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<document id="FFF93FB8275453B5033E66A8A5346199" ID-DOI="10.2478/aemnp-2019-0031" ID-ISSN="1804-6487" ID-Zenodo-Dep="5339942" ID-ZooBank="E04E98C4-9A0D-4285-9924-2A441CA756F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630185277844" checkinUser="marcus" docAuthor="Hartung, Viktor" docDate="2019" docId="03A6CE7CFFBAFFABC0CEFA60FB34FC4F" docLanguage="en" docName="ActaEntMusNatPra.2019.59.2.403-422.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (2)" docSource="http://dx.doi.org/10.2478/aemnp-2019-0031" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Symphylax handschini subsp. kinabaluensis Hartung 2019, subsp. nov." docType="treatment" docVersion="3" lastPageNumber="420" masterDocId="FF9FB604FFB4FFBAC076FFE1FFA3FFDA" masterDocTitle="New Oriental and Australasian taxa of Colobathristidae (Hemiptera: Heteroptera) in the collection of the State Natural History Museum Stuttgart" masterLastPageNumber="422" masterPageNumber="403" pageNumber="417" updateTime="1699179043883" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="D57DF6168302FEA7417CBAA3FC025B3E">New Oriental and Australasian taxa of Colobathristidae (Hemiptera: Heteroptera) in the collection of the State Natural History Museum Stuttgart</mods:title>
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<mods:roleTerm id="C6716AC099D30B636B8AED0C40E4DE65">Author</mods:roleTerm>
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<mods:namePart id="A251885278EAFEE2D74EA94148D2B440">Hartung, Viktor</mods:namePart>
<mods:affiliation id="ACD1E24EBA1CBDFAC21229CDC3A2F169">State Museum of Natural History Stuttgart (SMNS), Rosenstein &amp; Museum für Naturkunde - Leibniz-Institute for Evolution- and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany;</mods:affiliation>
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<mods:title id="E2E84669D448D83E033D627019571200">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="3E4B190957C960C1626CA6F39DEC6C6B">2019</mods:date>
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<mods:title id="381AB34A2285C2A4447109E2ACAA1C5C">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="C7E869D67E48D7802C1045F1FA260194">2019-08-15</mods:number>
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<mods:number id="7EF1ACCB41B4DCA245489A0355113C75">59</mods:number>
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<mods:classification id="F115E235055189861A941F47355BA129">journal article</mods:classification>
<mods:identifier id="29FD212F829967A21641E178EF5B34E6" type="DOI">10.2478/aemnp-2019-0031</mods:identifier>
<mods:identifier id="0EAB780E151A34BC8825D67DB302BAAB" type="ISSN">1804-6487</mods:identifier>
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<treatment id="03A6CE7CFFBAFFABC0CEFA60FB34FC4F" LSID="urn:lsid:plazi:treatment:03A6CE7CFFBAFFABC0CEFA60FB34FC4F" httpUri="http://treatment.plazi.org/id/03A6CE7CFFBAFFABC0CEFA60FB34FC4F" lastPageId="17" lastPageNumber="420" pageId="14" pageNumber="417">
<subSubSection id="C3152CE1FFBAFFB4C0CEFA60FD74FA43" box="[184,727,1409,1433]" pageId="14" pageNumber="417" type="nomenclature">
<paragraph id="8BB07F6AFFBAFFB4C0CEFA60FD74FA43" blockId="14.[184,727,1409,1433]" box="[184,727,1409,1433]" pageId="14" pageNumber="417">
<heading id="D0F8C806FFBAFFB4C0CEFA60FD74FA43" box="[184,727,1409,1433]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="417" reason="3">
<emphasis id="B97BA378FFBAFFB4C0CEFA60FD74FA43" bold="true" box="[184,727,1409,1433]" pageId="14" pageNumber="417">
<taxonomicName id="4C0F04E9FFBAFFB4C0CEFA60FDF0FA43" ID-CoL="8QM4W" authority="Hartung, 2019" authorityName="Hartung" authorityYear="2019" box="[184,595,1409,1433]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="417" phylum="Arthropoda" rank="subSpecies" species="handschini" status="subsp. nov." subSpecies="kinabaluensis">
<emphasis id="B97BA378FFBAFFB4C0CEFA60FDF0FA43" bold="true" box="[184,595,1409,1433]" italics="true" pageId="14" pageNumber="417">Symphylax handschini kinabaluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2481E03FFBAFFB4C22FFA63FD74FA43" box="[601,727,1410,1433]" pageId="14" pageNumber="417" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3152CE1FFBAFFB4C1F8FA41FDA2FA69" box="[398,513,1440,1459]" pageId="14" pageNumber="417" type="description">
<paragraph id="8BB07F6AFFBAFFB4C1F8FA41FDA2FA69" blockId="14.[151,761,1440,1608]" box="[398,513,1440,1459]" pageId="14" pageNumber="417">
(
<figureCitation id="133463EFFFBAFFB4C1E2FA41FE58FA69" box="[404,507,1440,1459]" captionStart-0="Figs 49–54" captionStart-1="Figs 55–61" captionStart-2="Fig" captionStartId-0="15.[151,188,1983,2002]" captionStartId-1="16.[151,188,1983,2002]" captionStartId-2="17.[150,181,868,887]" captionTargetBox-0="[151,1436,510,1965]" captionTargetBox-1="[316,1271,397,1971]" captionTargetBox-2="[176,755,212,852]" captionTargetId-0="figure-13@15.[151,1436,510,1965]" captionTargetId-1="figure-13@16.[316,1271,397,1971]" captionTargetId-2="figure-841@17.[151,762,189,852]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." captionText-1="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." captionText-2="Fig. 62. Symphylax handschini kinabaluensis subsp. nov., holotype, left paramere, inner view.Abbreviations:al = anterior lobe, pl = posterior lobe." figureDoi-0="http://doi.org/10.5281/zenodo.5339965" figureDoi-1="http://doi.org/10.5281/zenodo.5339967" figureDoi-2="http://doi.org/10.5281/zenodo.5339969" httpUri-0="https://zenodo.org/record/5339965/files/figure.png" httpUri-1="https://zenodo.org/record/5339967/files/figure.png" httpUri-2="https://zenodo.org/record/5339969/files/figure.png" pageId="14" pageNumber="417">Figs. 49–62</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3152CE1FFBAFFB4C0E1FA2BFD5AF992" pageId="14" pageNumber="417" type="materials_examined">
<paragraph id="8BB07F6AFFBAFFB4C0E1FA2BFD5AF992" blockId="14.[151,761,1440,1608]" pageId="14" pageNumber="417">
<emphasis id="B97BA378FFBAFFB4C0E1FA2BFE87FA07" bold="true" box="[151,292,1482,1501]" pageId="14" pageNumber="417">Type material.</emphasis>
<materialsCitation id="3B677537FFBAFFB4C15BFA2BFD5AF992" collectingDate="1996-11-30" collectionCode="SMNS" collectorName="D. Grimm" elevation="575" location="Poring" municipality="Kinabalu" pageId="14" pageNumber="417" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54B4C1C8FFBAFFB4C15BFA2BFE27FA01" box="[301,388,1482,1501]" pageId="14" pageNumber="417" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED1EE7AFFFBAFFB4C1C7FA2BFE52FA07" box="[433,497,1482,1501]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="14" pageNumber="417">SMNS</collectionCode>
), collection number
<collectionCode id="ED1EE7AFFFBAFFB4C2C7FA2BFD4FFA07" box="[689,748,1482,1501]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="14" pageNumber="417">SMNS</collectionCode>
_ HET_00008:“BORNEO:
<collectingRegion id="49CBB188FFBAFFB4C102FA04FE1DFA22" box="[372,446,1509,1528]" country="Malaysia" name="Sabah" pageId="14" pageNumber="417">SABAH</collectingRegion>
,
<collectingMunicipality id="6BD4E510FFBAFFB4C1B3FA04FDB0FA22" box="[453,531,1509,1528]" pageId="14" pageNumber="417">Kinabalu</collectingMunicipality>
/ N. P.,
<location id="8ED029B1FFBAFFB4C226FA04FD28FA22" LSID="urn:lsid:plazi:treatment:03A6CE7CFFBAFFABC0CEFA60FB34FC4F:8ED029B1FFBAFFB4C226FA04FD28FA22" box="[592,651,1509,1528]" municipality="Kinabalu" name="Poring" pageId="14" pageNumber="417" stateProvince="Sabah">Poring</location>
,
<quantity id="4CF7D28FFFBAFFB4C2E7FA04FD49FA22" box="[657,746,1509,1528]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="14" pageNumber="417" unit="m" value="575.0" valueMax="700.0" valueMin="450.0">
<elevation id="00229859FFBAFFB4C2E7FA04FD49FA22" box="[657,746,1509,1528]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="14" pageNumber="417" unit="m" value="575.0" valueMax="700.0" valueMin="450.0">450-700m</elevation>
</quantity>
; /
<date id="FFB159AAFFBAFFB4C0E1FA1EFF5BF9C8" box="[151,248,1535,1554]" pageId="14" pageNumber="417" value="1996-11-30">
<collectingDate id="EFF5A042FFBAFFB4C0E1FA1EFF5BF9C8" box="[151,248,1535,1554]" pageId="14" pageNumber="417" value="1996-11-30">30.XI.1996</collectingDate>
</date>
, leg.
<collectorName id="26FA1ABCFFBAFFB4C157FA1EFED5F9C8" box="[289,374,1535,1554]" pageId="14" pageNumber="417">D.Grimm</collectorName>
[printed in black ink on yellow label]”.Mount- ed on a triangle; pygophore with genitalia (left paramere separated) are dissected and stored in a plastic vial with glycerol attached to the same pin.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3152CE1FFBAFFB4C0E1F981FBF4FBA3" pageId="14" pageNumber="417" type="diagnosis">
<paragraph id="8BB07F6AFFBAFFB4C0E1F981FBF4FBA3" blockId="14.[151,760,1632,2071]" lastBlockId="14.[828,1437,738,2073]" pageId="14" pageNumber="417">
<emphasis id="B97BA378FFBAFFB4C0E1F981FE36F9AD" bold="true" box="[151,405,1632,1655]" pageId="14" pageNumber="417">Differential diagnosis.</emphasis>
With the key from ŠTYS &amp; EX-
<subScript id="178B7D2FFFBAFFB4C0E1F967FF44F94C" attach="left" box="[151,231,1670,1686]" fontSize="7" pageId="14" pageNumber="417">NEROVÁ</subScript>
(2013) the specimen is clearly identified as a
<taxonomicName id="4C0F04E9FFBAFFB4C0E1F941FE16F96D" authority="Horvath, 1904" authorityName="Horvath" authorityYear="1904" box="[151,437,1696,1719]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="417" phylum="Arthropoda" rank="genus">
<emphasis id="B97BA378FFBAFFB4C0E1F941FEAFF96D" box="[151,268,1696,1719]" italics="true" pageId="14" pageNumber="417">Symphylax</emphasis>
<bibRefCitation id="EF9E029BFFBAFFB4C160F941FE16F96D" author="HORVATH G." box="[278,437,1696,1719]" pageId="14" pageNumber="417" pagination="117 - 172" refId="ref13354" refString="HORVATH G. 1904: Monographia Colobathristinarum. Annales Musei Nationalis Hungarici 2: 117 - 172." type="journal article" year="1904">Horváth, 1904</bibRefCitation>
</taxonomicName>
and with the key from
<bibRefCitation id="EF9E029BFFBAFFB4C2B3F941FF4FF90D" author="STYS P." pageId="14" pageNumber="417" pagination="229 - 232" refId="ref13933" refString="STYS P. 1977 b: Symphylax horvathi sp. n. from Borneo (Heteroptera, Colobathristidae). Reichenbachia 16: 229 - 232." type="journal article" year="1977">ŠTYS (1977b)</bibRefCitation>
as
<taxonomicName id="4C0F04E9FFBAFFB4C166F921FD13F90D" authority="Kormilev, 1953" authorityName="Kormilev" authorityYear="1953" box="[272,688,1728,1751]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="handschini">
<emphasis id="B97BA378FFBAFFB4C166F921FE5CF90D" box="[272,511,1728,1751]" italics="true" pageId="14" pageNumber="417">Symphylax handschini</emphasis>
<bibRefCitation id="EF9E029BFFBAFFB4C270F921FD13F90D" author="KORMILEV N. A." box="[518,688,1728,1751]" pageId="14" pageNumber="417" pagination="287 - 292" refId="ref13748" refString="KORMILEV N. A. 1953: Notes on Colobathristidae from Borneo and Java in the collections of the Museum of Natural History in Basle (Hemiptera). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 26: 287 - 292." type="journal article" year="1953">Kormilev, 1953</bibRefCitation>
</taxonomicName>
, based on the structure of genitalia. However, although the hypandrium of the SMNS specimen is very much alike that of the
<typeStatus id="54B4C1C8FFBAFFB4C17CF8C1FEC4F8ED" box="[266,359,1824,1847]" pageId="14" pageNumber="417" type="holotype">holotype</typeStatus>
(and the only described specimen) of
<taxonomicName id="4C0F04E9FFBAFFB4C0E1F8A1FE89F88D" authorityName="Kormilev" authorityYear="1953" box="[151,298,1856,1879]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="handschini">
<emphasis id="B97BA378FFBAFFB4C0E1F8A1FE89F88D" box="[151,298,1856,1879]" italics="true" pageId="14" pageNumber="417">S. handschini</emphasis>
</taxonomicName>
, the structure of paramere shows distinct differences. The concavity on the dorsal surface seems to be closer to the posterior margin than in
<taxonomicName id="4C0F04E9FFBAFFB4C217F861FD52F84D" authorityName="Kormilev" authorityYear="1953" box="[609,753,1920,1943]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="handschini">
<emphasis id="B97BA378FFBAFFB4C217F861FD52F84D" box="[609,753,1920,1943]" italics="true" pageId="14" pageNumber="417">S. handschini</emphasis>
</taxonomicName>
, and the posterior lobe is less well developed than in
<taxonomicName id="4C0F04E9FFBAFFB4C292F841FEAFF80D" authorityName="Kormilev" authorityYear="1953" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="handschini">
<emphasis id="B97BA378FFBAFFB4C292F841FEAFF80D" italics="true" pageId="14" pageNumber="417">S. handschini</emphasis>
</taxonomicName>
. Although the type specimen of
<taxonomicName id="4C0F04E9FFBAFFB4C21EF821FD5BF80D" authorityName="Kormilev" authorityYear="1953" box="[616,760,1984,2007]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="handschini">
<emphasis id="B97BA378FFBAFFB4C21EF821FD5BF80D" box="[616,760,1984,2007]" italics="true" pageId="14" pageNumber="417">S. handschini</emphasis>
</taxonomicName>
was not available, KORMILEV’ s (1953) description of
<taxonomicName id="4C0F04E9FFBAFFB4C292F801FEAFF7CD" authorityName="Kormilev" authorityYear="1953" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="417" phylum="Arthropoda" rank="species" species="handschini">
<emphasis id="B97BA378FFBAFFB4C292F801FEAFF7CD" italics="true" pageId="14" pageNumber="417">S. handschini</emphasis>
</taxonomicName>
shows also differences in somatic characters from the SMNS specimen: “the hind border” of the head is black in the
<typeStatus id="54B4C1C8FFBAFFB4C3B6FCE3FB83FCC3" box="[960,1056,770,793]" pageId="14" pageNumber="417" type="holotype">holotype</typeStatus>
, according to
<bibRefCitation id="EF9E029BFFBAFFB4C4C0FCE3FACCFCC3" author="KORMILEV N. A." box="[1206,1391,770,793]" pageId="14" pageNumber="417" pagination="287 - 292" refId="ref13748" refString="KORMILEV N. A. 1953: Notes on Colobathristidae from Borneo and Java in the collections of the Museum of Natural History in Basle (Hemiptera). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 26: 287 - 292." type="journal article" year="1953">KORMILEV (1953)</bibRefCitation>
, but in the SMNS specimen the head is black as far anterior as the front margin of the eye (
<figureCitation id="133463EFFFBAFFB4C414FCA3FB13FC83" box="[1122,1200,834,857]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Fig. 51</figureCitation>
); supracoxal lobes are whitish-yellow in the
<typeStatus id="54B4C1C8FFBAFFB4C45BFC83FB29FCA3" box="[1069,1162,866,889]" pageId="14" pageNumber="417" type="holotype">holotype</typeStatus>
but brown in the SMNS specimen (
<figureCitation id="133463EFFFBAFFB4C3DBFC63FC5FFC43" box="[941,1020,898,921]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Fig. 52</figureCitation>
); the color of abdomen is orange anteriorly and yellow-brown or reddish-brown posteriorly in the
<typeStatus id="54B4C1C8FFBAFFB4C34AFC23FC3AFC03" box="[828,921,962,985]" pageId="14" pageNumber="417" type="holotype">holotype</typeStatus>
but brown to blackish anteriorly and yellowish posteriorly with some reddish hue medially in the SMNS specimen; labium “almost reaches the middle coxae” (
<bibRefCitation id="EF9E029BFFBAFFB4C335FBC3FC4DFBE3" author="KORMILEV N. A." box="[835,1006,1058,1081]" pageId="14" pageNumber="417" pagination="287 - 292" refId="ref13748" refString="KORMILEV N. A. 1953: Notes on Colobathristidae from Borneo and Java in the collections of the Museum of Natural History in Basle (Hemiptera). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 26: 287 - 292." type="journal article" year="1953">KORMILEV 1953</bibRefCitation>
) in the
<typeStatus id="54B4C1C8FFBAFFB4C43EFBC3FB06FBE3" box="[1096,1189,1058,1081]" pageId="14" pageNumber="417" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EF9E029BFFBAFFB4C4CEFBC3FA95FBE3" author="STYS P." box="[1208,1334,1058,1081]" pageId="14" pageNumber="417" pagination="229 - 232" refId="ref13933" refString="STYS P. 1977 b: Symphylax horvathi sp. n. from Borneo (Heteroptera, Colobathristidae). Reichenbachia 16: 229 - 232." type="journal article" year="1977">ŠTYS 1977b</bibRefCitation>
corrects: “reaches apices of mesocoxae”) and reaches beyond middle coxae in SMNS specimen.
</paragraph>
</subSubSection>
<subSubSection id="C3152CE1FFBAFFABC34AFB63FBB6FCEF" lastPageId="17" lastPageNumber="420" pageId="14" pageNumber="417" type="description">
<paragraph id="8BB07F6AFFBAFFB5C34AFB63FDADFEEF" blockId="14.[828,1437,738,2073]" lastBlockId="15.[151,760,190,469]" lastPageId="15" lastPageNumber="418" pageId="14" pageNumber="417">
<emphasis id="B97BA378FFBAFFB4C34AFB63FBB4FB43" bold="true" box="[828,1047,1153,1177]" pageId="14" pageNumber="417">
Description.
<emphasis id="B97BA378FFBAFFB4C3A0FB60FBB3FB43" bold="true" box="[982,1040,1153,1177]" italics="true" pageId="14" pageNumber="417">Male</emphasis>
.
</emphasis>
<emphasis id="B97BA378FFBAFFB4C454FB63FB00FB43" box="[1058,1187,1154,1177]" italics="true" pageId="14" pageNumber="417">Coloration.</emphasis>
Head (
<figureCitation id="133463EFFFBAFFB4C48CFB63FAEEFB43" box="[1274,1357,1154,1177]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Fig. 51</figureCitation>
) black dorsally from hind margin to anterior margin of eyes, gula black, rest of head yellow with some irregular light brown pattern. Ocelli and eyes red; labium (
<figureCitation id="133463EFFFBAFFB4C4B5FB03FAB2FB23" box="[1219,1297,1250,1273]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Fig. 52</figureCitation>
) yellow with black tip; antennae brown, with first antennomere lighter than the others, its base being almost yellow (
<figureCitation id="133463EFFFBAFFB4C561FAC3FA31FAE3" box="[1303,1426,1314,1337]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Figs 49–50</figureCitation>
). collar and middle lobe of pronotum black (
<figureCitation id="133463EFFFBAFFB4C490FAA3FA93FA83" box="[1254,1328,1346,1369]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Fig. 51</figureCitation>
), posterior lobe yellow except small brownish black macula medially on front margin indicating a start of longitudinal stripe present in some other species; two dark brown maculae on humeral angles; posterior margin of pronotum whitish-transparent. Punctation concolorous on collar, middle pronotal lobe and scutellum and dark on posterior pronotal lobe. Scutellum black at base, getting brown closer to top and yellow on top, as well as base of scutellar spine, its apical two thirds light to dark brown (
<figureCitation id="133463EFFFBAFFB4C3A0F983FB80F9A3" box="[982,1059,1634,1657]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Fig. 52</figureCitation>
). Meso- and metapleura and sternites greyish black (initially covered by dried rests of metathoracic gland secretion), supracoxal lobes dark brown anteriorly and lighter brown posteriorly, peritreme of metathoracic gland brownish black. Bases of fore wings (
<figureCitation id="133463EFFFBAFFB4C56EF903FA37F923" box="[1304,1428,1762,1785]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Figs 49, 51</figureCitation>
) dark brown, rest of fore wings mostly light-coloured: veins yellowish brown, except dark brown distal parts of Cu and Pcu, claval organ, proximal part of RP and distal part of dividing vein; membranous parts transparent. Coxae dark brown at base anteriorly and lighter brown posteriorly, yellow at apex; other leg segments yellowish brown with distal parts of middle and hind tibiae and all tarsi darker brown (
<figureCitation id="133463EFFFBAFFB4C3FDF803FB89F823" box="[907,1066,2018,2041]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="14" pageNumber="417">Figs 49–50, 52</figureCitation>
). Abdomen yellowish for most part, except segments I and II greyish black to brown, sternite and tergite III with large dark brown medial area and sternite IV and tergite IV with dark brown area medioanteriorly; connexivum yellow (
<figureCitation id="133463EFFFBBFFB5C10EFF1FFE50FECF" box="[376,499,254,277]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="15" pageNumber="418">Figs 55–56</figureCitation>
) except segment II dark brown. Pygophore yellow (
<figureCitation id="133463EFFFBBFFB5C1C5FEFFFDA1FEEF" box="[435,514,286,309]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="15" pageNumber="418">Fig. 57</figureCitation>
).
</paragraph>
<paragraph id="8BB07F6AFFBBFFAAC0C3FEDFFE74FEEF" blockId="15.[151,760,190,469]" lastBlockId="16.[151,760,190,373]" lastPageId="16" lastPageNumber="419" pageId="15" pageNumber="418">
<emphasis id="B97BA378FFBBFFB5C0C3FEDFFE29FE8F" box="[181,394,318,341]" italics="true" pageId="15" pageNumber="418">Vestiture and texture</emphasis>
. Head (except the two symmetrical depressions posterolaterally of ocelli and medial sulcus anteriad of them), pronotal collar and middle lobe as well as propleura and prosternum covered by short, relatively sparse, adhering, yellowish setae (
<figureCitation id="133463EFFFBBFFB5C18AFE5FFDE9FE0F" box="[508,586,446,469]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="15" pageNumber="418">Fig. 51</figureCitation>
). Pronotum with punctures, smaller on collar and middle lobe and larger on posterior lobe where each also with a short seta, surface between punctures glabrous. Punctation of scutellum similar to that of posterior lobe, setae missing. Meso- and metapleura and sterna (
<figureCitation id="133463EFFFBBFFB5C436FEDFFB31FE8F" box="[1088,1170,318,341]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="15" pageNumber="418">Fig. 52</figureCitation>
) as densely punctate as posterior pronotal lobe and with as little vestiture, but also carrying mycoid microsculpture typical of evaporatoria (such sculpture also present on abdominal sternite II (greyish coloured area). Antennae and legs with sparse cover of short light setae getting slightly longer and darker on tarsi; sparse short setae on veins. Abdomen with adhering light longer setae on distal segments and especially on hypandrium (
<figureCitation id="133463EFFFA4FFAAC150FEFFFE69FEEF" box="[294,458,286,309]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="16" pageNumber="419">Figs 56–57, 61</figureCitation>
).
</paragraph>
<caption id="DF702FE2FFBBFFB5C0E1F85EFE00F7F8" ID-DOI="http://doi.org/10.5281/zenodo.5339965" ID-Zenodo-Dep="5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="15" pageNumber="418" startId="15.[151,188,1983,2002]" targetBox="[151,1436,510,1965]" targetPageId="15">
<paragraph id="8BB07F6AFFBBFFB5C0E1F85EFE00F7F8" blockId="15.[151,1436,1983,2082]" pageId="15" pageNumber="418">
Figs 49–54.
<taxonomicName id="4C0F04E9FFBBFFB5C089F85EFD94F808" authority="Hartung, 2019" authorityName="Hartung" authorityYear="2019" box="[255,567,1983,2002]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="418" phylum="Arthropoda" rank="subSpecies" species="handschini" status="subsp. nov." subSpecies="kinabaluensis">
<emphasis id="B97BA378FFBBFFB5C089F85EFD94F808" box="[255,567,1983,2002]" italics="true" pageId="15" pageNumber="418">Symphylax handschini kinabaluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2481E03FFBBFFB5C24AF85EFD3DF808" box="[572,670,1983,2002]" pageId="15" pageNumber="418" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
, holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum.
</paragraph>
</caption>
<paragraph id="8BB07F6AFFA4FFABC0C3FEDFFAB2FE4F" blockId="16.[151,760,190,373]" lastBlockId="17.[828,1437,190,917]" lastPageId="17" lastPageNumber="420" pageId="16" pageNumber="419">
<emphasis id="B97BA378FFA4FFAAC0C3FEDFFE83FE8F" box="[181,288,318,341]" italics="true" pageId="16" pageNumber="419">Structure.</emphasis>
Head (
<figureCitation id="133463EFFFA4FFAAC102FEDFFE65FE8F" box="[372,454,318,341]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="16" pageNumber="419">Fig. 51</figureCitation>
) of a length comparable to
<taxonomicName id="4C0F04E9FFA4FFAAC0E1FEBFFE36FEAF" box="[151,405,350,373]" class="Insecta" family="Colobathristidae" genus="Phaenacantha" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="419" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97BA378FFA4FFAAC0E1FEBFFE94FEAF" box="[151,311,350,373]" italics="true" pageId="16" pageNumber="419">Phaenacantha</emphasis>
species
</taxonomicName>
, with a median sulcus starting anteriad of ocelli and reaching middle of vertex. Two other depressions on the head, lateroposteriad of each ocellus, posterior margin being steep and anterior slope smooth, symmetrical on each side; other relief on head not pronounced. Eyes substylate. Angle between gula and clypeus almost right, gula subparallel to horizontal (
<figureCitation id="133463EFFFA4FFABC507FEBFFF16FC3A" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" lastPageId="17" lastPageNumber="420" pageId="16" pageNumber="419">Fig. 52</figureCitation>
). Labium reaching beyond middle coxae. Antennae thin, much longer than body, most segments (especially I and IV) slightly to moderately curved, left antennomere IV vestigial (
<figureCitation id="133463EFFFA5FFABC176FBC8FEECFB9A" box="[256,335,1065,1088]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="17" pageNumber="420">Fig. 49</figureCitation>
). Pronotum with collar similar to that in
<taxonomicName id="4C0F04E9FFA5FFABC0E1FBA8FE20FBBA" box="[151,387,1097,1120]" class="Insecta" family="Colobathristidae" genus="Phaenacantha" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="420" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97BA378FFA5FFABC0E1FBA8FE92FBBA" box="[151,305,1097,1120]" italics="true" pageId="17" pageNumber="420">Phaenacantha</emphasis>
species
</taxonomicName>
(
<figureCitation id="133463EFFFA5FFABC1E6FBA8FE7DFBBA" box="[400,478,1097,1120]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="17" pageNumber="420">Fig. 51</figureCitation>
). Middle lobe of pronotum with convex lateral outline, broader than collar; posterior lobe with almost straight lateral outline, same width as middle lobe. Middle and posterior lobe ca. in the same plane (
<figureCitation id="133463EFFFA5FFABC097FB28FE90FB3A" box="[225,307,1225,1248]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="17" pageNumber="420">Fig. 52</figureCitation>
), both lobes clearly convex in dorsal outline, separated by a deep transverse impression; posterior lobe only slightly elevated above the head. Hind third of posterior pronotal lobe gradually roundedly sloped posteriad; hind margin with weakly concave median part and two small rounded lateral lobes (
<figureCitation id="133463EFFFA5FFABC180FA88FDE5FA5A" box="[502,582,1385,1408]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="17" pageNumber="420">Fig. 51</figureCitation>
). Scutellar spine clearly longer than posterior pronotal lobe, slightly inclined anteriad (the angle to the plane of meso- and metanotum 95–100°), somewhat curved posteriad. Legs generally at least as long as or longer than body, hind legs the longest (
<figureCitation id="133463EFFFA5FFABC0E9F9E8FEBEF9FA" box="[159,285,1545,1568]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="17" pageNumber="420">Figs 49–50</figureCitation>
); femora slightly bent and widened distally, fore femora distally with a long straight spine and two small accessory spinules (
<figureCitation id="133463EFFFA5FFABC11CF9A8FE1AF9BA" box="[362,441,1609,1632]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="17" pageNumber="420">Fig. 53</figureCitation>
). Fore tibiae widened distally, with a median groove carrying terminally a comb of setae (
<figureCitation id="133463EFFFA5FFABC0E9F968FF4CF97A" box="[159,239,1673,1696]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="17" pageNumber="420">Fig. 54</figureCitation>
). Fore and hind wings well developed (specimen macropterous), fore wings ending very close to tip of abdomen. Abdomen slimmest at segments II and III and broadest at V, difference between broadest and thinnest part larger than in studied
<taxonomicName id="4C0F04E9FFA5FFABC103F8E8FDFDF8FA" box="[373,606,1801,1824]" class="Insecta" family="Colobathristidae" genus="Phaenacantha" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="420" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97BA378FFA5FFABC103F8E8FDAEF8FA" box="[373,525,1801,1824]" italics="true" pageId="17" pageNumber="420">Phaenacantha</emphasis>
species
</taxonomicName>
(
<figureCitation id="133463EFFFA5FFABC21CF8E8FD14F8FA" box="[618,695,1801,1824]" captionStart="Figs 49–54" captionStartId="15.[151,188,1983,2002]" captionTargetBox="[151,1436,510,1965]" captionTargetId="figure-13@15.[151,1436,510,1965]" captionTargetPageId="15" captionText="Figs 49–54. Symphylax handschini kinabaluensis subsp. nov., holotype. 49 – habitus, dorsal view. 50 – habitus, lateral view. 51 – head and thorax, dorsal view. 52 – head and thorax, lateral view; meso- and metapleura covered by rests of dried metathoracal gland secretion. 53 – right fore femur with the spine. 54 – right fore tibia with a comb of setae. Insert – specimen label. Abbreviations: col = pronotal collar, mp = middle lobe of the pronotum, pp = posterior lobe of the pronotum." figureDoi="http://doi.org/10.5281/zenodo.5339965" httpUri="https://zenodo.org/record/5339965/files/figure.png" pageId="17" pageNumber="420">Fig. 49</figureCitation>
). Borders between abdominal segments clear, segment borders smooth and not elevated. Lateral segment outlines straight or almost straight to slightly convex. Posterior border of segment VII in lateral outline almost straight (
<figureCitation id="133463EFFFA5FFABC2EFF868FD48F87A" box="[665,747,1929,1952]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="17" pageNumber="420">Fig. 56</figureCitation>
). Segments VIII and IX (pygophore) sunken into segment VII, sternite VIII with posterior median process sticking out (
<figureCitation id="133463EFFFA5FFABC0B3F808FE9CF7DA" box="[197,319,2025,2048]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="17" pageNumber="420">Figs 56–57</figureCitation>
). Pilosity on pygophore short, except setae on dorsal region of hypandrium (
<figureCitation id="133463EFFFA5FFABC19FF7E8FDC1F7FA" box="[489,610,2057,2080]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="17" pageNumber="420">Figs 58–61</figureCitation>
). Hypandrium building sharply ending parandria directed dorsoanteriad (
<figureCitation id="133463EFFFA5FFABC335FF3FFC51FF2F" box="[835,1010,222,245]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="17" pageNumber="420">Figs 58, 60–61</figureCitation>
); pygophore rounded posteriorly (
<figureCitation id="133463EFFFA5FFABC506FF3FFCF9FECF" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="17" pageNumber="420">Fig. 59</figureCitation>
); posterior margin of hypandrium in lateral view almost building an angle (
<figureCitation id="133463EFFFA5FFABC476FEFFFBECFEEF" box="[1024,1103,286,309]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="17" pageNumber="420">Fig. 61</figureCitation>
). Paramere (
<figureCitation id="133463EFFFA5FFABC4A4FEFFFA87FEEF" box="[1234,1316,286,309]" captionStart="Figs 55–61" captionStartId="16.[151,188,1983,2002]" captionTargetBox="[316,1271,397,1971]" captionTargetId="figure-13@16.[316,1271,397,1971]" captionTargetPageId="16" captionText="Figs 55–61. Symphylax handschini kinabaluensis subsp. nov., holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers." figureDoi="http://doi.org/10.5281/zenodo.5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="17" pageNumber="420">Figs 59</figureCitation>
,
<figureCitation id="133463EFFFA5FFABC558FEFFFAE9FEEF" box="[1326,1354,286,309]" captionStart="Fig" captionStartId="17.[150,181,868,887]" captionTargetBox="[176,755,212,852]" captionTargetId="figure-841@17.[151,762,189,852]" captionTargetPageId="17" captionText="Fig. 62. Symphylax handschini kinabaluensis subsp. nov., holotype, left paramere, inner view.Abbreviations:al = anterior lobe, pl = posterior lobe." figureDoi="http://doi.org/10.5281/zenodo.5339969" httpUri="https://zenodo.org/record/5339969/files/figure.png" pageId="17" pageNumber="420">62</figureCitation>
) spadelike, with broad dorsal margin with a concavity slightly posteriad from the middle, not forming processes, with anterior and posterior lobes well developed.
</paragraph>
<caption id="DF702FE2FFA4FFAAC0E1F85EFD44F7F8" ID-DOI="http://doi.org/10.5281/zenodo.5339967" ID-Zenodo-Dep="5339967" httpUri="https://zenodo.org/record/5339967/files/figure.png" pageId="16" pageNumber="419" startId="16.[151,188,1983,2002]" targetBox="[316,1271,397,1971]" targetPageId="16">
<paragraph id="8BB07F6AFFA4FFAAC0E1F85EFD44F7F8" blockId="16.[151,1437,1983,2082]" pageId="16" pageNumber="419">
Figs 55–61.
<taxonomicName id="4C0F04E9FFA4FFAAC174F85EFD9EF808" authority="Hartung, 2019" authorityName="Hartung" authorityYear="2019" box="[258,573,1983,2002]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="419" phylum="Arthropoda" rank="subSpecies" species="handschini" status="subsp. nov." subSpecies="kinabaluensis">
<emphasis id="B97BA378FFA4FFAAC174F85EFD9EF808" box="[258,573,1983,2002]" italics="true" pageId="16" pageNumber="419">Symphylax handschini kinabaluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2481E03FFA4FFAAC235F85EFD05F808" box="[579,678,1983,2002]" pageId="16" pageNumber="419" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
, holotype, genitalia. 55 – abdomen tip, dorsal view. 56 – abdomen tip, lateral view. 57 – abdomen tip, caudal view. 58 – pygophore with genitalia, dorsal view. 59 – pygophore with genitalia, lateral view. 60 – pygophore with genitalia, caudal view. 61 – hypandrium, lateral view. Abbreviations: hyp = hypandrium, p = paramere, par = parandrium, pp = posterior process of sternite VIII, pyg = pygophore. Abdominal segments are indicated with Roman numbers.
</paragraph>
</caption>
<paragraph id="8BB07F6AFFA5FFABC32CFE7FFBB6FCEF" blockId="17.[828,1437,190,917]" pageId="17" pageNumber="420">
<emphasis id="B97BA378FFA5FFABC32CFE7FFBA1FE6F" bold="true" box="[858,1026,414,437]" pageId="17" pageNumber="420">
<emphasis id="B97BA378FFA5FFABC32CFE7FFC58FE6F" bold="true" box="[858,1019,414,437]" italics="true" pageId="17" pageNumber="420">Measurements</emphasis>
.
</emphasis>
Body length
<quantity id="4CF7D28FFFA5FFABC4E0FE7FFB59FE6F" box="[1174,1274,414,437]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="17" pageNumber="420" unit="mm" value="6.8">6.80 mm</quantity>
; head width (= max. body width)
<quantity id="4CF7D28FFFA5FFABC471FE5FFBCFFE0F" box="[1031,1132,446,469]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="17" pageNumber="420" unit="mm" value="1.31">1.31 mm</quantity>
. Total antenna length
<quantity id="4CF7D28FFFA5FFABC516FE5FFCC6FE2F" metricMagnitude="-2" metricUnit="m" metricValue="1.047" pageId="17" pageNumber="420" unit="mm" value="10.47">10.47 mm</quantity>
(average segment ratios, I to IV: 1.00-1.49-1.52-1.98). Ratio antenna: body 1.54. Labium length
<quantity id="4CF7D28FFFA5FFABC577FE1FFAC7FDCF" box="[1281,1380,510,533]" metricMagnitude="-3" metricUnit="m" metricValue="2.01" pageId="17" pageNumber="420" unit="mm" value="2.01">2.01 mm</quantity>
. Distance between ocelli
<quantity id="4CF7D28FFFA5FFABC454FDFFFB26FDEF" box="[1058,1157,542,565]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="420" unit="mm" value="0.12">0.12 mm</quantity>
, between ocellus and eye
<quantity id="4CF7D28FFFA5FFABC34AFDDFFC03FD8F" box="[828,928,574,597]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="420" unit="mm" value="0.22">0.22 mm</quantity>
, ratio 1.83. Length of collar + middle lobe of pronotum
<quantity id="4CF7D28FFFA5FFABC3DFFDBFFBA9FDAF" box="[937,1034,606,629]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="17" pageNumber="420" unit="mm" value="0.47">0.47 mm</quantity>
; length of posterior pronotal lobe
<quantity id="4CF7D28FFFA5FFABC518FDBFFCCBFD4F" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="17" pageNumber="420" unit="mm" value="0.82">0.82 mm</quantity>
, ratio of the two measurements 1.74. Scutellar spine tip broken, therefore the length not measurable. Measurements of the hind tarsus (segments I / II / III, respectively): 0.93 / 0.17 /
<quantity id="4CF7D28FFFA5FFABC3FBFD3FFC4DFD2F" box="[909,1006,734,757]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="420" unit="mm" value="0.25">0.25 mm</quantity>
, total length of the hind tarsus
<quantity id="4CF7D28FFFA5FFABC541FD3FFA3AFD2F" box="[1335,1433,734,757]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="17" pageNumber="420" unit="mm" value="1.35">1.35 mm</quantity>
. Maximum width of abdomen
<quantity id="4CF7D28FFFA5FFABC4F6FD1FFB40FCCF" box="[1152,1251,766,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="420" unit="mm" value="1.23">1.23 mm</quantity>
, minimum width
<quantity id="4CF7D28FFFA5FFABC34AFCFFFC3EFCEF" box="[828,925,798,821]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="17" pageNumber="420" unit="mm" value="0.43">0.43 mm</quantity>
, ratio 2.86.
</paragraph>
</subSubSection>
<subSubSection id="C3152CE1FFA5FFABC34AFCDFFA3EFCAF" pageId="17" pageNumber="420" type="etymology">
<paragraph id="8BB07F6AFFA5FFABC34AFCDFFA3EFCAF" blockId="17.[828,1437,190,917]" pageId="17" pageNumber="420">
<emphasis id="B97BA378FFA5FFABC34AFCDFFC1AFC8F" bold="true" box="[828,953,830,853]" pageId="17" pageNumber="420">Etymology.</emphasis>
The name of this subspecies refers to Kinabalu National Park where this specimen was collected; adjective.
</paragraph>
</subSubSection>
<caption id="DF702FE2FFA5FFABC0E0FC85FD5AFC48" ID-DOI="http://doi.org/10.5281/zenodo.5339969" ID-Zenodo-Dep="5339969" httpUri="https://zenodo.org/record/5339969/files/figure.png" pageId="17" pageNumber="420" startId="17.[150,181,868,887]" targetBox="[176,755,212,852]" targetPageId="17">
<paragraph id="8BB07F6AFFA5FFABC0E0FC85FD5AFC48" blockId="17.[150,761,868,914]" pageId="17" pageNumber="420">
Fig. 62.
<taxonomicName id="4C0F04E9FFA5FFABC0A8FC85FDB4FCAD" authority="Hartung, 2019" authorityName="Hartung" authorityYear="2019" box="[222,535,868,887]" class="Insecta" family="Colobathristidae" genus="Symphylax" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="420" phylum="Arthropoda" rank="subSpecies" species="handschini" status="subsp. nov." subSpecies="kinabaluensis">
<emphasis id="B97BA378FFA5FFABC0A8FC85FDB4FCAD" box="[222,535,868,887]" italics="true" pageId="17" pageNumber="420">Symphylax handschini kinabaluensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2481E03FFA5FFABC26BFC85FDDCFCAD" box="[541,639,868,887]" pageId="17" pageNumber="420" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
, holotype, left paramere, inner view.Abbreviations:al = anterior lobe, pl = posterior lobe.
</paragraph>
</caption>
<subSubSection id="C3152CE1FFA5FFABC34AFC9FFB34FC4F" box="[828,1175,894,917]" pageId="17" pageNumber="420" type="distribution">
<paragraph id="8BB07F6AFFA5FFABC34AFC9FFB34FC4F" blockId="17.[828,1437,190,917]" box="[828,1175,894,917]" pageId="17" pageNumber="420">
<emphasis id="B97BA378FFA5FFABC34AFC9FFC6DFC4F" bold="true" box="[828,974,894,917]" pageId="17" pageNumber="420">Distribution.</emphasis>
<collectingCountry id="F3183FFAFFA5FFABC3A2FC9FFB95FC4F" box="[980,1078,894,917]" name="Malaysia" pageId="17" pageNumber="420">Malaysia</collectingCountry>
(
<collectingRegion id="49CBB188FFA5FFABC432FC9FFB28FC4F" box="[1092,1163,894,917]" country="Malaysia" name="Sabah" pageId="17" pageNumber="420">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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