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<document id="3AC4AA91299C0CF78E146E183D122F38" ID-DOI="10.5852/ejt.2024.926.2469" ID-GBIF-Dataset="4511b4bd-f026-49d3-9d4d-a39875e75cc5" ID-ISSN="2118-9773" ID-Zenodo-Dep="10847806" ID-ZooBank="A00D7669-2151-46AE-A066-9AF4D0F2BEEE" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1711011351298" checkinUser="plazi" docAuthor="Jochum, Adrienne, Michalik, Peter, Inäbnit, Thomas, Kneubühler, Jeannette, Slapnik, Rajko, Vrabec, Marko, Schilthuizen, Menno &amp; Ruthensteiner, Bernhard" docDate="2024" docId="03B0879DFFADFFDBFE26D26EFDB6FBBF" docLanguage="en" docName="EJT.2024.926.1-62.pdf" docOrigin="European Journal of Taxonomy 926" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zospeum tortuosum Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner 2024, sp. nov." docType="treatment" docUuid="BDD1308E-7D4C-447C-8009-E60D76AB38D2" docUuidSource="ZooBank" docVersion="12" lastPageNumber="28" masterDocId="FF89FFE5FFBAFFC0FFFFD700FFD4FFD4" masterDocTitle="3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="24" updateTime="1722640713706" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7919BF85E421D941AE532AE8D5ED4D71">3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)</mods:title>
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<mods:namePart id="C861C8C2F531C58E101F444FA6BF9ECE">Jochum, Adrienne</mods:namePart>
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<mods:affiliation id="BD1C865ED871F03A742F7D2558D77123">Senckenberg Research Institute and Natural History Museum, Senckenberganlage 25, D- 60325 Frankfurt / M., Germany.</mods:affiliation>
<mods:nameIdentifier id="F8455933ACB82220928196A6659FA826" type="email">nne.jochum@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="0071E3D9A277070B936ABF0AF8822D82">Michalik, Peter</mods:namePart>
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<mods:affiliation id="4256945A2B87C15DCF46411333E7A8CA">Zoological Institute and Museum, University of Greifswald, Loitzer Strasse 26, D- 17489 Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="1F5C5857BCB2432236BF0411839753EB" type="email">michalik@uni-greifswald.de</mods:nameIdentifier>
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<mods:namePart id="77A168C5F8B999C2710368106CC8B022">Inäbnit, Thomas</mods:namePart>
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<mods:affiliation id="D0B77E9C0581CAD5ADA88AAD6340F0DF">Institute for Biochemistry &amp; Biology, University of Potsdam, Karl-Liebknecht-Strasse 24 - 25, House 26, D- 14476 Potsdam, Germany.</mods:affiliation>
<mods:nameIdentifier id="7F350DC3E5ABF9D7C31B2CBC0ED8FB3C" type="email">inaebnit.thomas@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9BB265C39B16F6D81E617BED63692CED">Kneubühler, Jeannette</mods:namePart>
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<mods:affiliation id="B503FF151A97E7E9782BCCF8AC3B5AE1">Naturhistorisches Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland. &amp; Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="EC483002DFEF7AC1F9C088DBD3B3E25B" type="email">jeannette.kneubuehler@nmbe.ch</mods:nameIdentifier>
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<mods:namePart id="4A0839932B91FE5E66FDB308252713F3">Slapnik, Rajko</mods:namePart>
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<mods:affiliation id="FD658FF19AD211DBE8E4B9D567BB14FB">ZOSPEUM, Molluscs, Cave &amp; Karst Biological Consulting, Drnovškova pot 2, Mekinje, SI 1240 Kamnik, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="AFC68EC08AA59629BFFB793CE9D47D06" type="email">rajko.slapnik@gmail.com</mods:nameIdentifier>
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<mods:namePart id="461E3D2D2B4735E6E60E001DC27D86A0">Vrabec, Marko</mods:namePart>
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<mods:affiliation id="97FB1AD226E25672E5E997549B2861C2">Department of Geology, Faculty of Natural Sciences and Engineering, Aškerčeva 12, University of Ljubljana, 1000 Ljubljana, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="0CBCF209623750EFC6D48FD770E969AF" type="email">marko.vrabec@geo.ntf.uni-lj.si</mods:nameIdentifier>
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<mods:namePart id="5A3B05FC44639C171290FC7135184C3B">Schilthuizen, Menno</mods:namePart>
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<mods:affiliation id="7051A2694DE2CB64694A6DE1A3A4B01A">Naturalis Biodiversity Center and Leiden University, Vondellaan 55, NL- 2332 AA Leiden, The Netherlands.</mods:affiliation>
<mods:nameIdentifier id="D4BFFBAA4CAE90316AA52B9E3AACE66F" type="email">menno.schilthuizen@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="0503B1D31C9142E58C21816641B04A7E">Ruthensteiner, Bernhard</mods:namePart>
<mods:nameIdentifier id="6C592C7CCDF0AD2005956712B91A7B5D" type="LSID">9A8A56B0-19FA-42AB-A85E-4FCA98BF4A6F</mods:nameIdentifier>
<mods:affiliation id="51D53526027689E5C03CC464B8A59BE1">SNSB - Zoologische Staatssammlung München (ZSM), 81247 München, Germany.</mods:affiliation>
<mods:nameIdentifier id="38192C2C661ED39E15835B6ADF2FC4E7" type="email">ruthensteiner@snsb.de</mods:nameIdentifier>
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<mods:title id="26D0993E790EDF6CAB684E60A630EA8A">European Journal of Taxonomy</mods:title>
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<mods:part id="4D5C99A48D36BF7D1C6F15EC723B3764">
<mods:date id="75B5B3082FB98AC74FBD4EBDA95E0134">2024</mods:date>
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<mods:number id="5DAD275FBECDD520D38909D5F1786A7F">2024-03-20</mods:number>
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<mods:number id="E0E58322885765E4C5E870B76C46F667">926</mods:number>
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<mods:start id="B3347EED65AB3E573F86D196AA2DB6C1">1</mods:start>
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<mods:url id="A5D59DAB4878D728875DD3CF540AD5FB">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973</mods:url>
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<mods:identifier id="2BC575223FD28D4BF29BAC7FA41DBDA1" type="DOI">10.5852/ejt.2024.926.2469</mods:identifier>
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<treatment id="03B0879DFFADFFDBFE26D26EFDB6FBBF" ID-DOI="http://doi.org/10.5281/zenodo.10854255" ID-GBIF-Taxon="224067508" ID-Zenodo-Dep="10854255" LSID="urn:lsid:zoobank.org:act:BDD1308E-7D4C-447C-8009-E60D76AB38D2" httpUri="http://treatment.plazi.org/id/03B0879DFFADFFDBFE26D26EFDB6FBBF" lastPageId="27" lastPageNumber="28" pageId="23" pageNumber="24">
<subSubSection id="C3036500FFADFFD7FE26D26EFB8FFA5C" box="[473,1115,1389,1416]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BA6368BFFADFFD7FE26D26EFB8FFA5C" blockId="23.[360,1227,1389,1486]" box="[473,1115,1389,1416]" pageId="23" pageNumber="24">
<heading id="D0EE81E7FFADFFD7FE26D26EFB8FFA5C" box="[473,1115,1389,1416]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="4C194D08FFADFFD7FE26D26EFC2DFA53" ID-CoL="C8NTF" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[473,1017,1389,1416]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="tortuosum" status="sp. nov.">
<emphasis id="B96DEA99FFADFFD7FE26D26EFD19FA5C" bold="true" box="[473,717,1390,1416]" italics="true" pageId="23" pageNumber="24">Zospeum tortuosum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFADFFD7FBFFD26EFB8FFA5C" box="[1024,1115,1390,1416]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFADFFD7FE97D290FC57FA1A" pageId="23" pageNumber="24" type="description">
<paragraph id="8BA6368BFFADFFD7FE97D290FB1FFA7E" blockId="23.[360,1227,1389,1486]" box="[360,1227,1424,1450]" pageId="23" pageNumber="24">
<uri id="FF883A89FFADFFD7FE97D290FB1FFA7E" box="[360,1227,1424,1450]" pageId="23" pageNumber="24">
urn:lsid:zoobank.org:act:
<uuid id="FFBF0C5EFFADFFD7FD78D290FB1FFA7E" box="[647,1227,1424,1450]" pageId="23" pageNumber="24">BDD1308E-7D4C-447C-8009-E60D76AB38D2</uuid>
</uri>
</paragraph>
<paragraph id="8BA6368BFFADFFD7FD4ED2B4FC57FA1A" blockId="23.[360,1227,1389,1486]" box="[689,899,1460,1486]" pageId="23" pageNumber="24">
<figureCitation id="13222A0EFFADFFD7FD4ED2B4FCE7FA1A" box="[689,819,1460,1486]" captionStart="Fig" captionStartId="24.[189,232,1787,1813]" captionTargetBox="[189,1398,1155,1748]" captionTargetId="figure-235@24.[189,1398,1155,1748]" captionTargetPageId="24" captionText="Fig. 3. Light microscopic images of shells from the caves Benetina pećina (SE Bosnia and Herzegovina) and Cetinska pećina (Montenegro) with specimen labels. A. Zospeum troglobalcanicum Absolon, 1916, lectotype (NHMW Moll.Coll.Edlauer 32.749) from Benetina pećina. B–C. Zospeum tortuosum Jochum &amp; Ruthensteiner sp. nov., paratypes (MCSMNH-PMSL-Moll.-FVelkovrh 34099[spm2-3]) from Cetinska pećina. D. Zospeum tortuosum Jochum &amp; Ruthensteiner sp. nov., holotype (MCSMNH- PMSL-Moll.-FVelkovrh 34099[spm1]) from Cetinska pećina. E. Deformed shell (MCSMNH-PMSL- Moll.-FVelkovrh 34099[spm4]) from Cetinska pećina." figureDoi="http://doi.org/10.5281/zenodo.10847812" httpUri="https://zenodo.org/record/10847812/files/figure.png" pageId="23" pageNumber="24">Figs 3B–D</figureCitation>
,
<figureCitation id="13222A0EFFADFFD7FCC0D2B4FC57FA1A" box="[831,899,1460,1486]" captionStart="Fig" captionStartId="25.[190,233,1752,1778]" captionTargetBox="[202,1385,472,1710]" captionTargetId="figure-73@25.[202,1385,471,1710]" captionTargetPageId="25" captionText="Fig. 4. 3D visualizations using Micro-CT data of shells from the caves Benetina pećina (SE Bosnia and Herzegovina) and Cetinska pećina (Montenegro). A–F. Zospeum troglobalcanicum Absolon, 1916, lectotype (NHMW Moll.Coll.Edlauer 32.749). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view showing umbilical depression. G–L. Zospeum tortuosum Jochum &amp; Ruthensteiner sp. nov., holotype (MCSMNH-PMSL- Moll.-FVelkovrh 34099[spm1]). G. Aperture view. H. Aperture facing right view. I. Apical view. J. Dorsal view. K. Aperture facing left view. L. Ventral view showing callused peristome positioned mostly on top of umbilicus." figureDoi="http://doi.org/10.5281/zenodo.10847814" httpUri="https://zenodo.org/record/10847814/files/figure.png" pageId="23" pageNumber="24">4G–L</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFADFFD7FF42D2F5FCC4F973" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BA6368BFFADFFD7FF42D2F5FEE3F9DB" blockId="23.[189,311,1525,1551]" box="[189,311,1525,1551]" pageId="23" pageNumber="24">
<heading id="D0EE81E7FFADFFD7FF42D2F5FEE3F9DB" bold="true" box="[189,311,1525,1551]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B96DEA99FFADFFD7FF42D2F5FEE3F9DB" bold="true" box="[189,311,1525,1551]" pageId="23" pageNumber="24">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFFADFFD7FF42D124FCC4F973" blockId="23.[189,1399,1570,1704]" pageId="23" pageNumber="24">
Shell ca
<quantity id="4CE19B6EFFADFFD7FEDAD124FE50F9EA" box="[293,388,1572,1598]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="24" unit="mm" value="1.4">1.4 mm</quantity>
, conical or elongate-conical with ca 5¼ regularly coiled, slightly shouldered whorls; last coiling of shell twists a fraction beyond the frontal plane ca ⅛ the width of body whorl; columella slanted and not centrally aligned; no lamella; ventral side swollen with columellar side of peristome heavily callused on top of and plugging umbilicus.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFADFFD7FF42D1CFFAC2F8EE" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BA6368BFFADFFD7FF42D1CFFE90F93D" blockId="23.[189,324,1743,1769]" box="[189,324,1743,1769]" pageId="23" pageNumber="24">
<heading id="D0EE81E7FFADFFD7FF42D1CFFE90F93D" bold="true" box="[189,324,1743,1769]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B96DEA99FFADFFD7FF42D1CFFE90F93D" bold="true" box="[189,324,1743,1769]" pageId="23" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFFADFFD7FF42D1FDFAC2F8EE" blockId="23.[189,1398,1788,1851]" pageId="23" pageNumber="24">This species is named after the characteristic ⅛ additional turn of the last whorl. Due to the twisted offcenter alignment of the columella, the last whorl and the aperture are slightly turned to the left.</paragraph>
</subSubSection>
<subSubSection id="C3036500FFA2FFD8FF42D609FEAFFD97" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BA6368BFFA2FFD8FF42D609FEB8FEF6" blockId="24.[189,364,264,291]" box="[189,364,264,291]" pageId="24" pageNumber="25">
<heading id="D0EE81E7FFA2FFD8FF42D609FEB8FEF6" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B96DEA99FFA2FFD8FF42D609FEB8FEF6" bold="true" box="[189,364,264,291]" pageId="24" pageNumber="25">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFFA2FFD8FF15D637FE88FE85" blockId="24.[186,1399,311,443]" box="[234,348,311,337]" pageId="24" pageNumber="25">
<emphasis id="B96DEA99FFA2FFD8FF15D637FE88FE85" bold="true" box="[234,348,311,337]" pageId="24" pageNumber="25">
<typeStatus id="54A28829FFA2FFD8FF15D637FE88FE85" box="[234,348,311,337]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA6368BFFA2FFD8FF42D65AFE8AFE6F" blockId="24.[186,1399,311,443]" pageId="24" pageNumber="25">
<materialsCitation id="3B713CD6FFA2FFD8FF42D65AFE8AFE6F" ID-GBIF-Occurrence="4598261328" collectingDate="1976-09" collectionCode="MCSMNH-PMSL-Moll." collectorName="F. Velkovrh &amp; FVelkovrh" country="Serbia" latitude="42.3886" location="Cetinska cave" longLatPrecision="6" longitude="18.9208" municipality="Cetinje" pageId="24" pageNumber="25" specimenCode="MCSMNH-PMSL-Moll.-FVelkovrh 34099" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F30E761BFFA2FFD8FF42D65AFE5EFEA0" box="[189,394,346,372]" name="Serbia" pageId="24" pageNumber="25">MONTENEGRO</collectingCountry>
• “Cetinska pećina, Cetinje, vhod; Mtg.” [
<location id="8EC66050FFA2FFD8FC31D65AFB50FEA1" LSID="urn:lsid:plazi:treatment:03B0879DFFADFFDBFE26D26EFDB6FBBF:8EC66050FFA2FFD8FC31D65AFB50FEA1" box="[974,1156,346,373]" country="Serbia" latitude="42.3886" longLatPrecision="6" longitude="18.9208" municipality="Cetinje" name="Cetinska cave" pageId="24" pageNumber="25">Cetinska cave</location>
,
<collectingMunicipality id="6BC2ACF1FFA2FFD8FB63D65AFB20FEA0" box="[1180,1268,346,372]" pageId="24" pageNumber="25">Cetinje</collectingMunicipality>
, entrance; Montenegro]; [
<geoCoordinate id="EE2D504CFFA2FFD8FE95D67DFE24FE43" box="[362,496,381,407]" degrees="42.3886" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="42.3886">42.3886°N</geoCoordinate>
,
<geoCoordinate id="EE2D504CFFA2FFD8FE06D67EFDA8FE4C" box="[505,636,382,408]" degrees="18.9208" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="18.9208">18.9208°E</geoCoordinate>
];
<date id="FFA7104BFFA2FFD8FD72D67EFCD7FE43" box="[653,771,381,408]" pageId="24" pageNumber="25" value="1976-09">
<collectingDate id="EFE3E9A3FFA2FFD8FD72D67EFCD7FE43" box="[653,771,381,408]" pageId="24" pageNumber="25" value="1976-09">Sep. 1976</collectingDate>
</date>
;
<collectorName id="26EC535DFFA2FFD8FCF3D67EFC46FE43" box="[780,914,381,408]" pageId="24" pageNumber="25">F. Velkovrh</collectorName>
leg.;
<collectionCode id="ED08AE4EFFA2FFD8FC31D67DFB24FE43" box="[974,1264,381,407]" pageId="24" pageNumber="25">MCSMNH-PMSL-Moll.</collectionCode>
-
<collectorName id="26EC535DFFA2FFD8FB08D67DFAA3FE43" box="[1271,1399,381,407]" pageId="24" pageNumber="25">FVelkovrh</collectorName>
34099[spm1].
</materialsCitation>
</paragraph>
<paragraph id="8BA6368BFFA2FFD8FF15D6E2FEB3FE28" blockId="24.[189,1399,482,579]" box="[234,359,482,508]" pageId="24" pageNumber="25">
<emphasis id="B96DEA99FFA2FFD8FF15D6E2FEB3FE28" bold="true" box="[234,359,482,508]" pageId="24" pageNumber="25">
<typeStatus id="54A28829FFA2FFD8FF15D6E2FEB3FE28" box="[234,359,482,508]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA6368BFFA2FFD8FF42D505FEAFFD97" blockId="24.[189,1399,482,579]" pageId="24" pageNumber="25">
<materialsCitation id="3B713CD6FFA2FFD8FF42D505FEAFFD97" ID-GBIF-Occurrence="4598261315" collectionCode="MCSMNH-PMSL-Moll." collectorName="FVelkovrh" country="Serbia" location="MONTENEGRO" pageId="24" pageNumber="25" specimenCode="MCSMNH-PMSL-Moll.-FVelkovrh 34099" specimenCount="2">
<collectingCountry id="F30E761BFFA2FFD8FF42D505FE5EFDCB" box="[189,394,517,543]" name="Serbia" pageId="24" pageNumber="25">MONTENEGRO</collectingCountry>
•
<specimenCount id="9D1FFD02FFA2FFD8FE59D505FDD5FDF4" box="[422,513,517,544]" count="2" pageId="24" pageNumber="25" type="generic">2 specs</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED08AE4EFFA2FFD8FC31D505FB24FDCB" box="[974,1264,517,543]" pageId="24" pageNumber="25">MCSMNH-PMSL-Moll.</collectionCode>
-
<collectorName id="26EC535DFFA2FFD8FB08D505FAA3FDCB" box="[1271,1399,517,543]" pageId="24" pageNumber="25">FVelkovrh</collectorName>
34099[spm2-3].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFA2FFDAFF42D56AFD35FEBE" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
<paragraph id="8BA6368BFFA2FFD8FF42D56AFE9AFD50" blockId="24.[189,334,618,644]" box="[189,334,618,644]" pageId="24" pageNumber="25">
<heading id="D0EE81E7FFA2FFD8FF42D56AFE9AFD50" bold="true" box="[189,334,618,644]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B96DEA99FFA2FFD8FF42D56AFE9AFD50" bold="true" box="[189,334,618,644]" pageId="24" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFFA2FFD8FF42D599FC73FD2D" blockId="24.[188,1398,664,762]" pageId="24" pageNumber="25">
<smallCapsWord id="8D40A057FFA2FFD8FF42D599FEBAFD65" baselines="684,684" box="[189,366,665,691]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="24" pageNumber="25">MEASUREMENTS</smallCapsWord>
.
<typeStatus id="54A28829FFA2FFD8FE83D598FE3AFD66" box="[380,494,664,690]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
: sh:
<quantity id="4CE19B6EFFA2FFD8FDDED598FD4EFD67" box="[545,666,664,691]" metricMagnitude="-3" metricUnit="m" metricValue="1.435" pageId="24" pageNumber="25" unit="mm" value="1.435">1.435 mm</quantity>
; sw:
<quantity id="4CE19B6EFFA2FFD8FD2AD599FC9AFD67" box="[725,846,665,691]" metricMagnitude="-4" metricUnit="m" metricValue="9.44" pageId="24" pageNumber="25" unit="mm" value="0.944">0.944 mm</quantity>
; ah:
<quantity id="4CE19B6EFFA2FFD8FC7BD598FC2AFD67" box="[900,1022,664,691]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" pageId="24" pageNumber="25" unit="mm" value="0.515">0.515 mm</quantity>
; aw:
<quantity id="4CE19B6EFFA2FFD8FBC5D598FB66FD67" box="[1082,1202,664,691]" metricMagnitude="-4" metricUnit="m" metricValue="5.11" pageId="24" pageNumber="25" unit="mm" value="0.511">0.511 mm</quantity>
; hlw:
<quantity id="4CE19B6EFFA2FFD8FB07D599FAA6FD67" box="[1272,1394,665,691]" metricMagnitude="-4" metricUnit="m" metricValue="8.41" pageId="24" pageNumber="25" unit="mm" value="0.841">0.841 mm</quantity>
; SA: 57.93 deg.
<typeStatus id="54A28829FFA2FFD8FE8CD5BCFE31FD02" box="[371,485,700,726]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
(N = 2): sh:
<quantity id="4CE19B6EFFA2FFD8FD84D5BCFCE3FD02" box="[635,823,700,726]" metricMagnitude="-3" metricUnit="m" metricValue="1.3895" metricValueMax="1.43" metricValueMin="1.349" pageId="24" pageNumber="25" unit="mm" value="1.3895" valueMax="1.43" valueMin="1.349">1.349–1.43 mm</quantity>
; sw:
<quantity id="4CE19B6EFFA2FFD8FC8CD5BBFBEAFD02" box="[883,1086,699,726]" metricMagnitude="-4" metricUnit="m" metricValue="9.615" metricValueMax="9.719999999999999" metricValueMin="9.51" pageId="24" pageNumber="25" unit="mm" value="0.9615" valueMax="0.972" valueMin="0.951">
<geoCoordinate id="EE2D504CFFA2FFD8FC8CD5BBFC67FD02" box="[883,947,699,726]" degrees="0.951" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="55" value="0.951">0.951</geoCoordinate>
<geoCoordinate id="EE2D504CFFA2FFD8FC4CD5BBFBD2FD02" box="[947,1030,699,726]" degrees="0.972" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="55" value="-0.972">–0.972</geoCoordinate>
mm
</quantity>
; ah:
<quantity id="4CE19B6EFFA2FFD8FB8BD5BBFAEBFD02" box="[1140,1343,699,726]" metricMagnitude="-4" metricUnit="m" metricValue="5.535" metricValueMax="5.59" metricValueMin="5.48" pageId="24" pageNumber="25" unit="mm" value="0.5535" valueMax="0.559" valueMin="0.548">
<geoCoordinate id="EE2D504CFFA2FFD8FB8BD5BBFB62FD02" box="[1140,1206,699,726]" degrees="0.548" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="55" value="0.548">0.548</geoCoordinate>
<geoCoordinate id="EE2D504CFFA2FFD8FB49D5BBFAD3FD02" box="[1206,1287,699,726]" degrees="0.559" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="55" value="-0.559">–0.559</geoCoordinate>
mm
</quantity>
; aw:
<quantity id="4CE19B6EFFA2FFD8FF42D5DFFE5CFD2D" box="[189,392,735,762]" metricMagnitude="-4" metricUnit="m" metricValue="5.92" metricValueMax="6.09" metricValueMin="5.75" pageId="24" pageNumber="25" unit="mm" value="0.592" valueMax="0.609" valueMin="0.575">
<geoCoordinate id="EE2D504CFFA2FFD8FF42D5DFFF2AFD2E" box="[189,254,735,762]" degrees="0.575" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="55" value="0.575">0.575</geoCoordinate>
<geoCoordinate id="EE2D504CFFA2FFD8FF01D5DFFE9BFD2E" box="[254,335,735,762]" degrees="0.609" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="55" value="-0.609">–0.609</geoCoordinate>
mm
</quantity>
; hlw:
<quantity id="4CE19B6EFFA2FFD8FE30D5DFFD4EFD2D" box="[463,666,735,761]" metricMagnitude="-4" metricUnit="m" metricValue="8.354999999999999" metricValueMax="8.44" metricValueMin="8.27" pageId="24" pageNumber="25" unit="mm" value="0.8354999999999999" valueMax="0.844" valueMin="0.827">
<geoCoordinate id="EE2D504CFFA2FFD8FE30D5DFFDC4FD2D" box="[463,528,735,761]" degrees="0.827" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="55" value="0.827">0.827</geoCoordinate>
<geoCoordinate id="EE2D504CFFA2FFD8FDEED5DFFDB6FD2D" box="[529,610,735,761]" degrees="0.844" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="55" value="-0.844">–0.844</geoCoordinate>
mm
</quantity>
; SA: 61.20–65.16 deg.
</paragraph>
<paragraph id="8BA6368BFFA2FFDAFF42D420FD35FEBE" blockId="24.[188,1400,799,1108]" lastBlockId="26.[188,1399,265,362]" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
Shell conical, height
<quantity id="4CE19B6EFFA2FFD8FE3DD420FDA5FCEF" box="[450,625,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="1.395" metricValueMax="1.44" metricValueMin="1.35" pageId="24" pageNumber="25" unit="mm" value="1.395" valueMax="1.44" valueMin="1.35">1.35–1.44 mm</quantity>
with 5¼ convex whorls. Whorls unevenly formed and slightly shouldered; protoconch bulbous; spire angle narrow; last whorl coils beyond the frontal plane ca ⅛width of body whorl, right side sometimes almost disappears under spire in apical view; aperture reniform, somewhat narrower than high, no aperture dentition, columellar side straight, not oblique, thickly and irregularly callused; peristome notch at upper parietal and palatal junction; parietal callus (shield) thick, long and straight; palatal rim uniformly thickened to halfway the height of peristome, palatal-basal lip thins to a ridgelike edge, reflected; suture moderately deep; teleoconch generally smooth with occasional surface irregularity; last whorl shows occasional, irregular blunt growth lines at suture with penultimate whorl, extending downwards to ca ⅓ the height of the last whorl; weak radial banding on last whorl (seen in fresh individuals); some ribbing and irregular growth lines immediately behind palatal lip; light irregular ribbing on last whorl; height of last whorl greater than half of shell height; upper peristome rim almost flush with left side of penultimate whorl (in aperture facing left view); columella unevenly formed, moderately thick, not centrally aligned, tightly twisted, abapically attenuated, swollen at base; no umbilical depression is present; umbilicus thickly callused at junction of peristome and base of columella, columellar side of peristome positioned on top of callused umbilicus, base swollen with some puckering; alignment of last ¼ whorl compact.
</paragraph>
</subSubSection>
<caption id="DF666603FFA2FFD8FF42D1FBFCEFF83C" ID-DOI="http://doi.org/10.5281/zenodo.10847812" ID-Zenodo-Dep="10847812" httpUri="https://zenodo.org/record/10847812/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1787,1813]" targetBox="[189,1398,1155,1748]" targetPageId="24">
<paragraph id="8BA6368BFFA2FFD8FF42D1FBFCEFF83C" blockId="24.[189,1399,1787,2024]" pageId="24" pageNumber="25">
<emphasis id="B96DEA99FFA2FFD8FF42D1FBFEDEF8C1" bold="true" box="[189,266,1787,1813]" pageId="24" pageNumber="25">Fig. 3.</emphasis>
Light microscopic images of shells from the caves Benetina pećina (SE Bosnia and Herzegovina) and Cetinska pećina (Montenegro) with specimen labels.
<emphasis id="B96DEA99FFA2FFD8FC74D01EFC74F8EC" bold="true" box="[907,928,1822,1848]" pageId="24" pageNumber="25">A</emphasis>
.
<taxonomicName id="4C194D08FFA2FFD8FC4AD01FFF2EF88F" authority="Absolon, 1916" authorityName="Absolon, 1916 Absolon" authorityYear="1916" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FFA2FFD8FC4AD01FFAD4F8EC" box="[949,1280,1822,1848]" italics="true" pageId="24" pageNumber="25">Zospeum troglobalcanicum</emphasis>
Absolon, 1916
</taxonomicName>
, lectotype (NHMW Moll.Coll.Edlauer 32.749) from Benetina pećina.
<emphasis id="B96DEA99FFA2FFD8FBBDD041FBADF88F" bold="true" box="[1090,1145,1857,1883]" pageId="24" pageNumber="25">B–C</emphasis>
.
<taxonomicName id="4C194D08FFA2FFD8FB75D042FE24F8AB" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="tortuosum" status="sp. nov.">
<emphasis id="B96DEA99FFA2FFD8FB75D042FAA2F88F" box="[1162,1398,1858,1883]" italics="true" pageId="24" pageNumber="25">Zospeum tortuosum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFA2FFD8FE00D065FDB0F8AB" box="[511,612,1893,1919]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, paratypes (MCSMNH-PMSL-Moll.-FVelkovrh 34099[spm2-3]) from Cetinska pećina.
<emphasis id="B96DEA99FFA2FFD8FE38D088FE08F876" bold="true" box="[455,476,1928,1954]" pageId="24" pageNumber="25">D</emphasis>
.
<taxonomicName id="4C194D08FFA2FFD8FE14D089FBD6F876" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[491,1026,1928,1954]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="tortuosum" status="sp. nov.">
<emphasis id="B96DEA99FFA2FFD8FE14D089FD02F876" box="[491,726,1929,1954]" italics="true" pageId="24" pageNumber="25">Zospeum tortuosum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFA2FFD8FBF6D088FBB2F876" box="[1033,1126,1928,1954]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (MCSMNHPMSL-Moll.-FVelkovrh 34099[spm1]) from Cetinska pećina.
<emphasis id="B96DEA99FFA2FFD8FC5BD0ABFC6CF811" bold="true" box="[932,952,1963,1989]" pageId="24" pageNumber="25">E</emphasis>
. Deformed shell (MCSMNH-PMSLMoll.-FVelkovrh 34099[spm4]) from Cetinska pećina.
</paragraph>
</caption>
<caption id="DF666603FFA3FFD9FF41D1D8FDD5F83C" ID-DOI="http://doi.org/10.5281/zenodo.10847814" ID-Zenodo-Dep="10847814" httpUri="https://zenodo.org/record/10847814/files/figure.png" pageId="25" pageNumber="26" startId="25.[190,233,1752,1778]" targetBox="[202,1385,472,1710]" targetPageId="25">
<paragraph id="8BA6368BFFA3FFD9FF41D1D8FDD5F83C" blockId="25.[189,1399,1752,2025]" pageId="25" pageNumber="26">
<emphasis id="B96DEA99FFA3FFD9FF41D1D8FEDBF926" bold="true" box="[190,271,1752,1778]" pageId="25" pageNumber="26">Fig. 4.</emphasis>
3D visualizations using Micro-CT data of shells from the caves Benetina pećina (SE Bosnia and Herzegovina) and Cetinska pećina (Montenegro).
<emphasis id="B96DEA99FFA3FFD9FC98D1FBFC48F8C1" bold="true" box="[871,924,1787,1813]" pageId="25" pageNumber="26">A–F</emphasis>
.
<taxonomicName id="4C194D08FFA3FFD9FC4CD1FCFF2EF8EC" authority="Absolon, 1916" authorityName="Absolon, 1916 Absolon" authorityYear="1916" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FFA3FFD9FC4CD1FCFB2BF8C1" box="[947,1279,1787,1813]" italics="true" pageId="25" pageNumber="26">Zospeum troglobalcanicum</emphasis>
Absolon, 1916
</taxonomicName>
, lectotype (NHMW Moll.Coll.Edlauer 32.749).
<emphasis id="B96DEA99FFA3FFD9FCC8D01EFC98F8EC" bold="true" box="[823,844,1822,1848]" pageId="25" pageNumber="26">A</emphasis>
. Aperture view.
<emphasis id="B96DEA99FFA3FFD9FBECD01EFBF3F8EC" bold="true" box="[1043,1063,1822,1848]" pageId="25" pageNumber="26">B</emphasis>
. Aperture facing right view.
<emphasis id="B96DEA99FFA3FFD9FF41D041FF06F88F" bold="true" box="[190,210,1857,1883]" pageId="25" pageNumber="26">C</emphasis>
. Apical view.
<emphasis id="B96DEA99FFA3FFD9FE74D042FE74F888" bold="true" box="[395,416,1858,1884]" pageId="25" pageNumber="26">D</emphasis>
. Dorsal view.
<emphasis id="B96DEA99FFA3FFD9FDA5D042FDBAF888" bold="true" box="[602,622,1858,1884]" pageId="25" pageNumber="26">E</emphasis>
. Aperture facing left view.
<emphasis id="B96DEA99FFA3FFD9FC34D042FC09F888" bold="true" box="[971,989,1858,1884]" pageId="25" pageNumber="26">F</emphasis>
. Ventral view showing umbilical depression.
<emphasis id="B96DEA99FFA3FFD9FEB5D064FE56F8AA" bold="true" box="[330,386,1892,1918]" pageId="25" pageNumber="26">G–L</emphasis>
.
<taxonomicName id="4C194D08FFA3FFD9FE6DD066FC7EF8AB" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[402,938,1893,1919]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="tortuosum" status="sp. nov.">
<emphasis id="B96DEA99FFA3FFD9FE6DD066FDA9F8AB" box="[402,637,1894,1919]" italics="true" pageId="25" pageNumber="26">Zospeum tortuosum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFA3FFD9FC4DD065FBDBF8AB" box="[946,1039,1893,1919]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (MCSMNH-PMSLMoll.-FVelkovrh 34099[spm1]).
<emphasis id="B96DEA99FFA3FFD9FDB4D088FDB6F876" bold="true" box="[587,610,1928,1954]" pageId="25" pageNumber="26">G</emphasis>
. Aperture view.
<emphasis id="B96DEA99FFA3FFD9FCC8D088FC9AF876" bold="true" box="[823,846,1928,1954]" pageId="25" pageNumber="26">H</emphasis>
. Aperture facing right view.
<emphasis id="B96DEA99FFA3FFD9FB41D088FB1DF876" bold="true" box="[1214,1225,1928,1954]" pageId="25" pageNumber="26">I</emphasis>
. Apical view.
<emphasis id="B96DEA99FFA3FFD9FF42D0ABFF18F811" bold="true" box="[189,204,1963,1989]" pageId="25" pageNumber="26">J</emphasis>
. Dorsal view.
<emphasis id="B96DEA99FFA3FFD9FE82D0ABFE40F811" bold="true" box="[381,404,1963,1989]" pageId="25" pageNumber="26">K</emphasis>
. Aperture facing left view.
<emphasis id="B96DEA99FFA3FFD9FD1DD0ABFD22F811" bold="true" box="[738,758,1963,1989]" pageId="25" pageNumber="26">L</emphasis>
. Ventral view showing callused peristome positioned mostly on top of umbilicus.
</paragraph>
</caption>
<subSubSection id="C3036500FFA0FFDAFF42D690FA95FE0E" box="[189,1345,400,474]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8BA6368BFFA0FFDAFF42D690FE82FE7E" blockId="26.[189,342,400,426]" box="[189,342,400,426]" pageId="26" pageNumber="27">
<heading id="D0EE81E7FFA0FFDAFF42D690FE82FE7E" bold="true" box="[189,342,400,426]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B96DEA99FFA0FFDAFF42D690FE82FE7E" bold="true" box="[189,342,400,426]" pageId="26" pageNumber="27">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFFA0FFDAFF42D6BFFA95FE0E" blockId="26.[189,1345,447,474]" box="[189,1345,447,474]" pageId="26" pageNumber="27">
This species is only known from its
<typeStatus id="54A28829FFA0FFDAFD9AD6C0FD4CFE0E" box="[613,664,448,474]" pageId="26" pageNumber="27">type</typeStatus>
locality, Cetinska pećina, Cetinje in western
<collectingCountry id="F30E761BFFA0FFDAFB55D6C0FAE9FE0E" box="[1194,1341,448,474]" name="Serbia" pageId="26" pageNumber="27">Montenegro</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFA0FFDAFF42D501FE30FBC0" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BA6368BFFA0FFDAFF42D501FEFBFDCF" blockId="26.[189,303,513,539]" box="[189,303,513,539]" pageId="26" pageNumber="27">
<heading id="D0EE81E7FFA0FFDAFF42D501FEFBFDCF" bold="true" box="[189,303,513,539]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B96DEA99FFA0FFDAFF42D501FEFBFDCF" bold="true" box="[189,303,513,539]" pageId="26" pageNumber="27">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFFA0FFDAFF42D52FFABDFD44" blockId="26.[189,1398,559,656]" pageId="26" pageNumber="27">
The species is most similar in shell form and dimensions to
<emphasis id="B96DEA99FFA0FFDAFC70D530FB5EFD9D" box="[911,1162,559,585]" italics="true" pageId="26" pageNumber="27">
<taxonomicName id="4C194D08FFA0FFDAFC70D530FB52FD9D" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[911,1158,559,585]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="troglobalcanicum">Z. troglobalcanicum</taxonomicName>
.
</emphasis>
It differs mostly by the extra turning of the final whorl and the associated sideways twisting of the columella. It differs from
<taxonomicName id="4C194D08FFA0FFDAFF42D576FE7AFD44" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[189,430,630,656]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FFA0FFDAFF42D576FE7AFD44" box="[189,430,630,656]" italics="true" pageId="26" pageNumber="27">Z. troglobalcanicum</emphasis>
</taxonomicName>
in that it does not bear a conspicuous threadlike lamella on its twisted columella.
</paragraph>
<paragraph id="8BA6368BFFA0FFDAFF42D5B7FDD9FD06" blockId="26.[189,525,695,722]" box="[189,525,695,722]" pageId="26" pageNumber="27">
<emphasis id="B96DEA99FFA0FFDAFF42D5B7FE58FD05" bold="true" box="[189,396,695,721]" pageId="26" pageNumber="27">Deformed shell (</emphasis>
<figureCitation id="13222A0EFFA0FFDAFE73D5B7FE3DFD05" box="[396,489,695,721]" captionStart="Fig" captionStartId="24.[189,232,1787,1813]" captionTargetBox="[189,1398,1155,1748]" captionTargetId="figure-235@24.[189,1398,1155,1748]" captionTargetPageId="24" captionText="Fig. 3. Light microscopic images of shells from the caves Benetina pećina (SE Bosnia and Herzegovina) and Cetinska pećina (Montenegro) with specimen labels. A. Zospeum troglobalcanicum Absolon, 1916, lectotype (NHMW Moll.Coll.Edlauer 32.749) from Benetina pećina. B–C. Zospeum tortuosum Jochum &amp; Ruthensteiner sp. nov., paratypes (MCSMNH-PMSL-Moll.-FVelkovrh 34099[spm2-3]) from Cetinska pećina. D. Zospeum tortuosum Jochum &amp; Ruthensteiner sp. nov., holotype (MCSMNH- PMSL-Moll.-FVelkovrh 34099[spm1]) from Cetinska pećina. E. Deformed shell (MCSMNH-PMSL- Moll.-FVelkovrh 34099[spm4]) from Cetinska pećina." figureDoi="http://doi.org/10.5281/zenodo.10847812" httpUri="https://zenodo.org/record/10847812/files/figure.png" pageId="26" pageNumber="27">Figs 3E</figureCitation>
,
<figureCitation id="13222A0EFFA0FFDAFE0AD5B7FDD0FD06" box="[501,516,695,722]" captionStart="Fig" captionStartId="26.[189,232,1892,1918]" captionTargetBox="[189,1398,1249,1852]" captionTargetId="figure-225@26.[189,1398,1248,1852]" captionTargetPageId="26" captionText="Fig. 5. 3D visualizations of Micro-CT data of the deformed shell, (MCSMNH-PMSL-Moll.-FVelkovrh 34099[spm4]) from Cetinska pećina. A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view with umbilicus covered by last quarter of shell whorl." figureDoi="http://doi.org/10.5281/zenodo.10847816" httpUri="https://zenodo.org/record/10847816/files/figure.png" pageId="26" pageNumber="27">5</figureCitation>
)
</paragraph>
<paragraph id="8BA6368BFFA0FFDAFF42D5E6FEE8FCBE" blockId="26.[189,1399,742,874]" pageId="26" pageNumber="27">
A deformed second shell, (MCSMNH-PMSL-Moll.-FVelkovrh 34099[spm4]), similar to that of
<taxonomicName id="4C194D08FFA0FFDAFF42D40AFD02FCF7" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[189,726,777,803]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="tortuosum" status="sp. nov.">
<emphasis id="B96DEA99FFA0FFDAFF42D40AFE7CFCF7" box="[189,424,778,803]" italics="true" pageId="26" pageNumber="27">Zospeum tortuosum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFA0FFDAFD22D409FCECFCF7" box="[733,824,777,803]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
and found in the same lot from Cetinska pećina, is additionally described and considered in chronological context with the image figures of
<taxonomicName id="4C194D08FFA0FFDAFAF3D42DFEE1FCBD" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="tortuosum">
<emphasis id="B96DEA99FFA0FFDAFAF3D42DFEE1FCBD" italics="true" pageId="26" pageNumber="27">Zospeum tortuosum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA6368BFFA0FFDAFF15D491FDE1FC7F" blockId="26.[189,1399,912,1044]" box="[234,565,912,939]" pageId="26" pageNumber="27">
<emphasis id="B96DEA99FFA0FFDAFF15D491FDE1FC7F" bold="true" box="[234,565,912,939]" pageId="26" pageNumber="27">Data of the deformed shell</emphasis>
</paragraph>
<paragraph id="8BA6368BFFA0FFDAFF42D4B4FE30FBC0" blockId="26.[189,1399,912,1044]" pageId="26" pageNumber="27">
The species data are: “Cetinska pećina, Cetinje, vhod; Mtg.” [Cetinska cave, Cetinje, entrance;
<collectingCountry id="F30E761BFFA0FFDAFF42D4D8FE87FC26" box="[189,339,984,1010]" name="Serbia" pageId="26" pageNumber="27">Montenegro</collectingCountry>
]; [
<geoCoordinate id="EE2D504CFFA0FFDAFE8AD4D7FDDFFC26" box="[373,523,983,1010]" degrees="42.3886" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="5" value="42.3886">42.3886° N</geoCoordinate>
,
<geoCoordinate id="EE2D504CFFA0FFDAFDDFD4D7FD65FC26" box="[544,689,983,1010]" degrees="18.9208" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="5" value="18.9208">18.9208° E</geoCoordinate>
];
<date id="FFA7104BFFA0FFDAFD30D4D7FC8DFC25" box="[719,857,983,1009]" pageId="26" pageNumber="27" value="1976-09">Sept. 1976</date>
; F. Velkovrh leg. MCSMNH-PMSL-Moll.- FVelkovrh 34099[spm4].
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFA0FFDBFF15D33CFAE6FCC4" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="8BA6368BFFA0FFDAFF15D33CFD64FB82" blockId="26.[189,1355,1083,1146]" box="[234,688,1083,1110]" pageId="26" pageNumber="27">
<emphasis id="B96DEA99FFA0FFDAFF15D33CFD64FB82" bold="true" box="[234,688,1083,1110]" pageId="26" pageNumber="27">Measurements of the deformed shell</emphasis>
</paragraph>
<paragraph id="8BA6368BFFA0FFDAFF42D35FFA9FFBAD" blockId="26.[189,1355,1083,1146]" box="[189,1355,1119,1146]" pageId="26" pageNumber="27">
Shell: sh:
<quantity id="4CE19B6EFFA0FFDAFECDD35FFE79FBAD" box="[306,429,1119,1145]" metricMagnitude="-3" metricUnit="m" metricValue="1.396" pageId="26" pageNumber="27" unit="mm" value="1.396">1.396 mm</quantity>
; sw:
<quantity id="4CE19B6EFFA0FFDAFE16D35FFDB7FBAD" box="[489,611,1119,1145]" metricMagnitude="-4" metricUnit="m" metricValue="9.22" pageId="26" pageNumber="27" unit="mm" value="0.922">0.922 mm</quantity>
; ah:
<quantity id="4CE19B6EFFA0FFDAFD64D35FFCC1FBAD" box="[667,789,1119,1146]" metricMagnitude="-4" metricUnit="m" metricValue="5.659999999999999" pageId="26" pageNumber="27" unit="mm" value="0.566">0.566 mm</quantity>
; aw:
<quantity id="4CE19B6EFFA0FFDAFCACD35FFC19FBAD" box="[851,973,1119,1146]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="26" pageNumber="27" unit="mm" value="0.59">0.590 mm</quantity>
; hlw:
<quantity id="4CE19B6EFFA0FFDAFBEAD35FFB5BFBAD" box="[1045,1167,1119,1145]" metricMagnitude="-4" metricUnit="m" metricValue="8.36" pageId="26" pageNumber="27" unit="mm" value="0.836">0.836 mm</quantity>
; SA: 60.01 deg.
</paragraph>
<caption id="DF666603FFA0FFDAFF42D064FE9CF83C" ID-DOI="http://doi.org/10.5281/zenodo.10847816" ID-Zenodo-Dep="10847816" httpUri="https://zenodo.org/record/10847816/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1892,1918]" targetBox="[189,1398,1249,1852]" targetPageId="26">
<paragraph id="8BA6368BFFA0FFDAFF42D064FE9CF83C" blockId="26.[188,1399,1892,2024]" pageId="26" pageNumber="27">
<emphasis id="B96DEA99FFA0FFDAFF42D064FEDFF8AB" bold="true" box="[189,267,1892,1919]" pageId="26" pageNumber="27">Fig. 5.</emphasis>
3D visualizations of Micro-CT data of the deformed shell, (MCSMNH-PMSL-Moll.-FVelkovrh 34099[spm4]) from Cetinska pećina.
<emphasis id="B96DEA99FFA0FFDAFD87D088FD59F876" bold="true" box="[632,653,1928,1954]" pageId="26" pageNumber="27">A</emphasis>
. Aperture view.
<emphasis id="B96DEA99FFA0FFDAFCACD088FCB3F876" bold="true" box="[851,871,1928,1954]" pageId="26" pageNumber="27">B</emphasis>
. Aperture facing right view.
<emphasis id="B96DEA99FFA0FFDAFB40D088FB07F876" bold="true" box="[1215,1235,1928,1954]" pageId="26" pageNumber="27">C</emphasis>
. Apical view.
<emphasis id="B96DEA99FFA0FFDAFF43D0ABFF05F811" bold="true" box="[188,209,1963,1989]" pageId="26" pageNumber="27">D</emphasis>
. Dorsal view.
<emphasis id="B96DEA99FFA0FFDAFE81D0ABFE45F811" bold="true" box="[382,401,1963,1989]" pageId="26" pageNumber="27">E</emphasis>
. Aperture facing left view.
<emphasis id="B96DEA99FFA0FFDAFD2AD0ABFD33F811" bold="true" box="[725,743,1963,1989]" pageId="26" pageNumber="27">F</emphasis>
. Ventral view with umbilicus covered by last quarter of shell whorl.
</paragraph>
</caption>
<paragraph id="8BA6368BFFA1FFDBFF15D608FD5EFEF7" blockId="27.[187,1401,264,784]" box="[234,650,264,291]" pageId="27" pageNumber="28">
<emphasis id="B96DEA99FFA1FFDBFF15D608FD5EFEF7" bold="true" box="[234,650,264,291]" pageId="27" pageNumber="28">Description of the deformed shell</emphasis>
</paragraph>
<paragraph id="8BA6368BFFA1FFDBFF42D62CFAE6FCC4" blockId="27.[187,1401,264,784]" pageId="27" pageNumber="28">
Shell conical, height
<quantity id="4CE19B6EFFA1FFDBFE4ED62CFDDEFE92" box="[433,522,300,326]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="28" unit="mm" value="1.4">1.4 mm</quantity>
with 5¼ whorls; transparent. Whorls convex, and slightly shouldered, spire regularly coiled; suture moderately deep; aperture oriform, elliptical ovate, wider than high, interior of aperture heavily callused with wartlike growths on palatal side and in mid parietal region, giving the impression of a wart-like tooth; columellar side of peristome straight and angular; no notch at upper parietal and palatal junction; parietal callus (shield) well defined, narrow, roundish; palatal rim unevenly thin, thickening only at basal-columellar zone; palatal-basal lip thins to a ridgelike edge, narrowly reflected; teleoconch with course irregular growth lines; weak radial banding on penultimate and body whorl; course irregular growth lines present immediately behind palatal-basal lip and extending some distance from peristome rim; height of last whorl greater than half of shell height; upper peristome rim recedes
<superScript id="7C6C9BC3FFA1FFDBFEDFD568FEFDFDA2" attach="right" box="[288,297,616,630]" fontSize="6" pageId="27" pageNumber="28">1</superScript>
/
<subScript id="179D34CEFFA1FFDBFECFD573FEEDFD55" attach="left" box="[304,313,627,641]" fontSize="6" pageId="27" pageNumber="28">5</subScript>
width of penultimate whorl (in aperture facing left view); columella oblique, moderately thick, not centrally aligned, swollen at base; incomplete callused lamella under penultimate whorl; no umbilical depression; umbilicus thickly callused at junction of peristome and base of columella, columellar side of peristome positioned on top of callused umbilicus, base swollen with some puckering; upper palatal rim not reflected (as seen from umbilical view), alignment of last ¼ whorl compact.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FFA1FFDBFF15D437FDB6FBBF" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BA6368BFFA1FFDBFF15D437FDA5FC85" blockId="27.[187,1399,822,1131]" box="[234,625,822,849]" pageId="27" pageNumber="28">
<emphasis id="B96DEA99FFA1FFDBFF15D437FDA5FC85" bold="true" box="[234,625,822,849]" pageId="27" pageNumber="28">Remarks on the deformed shell</emphasis>
</paragraph>
<paragraph id="8BA6368BFFA1FFDBFF42D45AFDB6FBBF" blockId="27.[187,1399,822,1131]" pageId="27" pageNumber="28">
Although this shell has wart-like deformation in the aperture, the elliptical ovate aperture shape and the columella configuration, which is unaffected by the deformation, largely corresponds to that of
<taxonomicName id="4C194D08FFA1FFDBFF42D4A1FD43FC6E" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[189,663,928,954]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="tortuosum" status="sp. nov.">
<emphasis id="B96DEA99FFA1FFDBFF42D4A1FE8EFC6E" box="[189,346,929,954]" italics="true" pageId="27" pageNumber="28">Z. tortuosum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFA1FFDBFD5BD4A1FCD0FC6F" box="[676,772,929,955]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
(MCSMNH-PMSL-Moll.-FVelkovrh 34099[spm1]) from the same cave. Though the aperture shape also resembles that of
<taxonomicName id="4C194D08FFA1FFDBFBD8D4C4FEB4FBD5" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="dubokidoense" status="sp. nov.">
<emphasis id="B96DEA99FFA1FFDBFBD8D4C4FBE3FC09" box="[1063,1079,964,989]" italics="true" pageId="27" pageNumber="28">Z</emphasis>
.
<emphasis id="B96DEA99FFA1FFDBFBB5D4C3FB39FC09" box="[1098,1261,963,989]" italics="true" pageId="27" pageNumber="28">dubokidoense</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFA1FFDBFE97D4E7FE17FBD5" box="[360,451,999,1025]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13222A0EFFA1FFDBFE2CD4E7FDE4FBD5" box="[467,560,999,1025]" captionStart="Fig" captionStartId="30.[188,231,1857,1883]" captionTargetBox="[190,1399,1183,1819]" captionTargetId="figure-365@30.[190,1399,1183,1819]" captionTargetPageId="30" captionText="Fig. 7. Lightmicroscopic images of shells from the caves Lipska pećina and Duboki do (Montenegro) with specimen labels. A–C. Zospeum troglobalcanicum Absolon, 1916 (MCSMNH-PMSL-Moll.-FVelkovrh 29603). D–F. Zospeum dubokidoense Jochum &amp; Ruthensteiner sp. nov. D. Holotype (MCSMNH- PMSL-Moll.-FVelkovrh 30360[spm1]). E–F. Paratypes, Duboki do, Montenegro (MCSMNH-PMSL- Moll.-FVelkovrh 30360[spm2-3])." figureDoi="http://doi.org/10.5281/zenodo.10847822" httpUri="https://zenodo.org/record/10847822/files/figure.png" pageId="27" pageNumber="28">Fig. 7D</figureCitation>
,
<figureCitation id="13222A0EFFA1FFDBFDC2D4E7FDB1FBD5" box="[573,613,999,1025]" captionStart="Fig" captionStartId="31.[189,232,1787,1813]" captionTargetBox="[187,1396,481,1747]" captionTargetId="figure-65@31.[187,1396,481,1747]" captionTargetPageId="31" captionText="Fig. 8. 3D visualizations of Micro-CT data of shells from the caves Lipska pećina and Duboki do (Montenegro). A–F. Zospeum troglobalcanicum Absolon, 1916 (MCSMNH-PMSL-Moll.-FVelkovrh 29603) (Fig. 7A). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view showing reduced umbilical depression. G–L. Zospeum dubokidoense Jochum &amp; Ruthensteiner sp. nov., holotype (MCSMNH-PMSL-Moll.-FVelkovrh 30360[spm1]). G. Aperture view. H. Aperture facing right view. I. Apical view. J. Dorsal view. K. Aperture facing left view. L. Ventral view showing no umbilical depression." figureDoi="http://doi.org/10.5281/zenodo.10847826" httpUri="https://zenodo.org/record/10847826/files/figure.png" pageId="27" pageNumber="28">8G</figureCitation>
), the deformed shell shows narrower and smaller dimensions. Due to the compactness of the last quarter whorl, the straight alignment of the left side of the peristome on top of the umbilical depression and the narrowness of the parietal shield, this shell cannot be unequivocally determined as
<taxonomicName id="4C194D08FFA1FFDBFE97D351FE2BFBBE" authority="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner, 2024" authorityName="Jochum &amp; Michalik &amp; Inäbnit &amp; Kneubühler &amp; Slapnik &amp; Vrabec &amp; Schilthuizen &amp; Ruthensteiner" authorityYear="2024" box="[360,511,1105,1130]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="tortuosum" status="sp. nov.">
<emphasis id="B96DEA99FFA1FFDBFE97D351FE2BFBBE" box="[360,511,1105,1130]" italics="true" pageId="27" pageNumber="28">Z. tortuosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FFA1FFDBFDF8D351FDB6FBBF" box="[519,610,1105,1131]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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