>
obsolete, revealed only by presence of dental canals; coupled sensilla 
<emphasis box="[1320,1346,189,212]" italics="true" pageId="22" pageNumber="123">pc</emphasis>
usually nearer to 
<emphasis box="[257,282,223,248]" italics="true" pageId="22" pageNumber="123">sb</emphasis>
than to 
<emphasis box="[374,388,223,248]" italics="true" pageId="22" pageNumber="123">b</emphasis>
, rarely halfway between them; tip of movable finger with apical sensilla 
<emphasis box="[1211,1245,225,248]" italics="true" pageId="22" pageNumber="123">am</emphasis>
<emphasis box="[1245,1253,238,252]" italics="true" pageId="22" pageNumber="123">
<number box="[1245,1253,238,252]" pageId="22" pageNumber="123" value="1.0">1</number>
</emphasis>
‾ 
<emphasis box="[1268,1276,238,252]" italics="true" pageId="22" pageNumber="123">
<number box="[1268,1276,238,252]" pageId="22" pageNumber="123" value="2.0">2</number>
</emphasis>
; trichobothria as in 
<figureCitation box="[213,342,262,287]" captionStart="FIGURES 50 – 58" captionStartId="22.[151,264,1628,1651]" captionTargetBox="[255,1332,407,1626]" captionTargetId="figure@22.[226,1360,378,1638]" captionTargetPageId="22" captionText="FIGURES 50 – 58. Chthonius (C.) pusillus Beier, 1947. 50 – 52, Anterior margin of carapace of ♂ (Austria) (50), ♀ (Austria) (51) and ♂ (Slovenia) (52). 53, Anterolateral portion of carapace of ♂ (Austria). 54 – 55, Right chelicera of ♂ (54) and ♀ (55) (Austria). 56 – 58, Right chela, lateral view, of ♂ (Austria) (56), ♀ (Austria) (57) and ♂ (Slovenia) (58). Figures 50 – 51, 53 – 57: adults from Austria, Weizklamm; 52, 58: male from Slovenia, Šturmova graba." httpUri="https://zenodo.org/record/228063/files/figure.png" pageId="22" pageNumber="123">
Figs 
<number box="[269,342,262,287]" pageId="22" pageNumber="123" value="57.0" valueMax="58.0" valueMin="56.0">56–58</number>
</figureCitation>
, 
<emphasis box="[355,382,264,287]" italics="true" pageId="22" pageNumber="123">ist</emphasis>
distad of the line 
<emphasis box="[589,661,262,287]" italics="true" pageId="22" pageNumber="123">eb/esb</emphasis>
; ratio of movable finger/hand of chela 
<number box="[1110,1208,262,287]" pageId="22" pageNumber="123" value="1.825" valueMax="1.9" valueMin="1.75">1.75–1.9</number>
(♂) or 
<number box="[1296,1380,263,287]" pageId="22" pageNumber="123" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8</number>
(♀); ratio of pedipalpal femur/movable finger 
<number box="[649,733,299,323]" pageId="22" pageNumber="123" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2</number>
(♂♀); ratio of pedipalpal femur/carapace 
<number box="[1247,1345,298,323]" pageId="22" pageNumber="123" value="1.275" valueMax="1.4" valueMin="1.15">1.15–1.4</number>
(♂) or 
<number box="[151,248,334,359]" pageId="22" pageNumber="123" value="1.225" valueMax="1.25" valueMin="1.2">1.2–1.25</number>
(♀).
</paragraph>
<caption httpUri="https://zenodo.org/record/228063/files/figure.png" pageId="22" pageNumber="123" targetBox="[255,1332,407,1626]" targetPageId="22">
<paragraph blockId="22.[151,1436,1628,1742]" pageId="22" pageNumber="123">
<emphasis bold="true" box="[151,342,1628,1651]" pageId="22" pageNumber="123">FIGURES 50–58.</emphasis>
<taxonomicName authority="Beier, 1947" authorityName="Beier" authorityYear="1947" box="[349,718,1628,1650]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="22" pageNumber="123" phylum="Arthropoda" rank="species" species="pusillus" subGenus="Chthonius">
<emphasis box="[349,588,1628,1650]" italics="true" pageId="22" pageNumber="123">
Chthonius 
<taxonomicName authority="Beier, 1947" authorityName="Beier" authorityYear="1947" box="[461,588,1628,1650]" pageId="22" pageNumber="123" rank="species" species="pusillus" subGenus="C.">
(C.) 
<taxonomicName authority="Beier" authorityName="Beier" authorityYear="1947" box="[509,588,1628,1650]" pageId="22" pageNumber="123" rank="species" species="pusillus">pusillus</taxonomicName>
</taxonomicName>
</emphasis>
<bibRefCitation author="Beier, M." box="[596,718,1628,1650]" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="22" pageNumber="123" pagination="296 - 301" part="55" refId="ref36418" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." title="Neue Pseudoscorpione aus der Steiermark" type="journal article" year="1947">Beier, 1947</bibRefCitation>
</taxonomicName>
. 50–52, Anterior margin of carapace of ♂ (Austria) (50), ♀ (Austria) (51) and ♂ (Slovenia) (52). 53, Anterolateral portion of carapace of ♂ (Austria). 54–55, Right chelicera of ♂ (54) and ♀ (55) (Austria). 56–58, Right chela, lateral view, of ♂ (Austria) (56), ♀ (Austria) (57) and ♂ (Slovenia) (58). Figures 50–51, 53–57: adults from Austria, Weizklamm; 52, 58: male from Slovenia, Šturmova graba.
</paragraph>
</caption>
<paragraph blockId="22.[151,1436,1781,2022]" pageId="22" pageNumber="123">
Measurements (in mm). Body length 
<number box="[623,707,1782,1806]" pageId="22" pageNumber="123" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4</number>
(♂) or 
<number box="[793,878,1781,1806]" pageId="22" pageNumber="123" value="1.4500000000000002" valueMax="1.6" valueMin="1.3">1.3–1.6</number>
(♀). Carapace 
<number box="[1052,1164,1782,1806]" pageId="22" pageNumber="123" value="0.41000000000000003" valueMax="0.42" valueMin="0.4">0.40–0.42</number>
× 
<number box="[1195,1307,1782,1806]" pageId="22" pageNumber="123" value="0.4" valueMax="0.42" valueMin="0.38">0.38–0.42</number>
(
<number box="[1322,1436,1782,1806]" pageId="22" pageNumber="123" value="0.35" valueMax="0.37" valueMin="0.33">0.33–0.37</number>
anteriorly) (♂) or 
<number box="[368,480,1818,1843]" pageId="22" pageNumber="123" value="0.435" valueMax="0.45" valueMin="0.42">0.42–0.45</number>
× 
<number box="[520,632,1818,1842]" pageId="22" pageNumber="123" value="0.43" valueMax="0.44" valueMin="0.42">0.42–0.44</number>
(
<number box="[652,766,1818,1842]" pageId="22" pageNumber="123" value="0.395" valueMax="0.41" valueMin="0.38">0.38–0.41</number>
anteriorly) (♀). Chelicera 
<number box="[1085,1197,1818,1843]" pageId="22" pageNumber="123" value="0.32499999999999996" valueMax="0.35" valueMin="0.3">0.30–0.35</number>
× 
<number box="[1237,1349,1818,1843]" pageId="22" pageNumber="123" value="0.15000000000000002" valueMax="0.16" valueMin="0.14">0.14–0.16</number>
(♂) or 
<number box="[151,263,1853,1878]" pageId="22" pageNumber="123" value="0.36" valueMax="0.37" valueMin="0.35">0.35–0.37</number>
× 
<number box="[296,408,1854,1878]" pageId="22" pageNumber="123" value="0.175" valueMax="0.18" valueMin="0.17">0.17–0.18</number>
(♀); movable finger length 
<number box="[732,844,1853,1878]" pageId="22" pageNumber="123" value="0.16499999999999998" valueMax="0.18" valueMin="0.15">0.15–0.18</number>
(♂) or 
<number box="[933,1045,1854,1878]" pageId="22" pageNumber="123" value="0.195" valueMax="0.2" valueMin="0.19">0.19–0.20</number>
(♀). Pedipalp: femur 
<number box="[1300,1411,1853,1878]" pageId="22" pageNumber="123" value="0.515" valueMax="0.57" valueMin="0.46">0.46–0.57</number>
× 
<number box="[151,291,1890,1915]" pageId="22" pageNumber="123" value="0.1" valueMax="0.105" valueMin="0.095">0.095–0.105</number>
(♂) or 
<number box="[376,487,1890,1915]" pageId="22" pageNumber="123" value="0.53" valueMax="0.55" valueMin="0.51">0.51–0.55</number>
× 0.100– 
<number box="[596,658,1890,1915]" pageId="22" pageNumber="123" value="0.115">0.115</number>
(♀); chela 
<number box="[786,898,1890,1915]" pageId="22" pageNumber="123" value="0.74" valueMax="0.81" valueMin="0.67">0.67–0.81</number>
× 
<number box="[929,1040,1890,1914]" pageId="22" pageNumber="123" value="0.15500000000000003" valueMax="0.17" valueMin="0.14">0.14–0.17</number>
(♂) or 
<number box="[1126,1238,1890,1915]" pageId="22" pageNumber="123" value="0.77" valueMax="0.79" valueMin="0.75">0.75–0.79</number>
× 
<number box="[1269,1381,1890,1915]" pageId="22" pageNumber="123" value="0.16999999999999998" valueMax="0.18" valueMin="0.16">0.16–0.18</number>
(♀); hand length 
<number box="[303,443,1925,1950]" pageId="22" pageNumber="123" value="0.2575" valueMax="0.285" valueMin="0.23">0.230–0.285</number>
(♂) or 
<number box="[548,688,1925,1950]" pageId="22" pageNumber="123" value="0.275" valueMax="0.285" valueMin="0.265">0.265–0.285</number>
(♀); fixed finger length (from tip to 
<emphasis box="[1156,1182,1925,1950]" italics="true" pageId="22" pageNumber="123">eb</emphasis>
) 
<number box="[1205,1345,1925,1950]" pageId="22" pageNumber="123" value="0.475" valueMax="0.515" valueMin="0.435">0.435–0.515</number>
(♂) or 
<number box="[151,213,1962,1987]" pageId="22" pageNumber="123" value="0.475">0.475</number>
–0.500 (♀); movable finger length 
<number box="[607,719,1962,1987]" pageId="22" pageNumber="123" value="0.45999999999999996" valueMax="0.5" valueMin="0.42">0.42–0.50</number>
(♂) or 
<number box="[802,914,1962,1987]" pageId="22" pageNumber="123" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.45–0.48</number>
(♀).
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="124" pageId="22" pageNumber="123" type="discussion">
<paragraph blockId="22.[151,1436,1781,2022]" lastBlockId="23.[151,1436,151,429]" lastPageId="23" lastPageNumber="124" pageId="22" pageNumber="123">
<emphasis bold="true" box="[199,315,1997,2022]" pageId="22" pageNumber="123">Remarks.</emphasis>
The above redescription of 
<taxonomicName box="[659,784,1997,2022]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="22" pageNumber="123" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis box="[659,784,1997,2022]" italics="true" pageId="22" pageNumber="123">C. pusillus</emphasis>
</taxonomicName>
partially incorporates the original description of 
<bibRefCitation author="Beier" firstAuthor="Beier" lastPageId="23" lastPageNumber="124" pageId="22" pageNumber="123" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
Beier (
<number box="[159,219,152,176]" pageId="23" pageNumber="124" value="1947.0">1947</number>
)
</bibRefCitation>
, which is nearly identical to that in 
<bibRefCitation author="Beier" box="[639,784,151,176]" firstAuthor="Beier" pageId="23" pageNumber="124" refString="Beier, M. (1963) Ordnung Pseudoscorpionidea (Afterskorpione). In: d'Aguilar, J., Beier, M., Franz, H. &amp; Raw, F. (Eds.), Bestimmungsbucher zur Bodenfauna Europas. Vol. 1. Akademie-Verlag, Berlin, VI + 313 pp." type="book" year="1963">
Beier (
<number box="[716,776,151,176]" pageId="23" pageNumber="124" value="1963.0">1963</number>
)
</bibRefCitation>
, with the exception of the discrepancies noted above and the chaetotaxy of the posterior margin of the carapace [lateral setae of posterior row nearly as long as the median ones according to 
<bibRefCitation author="Beier" box="[359,504,223,248]" firstAuthor="Beier" pageId="23" pageNumber="124" refString="Beier, M. (1947) Neue Pseudoscorpione aus der Steiermark. Annalen des Naturhistorischen Museums in Wien, 55, 296 - 301." type="journal article" year="1947">
Beier (
<number box="[436,496,224,248]" pageId="23" pageNumber="124" value="1947.0">1947</number>
)
</bibRefCitation>
, as opposed to being very short microseta or, rarely, half the length of the median macrosetae].
</paragraph>
<paragraph blockId="23.[151,1436,151,429]" pageId="23" pageNumber="124">
<taxonomicName box="[199,408,295,320]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis box="[199,408,295,320]" italics="true" pageId="23" pageNumber="124">Chthonius pusillus</emphasis>
</taxonomicName>
was previously known only from the Austrian states of Styria, Lower 
<collectingCountry box="[1195,1277,295,320]" name="Austria" pageId="23" pageNumber="124">Austria</collectingCountry>
and Carinthia (
<bibRefCitation author="Mahnert" box="[158,320,332,357]" firstAuthor="Mahnert" pageId="23" pageNumber="124" refString="Mahnert, V. (2009) Pseudoscorpiones (Pseudoskorpione). In: Rabitsch, W. &amp; Essl, F. (Eds.), Endemiten-Kostbarkeiten in Osterreichs Pflanzen- und Tierwelt. Naturwissenschaftlicher Verein Kaernten und Umweltbundesamt GmbH, Klagenfurt and Vienna, 924 pp." type="book" year="2009">Mahnert 2009</bibRefCitation>
, 
<bibRefCitation author="Mahnert" box="[331,391,332,356]" firstAuthor="Mahnert" pageId="23" pageNumber="124" refString="Mahnert, V. (2011) Pseudoscorpiones (Arachnida). Checklisten der Fauna Osterreich, 5, 28 - 39." type="journal article" year="2011">2011</bibRefCitation>
). It is newly recorded here from 
<collectingCountry box="[753,851,332,357]" name="Slovenia" pageId="23" pageNumber="124">Slovenia</collectingCountry>
(
<figureCitation box="[866,950,332,357]" captionStart="FIGURE 90" captionStartId="33.[153,252,1653,1675]" captionTargetBox="[167,1422,866,1638]" captionTargetId="figure@33.[165,1424,863,1640]" captionTargetPageId="33" captionText="FIGURE 90. Distribution map for Chthonius (C.) cavernarum Ellingsen, 1909 (▲), C. (C.) heterodactylus Tömösváry, 1882 (Δ), C. (C.) hungaricus Mahnert, 1981 (■), C. (C.) lessiniensis Schawaller, 1982 (□), C. (C.) pusillus Beier, 1947 (●), C. (C.) raridentatus Hadži, 1930 (○)." httpUri="https://zenodo.org/record/228068/files/figure.png" pageId="23" pageNumber="124">
Fig. 
<number box="[918,950,332,356]" pageId="23" pageNumber="124" value="90.0">90</number>
</figureCitation>
). Among the species of 
<taxonomicName box="[1218,1333,332,357]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis box="[1218,1333,332,357]" italics="true" pageId="23" pageNumber="124">Chthonius</emphasis>
</taxonomicName>
of the 
<taxonomicName class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="heterodactylus">
<emphasis italics="true" pageId="23" pageNumber="124">C. heterodactylus</emphasis>
</taxonomicName>
group, the affinities of 
<taxonomicName box="[577,697,367,392]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="124" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis box="[577,697,367,392]" italics="true" pageId="23" pageNumber="124">C. pusillus</emphasis>
</taxonomicName>
are uncertain. It can be distinguished from the other species using the key given above.
</paragraph>
</subSubSection>
</treatment>
</document>