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<document id="CED0AC69BD1258A11CF94006F6014260" ID-CLB-Dataset="29087" ID-DOI="10.11646/zootaxa.4455.3.4" ID-GBIF-Dataset="c9bf5059-abe4-414e-a7c2-f3f9e75f2979" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457467" ID-ZooBank="69209FF7-80B0-409D-A257-0B88C65E903F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539262799757" checkinUser="plazi" docAuthor="Kamiński, Marcin J. &amp; Schoeman, Colin S." docDate="2018" docId="038BC66EFFDC965EF7ABF916FDE4FE7A" docLanguage="en" docName="zootaxa.4455.3.4.pdf" docOrigin="Zootaxa 4455 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anaxius obesus Fahraeus 1870" docType="treatment" docVersion="7" lastPageNumber="481" masterDocId="FFB2BE16FFD49654F73CFFF7FF95FFF2" masterDocTitle="Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina)" masterLastPageNumber="485" masterPageNumber="471" pageNumber="479" updateTime="1698576062251" updateUser="ExternalLinkService">
<mods:mods id="08561C7795E47E773B54E952EB52FB65" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="54F3C9C093A2AB4F8D791FB929EDD77C">
<mods:title id="4894278A1C5C3897B74059B954ED72AC">Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina)</mods:title>
</mods:titleInfo>
<mods:name id="3CC62B3CFDA81824201701AC130E6BED" type="personal">
<mods:role id="0903DD1C1FF3C4622E51917C649DB750">
<mods:roleTerm id="F232506DA5C0672AECAE8036D7A532FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B3E57900BC941ADF8ADBE5144F57E8D">Kamiński, Marcin J.</mods:namePart>
</mods:name>
<mods:name id="53F17B0DC43C7C261F6593A1B9D4B4C0" type="personal">
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</mods:role>
<mods:namePart id="B5BDAB04BBE1CB2425483D1BA09BADBE">Schoeman, Colin S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D1D030B5228C9643279E19E1AF478B89">text</mods:typeOfResource>
<mods:relatedItem id="301C2640A398E269D95D8CEA744A3A04" type="host">
<mods:titleInfo id="8166892CA06BCA209F06AF58DB82C544">
<mods:title id="46B40A21FF7825A9F7249F9036FC5B67">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="221FF0E5569D17FCD20CD6EA7EEF7F8C">
<mods:date id="569BFE6D30E2BF093A7FDD63D955D094">2018</mods:date>
<mods:detail id="CC69826D3663C07C158144CCA17A3E20" type="pubDate">
<mods:number id="3B1664015A4EA06D7EEEC348191EBE3F">2018-08-03</mods:number>
</mods:detail>
<mods:detail id="8D386F51F83A4BAFAF94BB02C60096BA" type="volume">
<mods:number id="2D48D45AF75403B068E17B7A3FF314FA">4455</mods:number>
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<mods:detail id="AE5165339F0AE62399640ED5DCCD1FEA" type="issue">
<mods:number id="2E52CC7CC6499ED74CC073F638D3675A">3</mods:number>
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<mods:classification id="F77A330E9DB26F38C5E66E717C8F91FD">journal article</mods:classification>
<mods:identifier id="25E7CA6617F2EB583AC578CEDD293310" type="CLB-Dataset">29087</mods:identifier>
<mods:identifier id="1B73C2DAE10F9F8957005BDA49AF184C" type="DOI">10.11646/zootaxa.4455.3.4</mods:identifier>
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<mods:identifier id="736E32B72A6E627A27AA0CB0C841C472" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AB1C6F9FE5BFF6C6D9929F3609F64A0A" type="Zenodo-Dep">1457467</mods:identifier>
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<treatment id="038BC66EFFDC965EF7ABF916FDE4FE7A" ID-DOI="http://doi.org/10.5281/zenodo.5975961" ID-GBIF-Taxon="148691417" ID-Zenodo-Dep="5975961" LSID="urn:lsid:plazi:treatment:038BC66EFFDC965EF7ABF916FDE4FE7A" httpUri="http://treatment.plazi.org/id/038BC66EFFDC965EF7ABF916FDE4FE7A" lastPageId="10" lastPageNumber="481" pageId="8" pageNumber="479">
<subSubSection id="C33824F3FFDC965CF7ABF916FEB6F8EF" pageId="8" pageNumber="479" type="nomenclature">
<paragraph id="8B9D7778FFDC965CF7ABF916FD8EF909" blockId="8.[151,539,1761,1821]" box="[151,539,1761,1787]" pageId="8" pageNumber="479">
<heading id="D0D5C014FFDC965CF7ABF916FD8EF909" bold="true" box="[151,539,1761,1787]" fontSize="11" level="1" pageId="8" pageNumber="479" reason="1">
<taxonomicName id="4C220CFBFFDC965CF7ABF916FD8EF909" authority="Fahraeus, 1870" authorityName="Fahraeus" authorityYear="1870" box="[151,539,1761,1787]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956AB6AFFDC965CF7ABF916FD8EF909" bold="true" box="[151,539,1761,1787]" pageId="8" pageNumber="479">
<emphasis id="B956AB6AFFDC965CF7ABF916FEC1F909" bold="true" box="[151,340,1761,1787]" italics="true" pageId="8" pageNumber="479">Anaxius obesus</emphasis>
<bibRefCitation id="EFB30A89FFDC965CF660F916FD8EF909" box="[348,539,1761,1787]" pageId="8" pageNumber="479" refId="ref8622" refString="Fahraeus, O. I. (1870) Coleoptera Caffrariae a J. A. Walhberg collecta. - Ofversigt af Kongl. Akademiens Ftirhandlingar 4: 243 - 358." type="journal article">Fåhraeus, 1870</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9D7778FFDC965CF7ABF8F3FEB6F8EF" blockId="8.[151,539,1761,1821]" box="[151,291,1796,1821]" pageId="8" pageNumber="479">
(
<figureCitation id="13196BFDFFDC965CF7A3F8F3FF6AF8EE" box="[159,255,1796,1821]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="8" pageNumber="479">Figs. 1E</figureCitation>
,
<figureCitation id="13196BFDFFDC965CF630F8F3FE8FF8EF" box="[268,282,1796,1821]" captionStart="FIGURE 5" captionStartId="11.[151,250,1731,1753]" captionTargetBox="[161,1437,844,1704]" captionTargetId="figure@11.[151,1437,844,1710]" captionTargetPageId="11" captionText="FIGURE 5. Distribution of the species representing Anaxius." httpUri="https://zenodo.org/record/1457477/files/figure.png" pageId="8" pageNumber="479">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33824F3FFDC965CF7ABF8B0FD09F817" pageId="8" pageNumber="479" type="reference_group">
<paragraph id="8B9D7778FFDC965CF7ABF8B0FD59F8AE" blockId="8.[151,716,1862,1884]" box="[151,716,1862,1884]" pageId="8" pageNumber="479">
<treatmentCitationGroup id="AB325056FFDC965CF7ABF8B0FD59F8AE" box="[151,716,1862,1884]" pageId="8" pageNumber="479">
<taxonomicName id="4C220CFBFFDC965CF7ABF8B0FD99F8AE" authority="Fahraeus, 1870: 307" authorityName="Fahraeus" authorityPageNumber="307" authorityYear="1870" box="[151,524,1862,1884]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956AB6AFFDC965CF7ABF8B0FEA1F8AE" box="[151,308,1862,1884]" italics="true" pageId="8" pageNumber="479">Anaxius obesus</emphasis>
<treatmentCitation id="0A835169FFDC965CF606F8B1FD99F8AE" author="" box="[314,524,1862,1884]" page="307" pageId="8" pageNumber="479">
<bibRefCitation id="EFB30A89FFDC965CF606F8B1FE4DF8AE" box="[314,472,1862,1884]" pageId="8" pageNumber="479" refId="ref8622" refString="Fahraeus, O. I. (1870) Coleoptera Caffrariae a J. A. Walhberg collecta. - Ofversigt af Kongl. Akademiens Ftirhandlingar 4: 243 - 358." type="journal article">Fåhraeus, 1870</bibRefCitation>
: 307
</treatmentCitation>
</taxonomicName>
.-
<treatmentCitation id="0A835169FFDC965CF521F8B1FD5DF8AE" author="" box="[541,712,1862,1884]" page="211" pageId="8" pageNumber="479">
<bibRefCitation id="EFB30A89FFDC965CF521F8B1FD03F8AE" box="[541,662,1862,1884]" pageId="8" pageNumber="479" refId="ref9041" refString="Koch, C. (1958) Tenebrionidae of Angola. Publicacoes Culturais da Companhia de Diamantes de Angola, 39, 1 - 231." type="journal article">Koch, 1958</bibRefCitation>
: 211
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9D7778FFDC965CF7ABF873FD09F817" blockId="8.[151,1436,1924,2021]" pageId="8" pageNumber="479">
<emphasis id="B956AB6AFFDC965CF7ABF873FF74F869" bold="true" box="[151,225,1924,1949]" pageId="8" pageNumber="479">Notes.</emphasis>
<bibRefCitation id="EFB30A89FFDC965CF7D4F873FE0AF86E" box="[232,415,1924,1949]" pageId="8" pageNumber="479" refId="ref8622" refString="Fahraeus, O. I. (1870) Coleoptera Caffrariae a J. A. Walhberg collecta. - Ofversigt af Kongl. Akademiens Ftirhandlingar 4: 243 - 358." type="journal article">Fåhraeus (1870)</bibRefCitation>
did not specify the number of studied specimens, however from the context it seems to be clear that he was dealing with at least two specimens. Therefore, a
<typeStatus id="5499C9DAFFDC965CF484F85FFB8AF833" box="[952,1055,1960,1985]" pageId="8" pageNumber="479" type="lectotype">lectotype</typeStatus>
designation is needed to fix the taxonomic status of the genus and the species.
</paragraph>
</subSubSection>
<caption id="DF5D27F0FFDD965DF7ABF84FFCF4F83C" box="[151,865,1976,1998]" httpUri="https://zenodo.org/record/1457475/files/figure.png" pageId="9" pageNumber="480" startId="9.[151,250,1976,1998]" targetBox="[273,1318,193,1958]" targetPageId="9">
<paragraph id="8B9D7778FFDD965DF7ABF84FFCF4F83C" blockId="9.[151,865,1976,1998]" box="[151,865,1976,1998]" pageId="9" pageNumber="480">
<emphasis id="B956AB6AFFDD965DF7ABF84FFE98F83F" bold="true" box="[151,269,1976,1998]" pageId="9" pageNumber="480">FIGURE 4</emphasis>
. Morphology of female terminalia of
<taxonomicName id="4C220CFBFFDD965DF5B1F84EFCC9F83C" authorityName="Koch" authorityYear="1958" box="[653,860,1976,1998]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="480" phylum="Arthropoda" rank="species" species="campbellae">Anaxius campbellae</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C33824F3FFDE965EF7FBFF60FC25FEB2" pageId="10" pageNumber="481" type="materials_examined">
<paragraph id="8B9D7778FFDE965EF7FBFF60FC25FEB2" blockId="10.[151,1436,151,392]" pageId="10" pageNumber="481">
<emphasis id="B956AB6AFFDE965EF7FBFF60FD81FF42" bold="true" box="[199,532,151,176]" pageId="10" pageNumber="481">Material studied. Lectotype</emphasis>
, designated here, male (TMSA): “Caffra- / ria”.
<materialsCitation id="3B4A7D25FFDE965EF37DFF60FED3FEB2" ID-GBIF-Occurrence="1917075923" collectingDate="1950-11" collectingDateMax="2009-12-12" collectingDateMin="1950-11" collectionCode="TMSA" collectorName="Other &amp; Lichtenburg &amp; Transvaal &amp; Dr. Brauns &amp; Plansberg &amp; P. L. Breutz &amp; S. Afr. &amp; Ruth Muller" county="Mabote" latitude="-25.17" location="Transvaal" longLatPrecision="748" longitude="28.17" municipality="Waterberg" pageId="10" pageNumber="481" specimenCount="2" specimenCount-male="2" stateProvince="Limpopo" typeStatus="paralectotype">
<typeStatus id="5499C9DAFFDE965EF37DFF60FB73FF42" box="[1089,1254,151,176]" pageId="10" pageNumber="481" type="paralectotype">
<emphasis id="B956AB6AFFDE965EF37DFF60FB73FF42" bold="true" box="[1089,1254,151,176]" pageId="10" pageNumber="481">Paralectotype</emphasis>
</typeStatus>
, male (
<collectionCode id="ED33EFBDFFDE965EF200FF6FFA04FF42" box="[1340,1425,152,176]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="10" pageNumber="481">TMSA</collectionCode>
): same data as holotype.
<emphasis id="B956AB6AFFDE965EF69BFF4BFDCBFF27" bold="true" box="[423,606,188,213]" pageId="10" pageNumber="481">
<collectorName id="26D712AEFFDE965EF69BFF4BFE7BFF27" box="[423,494,188,213]" pageId="10" pageNumber="481">Other</collectorName>
material
</emphasis>
, male (
<collectionCode id="ED33EFBDFFDE965EF58BFF4BFC99FF26" box="[695,780,188,212]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="10" pageNumber="481">TMSA</collectionCode>
): &quot;
<collectorName id="26D712AEFFDE965EF416FF4BFC23FF27" box="[810,950,188,213]" pageId="10" pageNumber="481">Lichtenburg</collectorName>
/
<collectorName id="26D712AEFFDE965EF4EDFF4BFBABFF27" box="[977,1086,188,213]" pageId="10" pageNumber="481">Transvaal</collectorName>
/
<collectorName id="26D712AEFFDE965EF366FF4AFB49FF26" box="[1114,1244,188,212]" pageId="10" pageNumber="481">Dr. Brauns</collectorName>
&quot;, male (
<collectionCode id="ED33EFBDFFDE965EF207FF4BFA05FF26" box="[1339,1424,188,212]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="10" pageNumber="481">TMSA</collectionCode>
): &quot;
<collectorName id="26D712AEFFDE965EF7A3FF28FE85FF0A" box="[159,272,223,248]" pageId="10" pageNumber="481">Plansberg</collectorName>
/
<collectingDate id="EFD8A850FFDE965EF614FF28FE10FF0A" box="[296,389,223,248]" pageId="10" pageNumber="481" value="1950-11">
<date id="FF9C51B8FFDE965EF614FF28FE10FF0A" box="[296,389,223,248]" pageId="10" pageNumber="481" value="1950-11">XI.1950</date>
</collectingDate>
/
<collectorName id="26D712AEFFDE965EF6A0FF17FDB5FF0A" box="[412,544,224,248]" pageId="10" pageNumber="481">P.L. Breutz</collectorName>
&quot;;
<specimenCount id="9D24BCF1FFDE965EF509FF17FD19FF0A" box="[565,652,223,248]" pageId="10" pageNumber="481" type="male">2 males</specimenCount>
(
<collectionCode id="ED33EFBDFFDE965EF5A1FF17FD67FF0A" box="[669,754,224,248]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="10" pageNumber="481">TMSA</collectionCode>
): &quot;
<collectorName id="26D712AEFFDE965EF42CFF17FCCAFF0A" box="[784,863,223,248]" pageId="10" pageNumber="481">S. Afr.</collectorName>
;
<collectingRegion id="49E6B99AFFDE965EF457FF28FC47FF0A" box="[875,978,223,248]" country="South Africa" name="Limpopo" pageId="10" pageNumber="481">Limpopo</collectingRegion>
Prov. /
<collectingCounty id="62FC0FF4FFDE965EF317FF28FB15FF0A" box="[1067,1152,223,248]" pageId="10" pageNumber="481">Mabote</collectingCounty>
farm /
<geoCoordinate id="EE1611BFFFDE965EF3E9FF17FABFFF0A" box="[1237,1322,224,248]" direction="south" orientation="latitude" pageId="10" pageNumber="481" precision="555" value="-24.07">24.07 S</geoCoordinate>
–
<geoCoordinate id="EE1611BFFFDE965EF205FF17FA1AFF0A" box="[1337,1423,224,248]" direction="east" orientation="longitude" pageId="10" pageNumber="481" precision="555" value="28.39">28.39E</geoCoordinate>
&quot;, &quot;
<collectingDate id="EFD8A850FFDE965EF7A3FEF3FEB5FEEE" box="[159,288,260,284]" pageId="10" pageNumber="481" value="2009-12-12">
<date id="FF9C51B8FFDE965EF7A3FEF3FEB5FEEE" box="[159,288,260,284]" pageId="10" pageNumber="481" value="2009-12-12">12.12.2009</date>
</collectingDate>
/ in the
<collectingMunicipality id="6BF9ED02FFDE965EF6B7FEF3FE6AFEEF" box="[395,511,260,285]" pageId="10" pageNumber="481">Waterberg</collectingMunicipality>
/ leg.
<collectorName id="26D712AEFFDE965EF573FEF3FD77FEEF" box="[591,738,260,285]" pageId="10" pageNumber="481">Ruth Müller</collectorName>
&quot;; male (
<collectionCode id="ED33EFBDFFDE965EF478FEF3FC0DFEEE" box="[836,920,260,284]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="10" pageNumber="481">TMSA</collectionCode>
): “
<collectorName id="26D712AEFFDE965EF480FEF3FB98FEEF" box="[956,1037,260,285]" pageId="10" pageNumber="481">S. Afr.</collectorName>
:
<location id="8EFD21A3FFDE965EF327FEF3FB1DFEEF" LSID="urn:lsid:plazi:treatment:038BC66EFFDC965EF7ABF916FDE4FE7A:8EFD21A3FFDE965EF327FEF3FB1DFEEF" box="[1051,1160,260,285]" county="Mabote" latitude="-25.17" longLatPrecision="748" longitude="28.17" municipality="Waterberg" name="Transvaal" pageId="10" pageNumber="481" stateProvince="Limpopo">Transvaal</location>
/
<location id="8EFD21A3FFDE965EF399FEF3FAD8FEEF" LSID="urn:lsid:plazi:treatment:038BC66EFFDC965EF7ABF916FDE4FE7A:8EFD21A3FFDE965EF399FEF3FAD8FEEF" box="[1189,1357,260,285]" county="Mabote" latitude="-25.17" longLatPrecision="748" longitude="28.17" municipality="Waterberg" name="Pienaars River" pageId="10" pageNumber="481" stateProvince="Limpopo">Pienaars River</location>
<quantity id="4CDADA9DFFDE965EF264FEF3FA1CFEEF" box="[1368,1417,260,285]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="481" unit="km" value="8.0">8km</quantity>
/
<geoCoordinate id="EE1611BFFFDE965EF7ABFED0FF70FEB2" box="[151,229,295,320]" direction="south" orientation="latitude" pageId="10" pageNumber="481" precision="555" value="-25.17">25.17S</geoCoordinate>
–
<geoCoordinate id="EE1611BFFFDE965EF7C8FED0FED3FEB2" box="[244,326,295,320]" direction="east" orientation="longitude" pageId="10" pageNumber="481" precision="555" value="28.17">28.17E</geoCoordinate>
</materialsCitation>
,
<date id="FF9C51B8FFDE965EF66DFEDFFE46FEB2" box="[337,467,296,320]" pageId="10" pageNumber="481" value="1994-12-28">28.12.1994</date>
; E-Y 3234, on ground / leg. CL Bellamy”.
</paragraph>
</subSubSection>
<subSubSection id="C33824F3FFDE965EF7FBFEBBFDE4FE7A" pageId="10" pageNumber="481" type="distribution">
<paragraph id="8B9D7778FFDE965EF7FBFEBBFDE4FE7A" blockId="10.[151,1436,151,392]" pageId="10" pageNumber="481">
<emphasis id="B956AB6AFFDE965EF7FBFEBBFEF5FE97" bold="true" box="[199,352,332,357]" pageId="10" pageNumber="481">Distribution.</emphasis>
This species has been collected in the following ecoregions of
<collectingCountry id="F33537E8FFDE965EF308FEBBFB50FE97" box="[1076,1221,332,357]" name="South Africa" pageId="10" pageNumber="481">
<collectingRegion id="49E6B99AFFDE965EF308FEBBFBE3FE97" box="[1076,1142,332,357]" country="Tanzania" name="Kusini Unguja" pageId="10" pageNumber="481">South</collectingRegion>
Africa
</collectingCountry>
(
<figureCitation id="13196BFDFFDE965EF3EBFEBBFA89FE97" box="[1239,1308,332,357]" captionStart="FIGURE 5" captionStartId="11.[151,250,1731,1753]" captionTargetBox="[161,1437,844,1704]" captionTargetId="figure@11.[151,1437,844,1710]" captionTargetPageId="11" captionText="FIGURE 5. Distribution of the species representing Anaxius." httpUri="https://zenodo.org/record/1457477/files/figure.png" pageId="10" pageNumber="481">Fig. 5</figureCitation>
): Highveld grasslands; and Southern Africa bushveld.
</paragraph>
</subSubSection>
</treatment>
</document>
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