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<document id="70362B6609A749D15D34838CF9869615" ID-CLB-Dataset="29087" ID-DOI="10.11646/zootaxa.4455.3.4" ID-GBIF-Dataset="c9bf5059-abe4-414e-a7c2-f3f9e75f2979" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457467" ID-ZooBank="69209FF7-80B0-409D-A257-0B88C65E903F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539262799757" checkinUser="plazi" docAuthor="Kamiński, Marcin J. &amp; Schoeman, Colin S." docDate="2018" docId="038BC66EFFD3965CF7ABF9D2FEC7F96A" docLanguage="en" docName="zootaxa.4455.3.4.pdf" docOrigin="Zootaxa 4455 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anaxius meletsensis Kamiński &amp; Schoeman 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="479" masterDocId="FFB2BE16FFD49654F73CFFF7FF95FFF2" masterDocTitle="Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina)" masterLastPageNumber="485" masterPageNumber="471" pageNumber="478" updateTime="1698576062251" updateUser="ExternalLinkService">
<mods:mods id="770B3CCE54D88C1DB6B6730D6BB26281" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0268C3FF5B307EC73CE9C2F1359678AE">
<mods:title id="A695BC8C66171780F3AA5B2120688FF1">Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina)</mods:title>
</mods:titleInfo>
<mods:name id="DC60F4B2F6C520F1B69E105D2D0319F3" type="personal">
<mods:role id="921AD286121B1171FDBB34F7ACA0DF4A">
<mods:roleTerm id="EB181423C6A1D02162CC6AE8E475858E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC617539C4775EB3A726A5E2958AE64A">Kamiński, Marcin J.</mods:namePart>
</mods:name>
<mods:name id="40774E9E5C243244D82BD9FFD377F64F" type="personal">
<mods:role id="CF5C76C71E8921FEB852EA238DC89D3F">
<mods:roleTerm id="A349E446A5643678C1D007561134456D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75B7D5B3C13250FDEF21D8838EFA1C4B">Schoeman, Colin S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4BAA8CA1664F9632F4FA70786A03F414">text</mods:typeOfResource>
<mods:relatedItem id="BB63FB8038A8C3CFC07A4E17A2987770" type="host">
<mods:titleInfo id="D0E7209082651D59DB9660FE3AD3E82C">
<mods:title id="0A092A41B755680DD10E532DD71EF25B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8DD0EBAB64038275C87C26692F5C356B">
<mods:date id="967C99FB8CCB80545A418A561E9304FF">2018</mods:date>
<mods:detail id="007E45F70E42D7D90A562E9F40338BAD" type="pubDate">
<mods:number id="141806F985D31952F057BA93135BE9E2">2018-08-03</mods:number>
</mods:detail>
<mods:detail id="EA90570DA66CDECED244020F8DA52388" type="volume">
<mods:number id="4317B200A2871D595ECE85ADE067AF3D">4455</mods:number>
</mods:detail>
<mods:detail id="36DB239D8202C5CE432F4A76F470422C" type="issue">
<mods:number id="FEDD1361CF83AF708FE69BAAC9B003DA">3</mods:number>
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<mods:start id="B130C1F98F2EB6F3B46DFC6C120B38B4">471</mods:start>
<mods:end id="9FBBF538E6D0F9FCC56CD0BFE83EA989">485</mods:end>
</mods:extent>
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<mods:classification id="4F714F0FC013C4AC20AC18703B513061">journal article</mods:classification>
<mods:identifier id="6384343C8E84C404FD4B8A8E44DB5814" type="CLB-Dataset">29087</mods:identifier>
<mods:identifier id="46B91813ED4DBA769397E1B0DDD7353A" type="DOI">10.11646/zootaxa.4455.3.4</mods:identifier>
<mods:identifier id="D9775671C2B7E14B36D3A737E83C93A1" type="GBIF-Dataset">c9bf5059-abe4-414e-a7c2-f3f9e75f2979</mods:identifier>
<mods:identifier id="FBFA4F4CD609DE63DD2C1DAD9440EF4C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0C912E72CAB75EBA82657D0169E19E90" type="Zenodo-Dep">1457467</mods:identifier>
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</mods:mods>
<treatment id="038BC66EFFD3965CF7ABF9D2FEC7F96A" ID-DOI="http://doi.org/10.5281/zenodo.5975959" ID-GBIF-Taxon="148691420" ID-Zenodo-Dep="5975959" LSID="urn:lsid:plazi:treatment:038BC66EFFD3965CF7ABF9D2FEC7F96A" httpUri="http://treatment.plazi.org/id/038BC66EFFD3965CF7ABF9D2FEC7F96A" lastPageId="8" lastPageNumber="479" pageId="7" pageNumber="478">
<subSubSection id="C33824F3FFD39653F7ABF9D2FB0FF993" pageId="7" pageNumber="478" type="nomenclature">
<paragraph id="8B9D7778FFD39653F7ABF9D2FE65F9B2" blockId="7.[151,1178,1573,1669]" box="[151,496,1573,1600]" pageId="7" pageNumber="478">
<heading id="D0D5C014FFD39653F7ABF9D2FE65F9B2" bold="true" box="[151,496,1573,1600]" fontSize="11" level="1" pageId="7" pageNumber="478" reason="1">
<emphasis id="B956AB6AFFD39653F7ABF9D2FE65F9B2" bold="true" box="[151,496,1573,1600]" pageId="7" pageNumber="478">
<taxonomicName id="4C220CFBFFD39653F7ABF9D2FE1EF9CD" authority="Kamiński &amp; Schoeman, 2018" authorityName="Kamiński &amp; Schoeman" authorityYear="2018" box="[151,395,1573,1599]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="478" phylum="Arthropoda" rank="species" species="meletsensis" status="sp. nov.">
<emphasis id="B956AB6AFFD39653F7ABF9D2FE1EF9CD" bold="true" box="[151,395,1573,1599]" italics="true" pageId="7" pageNumber="478">Anaxius meletsensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2651611FFD39653F6AFF9D1FE65F9B2" box="[403,496,1574,1600]" pageId="7" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D7778FFD39653F7ABF9BFFB0FF993" blockId="7.[151,1178,1573,1669]" box="[151,1178,1608,1633]" pageId="7" pageNumber="478">http://zoobank.org/urn:lsid:zoobank.org:act:79A5A46A-8614-410A-A322-AC3CF9933242</paragraph>
</subSubSection>
<subSubSection id="C33824F3FFD39653F7ABF99BFE0FF977" box="[151,410,1644,1669]" pageId="7" pageNumber="478" type="description">
<paragraph id="8B9D7778FFD39653F7ABF99BFE0FF977" blockId="7.[151,1178,1573,1669]" box="[151,410,1644,1669]" pageId="7" pageNumber="478">
(
<figureCitation id="13196BFDFFD39653F7A3F99BFE97F977" box="[159,258,1644,1669]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="7" pageNumber="478">Figs. 1D</figureCitation>
,
<figureCitation id="13196BFDFFD39653F632F99AFED2F976" box="[270,327,1645,1669]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[151,1435,681,1847]" captionTargetId="figure@5.[151,1435,681,1847]" captionTargetPageId="5" captionText="FIGURE 2. Diagnostic characters proposed for Anaxius: mentum (A), proleg (B, G), mesoleg (C, F), metaleg (D), metaventrite (H, I). Anaxius bloubergensis (E, F), A. campbellae (A, H), A. meletsensis (G, I), and A. prozeskyi (B, C, D). Abbreviations: d—denticles, ld—surface between labial palpi, lk—longitudinal keel, lw—lateral wing, p—palpifer, t—apical denticle, tub—tubercles." httpUri="https://zenodo.org/record/1457471/files/figure.png" pageId="7" pageNumber="478">2G, I</figureCitation>
,
<figureCitation id="13196BFDFFD39653F66FF99AFEE2F977" box="[339,375,1645,1669]" captionStart="FIGURE 3" captionStartId="7.[151,250,1440,1462]" captionTargetBox="[200,1386,725,1434]" captionTargetId="figure@7.[200,1386,725,1434]" captionTargetPageId="7" captionText="FIGURE 3. Aedeagal tegmen morphology. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457473/files/figure.png" pageId="7" pageNumber="478">3D</figureCitation>
,
<figureCitation id="13196BFDFFD39653F6BFF99BFE04F977" box="[387,401,1644,1669]" captionStart="FIGURE 5" captionStartId="11.[151,250,1731,1753]" captionTargetBox="[161,1437,844,1704]" captionTargetId="figure@11.[151,1437,844,1710]" captionTargetPageId="11" captionText="FIGURE 5. Distribution of the species representing Anaxius." httpUri="https://zenodo.org/record/1457477/files/figure.png" pageId="7" pageNumber="478">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33824F3FFD39653F7ABF943FB8FF8CB" pageId="7" pageNumber="478" type="materials_examined">
<paragraph id="8B9D7778FFD39653F7ABF943FB8FF8CB" blockId="7.[151,1436,1716,2029]" pageId="7" pageNumber="478">
<materialsCitation id="3B4A7D25FFD39653F7ABF943FC1EF903" ID-GBIF-Occurrence="1917075887" collectingDate="2014-11-27" collectionCode="TMSA" collectorName="S. Afr. &amp; Ruth Muller" elevation="1003" latitude="-24.36" location="Meletse Reserve" longLatPrecision="751" longitude="27.39" pageId="7" pageNumber="478" specimenCount="2" stateProvince="Limpopo" typeStatus="holotype">
<emphasis id="B956AB6AFFD39653F7ABF943FE04F93F" bold="true" box="[151,401,1716,1741]" pageId="7" pageNumber="478">
<typeStatus id="5499C9DAFFD39653F7ABF943FF47F93F" box="[151,210,1716,1741]" pageId="7" pageNumber="478">Type</typeStatus>
data.
<typeStatus id="5499C9DAFFD39653F618F943FE04F93F" box="[292,401,1716,1741]" pageId="7" pageNumber="478" type="holotype">Holotype</typeStatus>
</emphasis>
, male (
<collectionCode id="ED33EFBDFFD39653F6D1F942FDD7F93F" box="[493,578,1717,1741]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="478">TMSA</collectionCode>
): “
<collectorName id="26D712AEFFD39653F559F942FD23F93F" box="[613,694,1716,1741]" pageId="7" pageNumber="478">S. Afr.</collectorName>
;
<collectingRegion id="49E6B99AFFD39653F5FAF943FCB9F93F" box="[710,812,1716,1741]" country="South Africa" name="Limpopo" pageId="7" pageNumber="478">Limpopo</collectingRegion>
Prov. /
<location id="8EFD21A3FFD39653F4ACF943FBDBF93E" LSID="urn:lsid:plazi:treatment:038BC66EFFD3965CF7ABF9D2FEC7F96A:8EFD21A3FFD39653F4ACF943FBDBF93E" box="[912,1102,1716,1741]" latitude="-24.36" longLatPrecision="751" longitude="27.39" name="Meletse Reserve" pageId="7" pageNumber="478" stateProvince="Limpopo">Meletse Reserve</location>
<elevation id="000F904BFFD39653F365F942FB32F93F" box="[1113,1191,1717,1741]" metricMagnitude="3" metricUnit="m" metricValue="1.003" pageId="7" pageNumber="478" unit="m" value="1003.0">
<quantity id="4CDADA9DFFD39653F365F942FB32F93F" box="[1113,1191,1717,1741]" metricMagnitude="3" metricUnit="m" metricValue="1.003" pageId="7" pageNumber="478" unit="m" value="1003.0">1003m</quantity>
</elevation>
/
<geoCoordinate id="EE1611BFFFD39653F3FAF943FA88F93F" box="[1222,1309,1716,1741]" direction="south" orientation="latitude" pageId="7" pageNumber="478" precision="555" value="-24.36">24.36 S</geoCoordinate>
–
<geoCoordinate id="EE1611BFFFD39653F210F942FA1EF93F" box="[1324,1419,1717,1741]" direction="east" orientation="longitude" pageId="7" pageNumber="478" precision="555" value="27.39">27.39 E</geoCoordinate>
”, “
<collectingDate id="EFD8A850FFD39653F79DF92EFEB1F903" box="[161,292,1753,1777]" pageId="7" pageNumber="478" value="2014-11-27">
<date id="FF9C51B8FFD39653F79DF92EFEB1F903" box="[161,292,1753,1777]" pageId="7" pageNumber="478" value="2014-11-27">27.11.2014</date>
</collectingDate>
; E-Y:3954 / on ground, bushveld / leg.
<collectorName id="26D712AEFFD39653F5D3F92FFC14F903" box="[751,897,1752,1777]" pageId="7" pageNumber="478">Ruth Müller</collectorName>
”
</materialsCitation>
.
<materialsCitation id="3B4A7D25FFD39653F4A4F92FFDFBF8E7" ID-GBIF-Occurrence="1917075916" collectingDate="2014-11-27" collectionCode="TMSA, MIZ, PAN" collectorName="S. Afr. &amp; Ruth Muller" elevation="1003" latitude="-24.36" location="Meletse Reserve" longLatPrecision="751" longitude="27.39" pageId="7" pageNumber="478" specimenCount="2" specimenCount-male="2" stateProvince="Limpopo" typeStatus="paratype">
<typeStatus id="5499C9DAFFD39653F4A4F92FFB85F903" box="[920,1040,1752,1777]" pageId="7" pageNumber="478" type="paratype">
<emphasis id="B956AB6AFFD39653F4A4F92FFB85F903" bold="true" box="[920,1040,1752,1777]" pageId="7" pageNumber="478">Paratypes</emphasis>
</typeStatus>
,
<specimenCount id="9D24BCF1FFD39653F31CF92EFBECF903" box="[1056,1145,1752,1777]" pageId="7" pageNumber="478" type="male">2 males</specimenCount>
and female (
<collectionCode id="ED33EFBDFFD39653F22DF92EFAF7F903" box="[1297,1378,1753,1777]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="478">TMSA</collectionCode>
) and male (
<collectionCode id="ED33EFBDFFD39653F7E2F90AFE86F8E7" box="[222,275,1789,1813]" pageId="7" pageNumber="478">MIZ</collectionCode>
<collectionCode id="ED33EFBDFFD39653F621F90AFECDF8E7" box="[285,344,1789,1813]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13841" name="Panjab University" pageId="7" pageNumber="478">PAN</collectionCode>
): same data as holotype
</materialsCitation>
.
<materialsCitation id="3B4A7D25FFD39653F547F90BFB8FF8CB" ID-GBIF-Occurrence="1917075872" collectingDate="1955-03-10" collectionCode="TMSA" collectorName="S. Afr. &amp; Endrody &amp; Bellamy" county="Other" latitude="-24.22" location="Geelhoutbush" longLatPrecision="751" longitude="27.33" municipality="Waterberg" pageId="7" pageNumber="478" specimenCount="1" stateProvince="Limpopo" typeStatus="paratype">
<emphasis id="B956AB6AFFD39653F547F90BFCA4F8E7" bold="true" box="[635,817,1788,1813]" pageId="7" pageNumber="478">
<collectingCounty id="62FC0FF4FFD39653F547F90BFD57F8E7" box="[635,706,1788,1813]" pageId="7" pageNumber="478">Other</collectingCounty>
material
</emphasis>
, female (
<collectionCode id="ED33EFBDFFD39653F4A3F90AFC61F8E7" box="[927,1012,1789,1813]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="7" pageNumber="478">TMSA</collectionCode>
): “
<collectorName id="26D712AEFFD39653F32AF90AFBF3F8E7" box="[1046,1126,1788,1813]" pageId="7" pageNumber="478">S. Afr.</collectorName>
:
<collectingMunicipality id="6BF9ED02FFD39653F34EF90BFB73F8E7" box="[1138,1254,1788,1813]" pageId="7" pageNumber="478">Waterberg</collectingMunicipality>
/
<location id="8EFD21A3FFD39653F23CF90BFA09F8E7" LSID="urn:lsid:plazi:treatment:038BC66EFFD3965CF7ABF9D2FEC7F96A:8EFD21A3FFD39653F23CF90BFA09F8E7" box="[1280,1436,1788,1813]" county="Other" latitude="-24.22" longLatPrecision="751" longitude="27.33" municipality="Waterberg" name="Geelhoutbush" pageId="7" pageNumber="478" stateProvince="Limpopo">Geelhoutbush</location>
farm /
<geoCoordinate id="EE1611BFFFD39653F7DDF8D6FEBAF8CB" box="[225,303,1825,1849]" direction="south" orientation="latitude" pageId="7" pageNumber="478" precision="555" value="-24.22">24.22S</geoCoordinate>
–
<geoCoordinate id="EE1611BFFFD39653F602F8D6FE0DF8CB" box="[318,408,1825,1849]" direction="east" orientation="longitude" pageId="7" pageNumber="478" precision="555" value="27.33">27.33 E</geoCoordinate>
”, “
<collectingDate id="EFD8A850FFD39653F686F8D7FDA5F8CB" box="[442,560,1824,1849]" pageId="7" pageNumber="478" value="1955-03-10">
<date id="FF9C51B8FFD39653F686F8D7FDA5F8CB" box="[442,560,1824,1849]" pageId="7" pageNumber="478" value="1955-03-10">3.10.1955</date>
</collectingDate>
; E-Y:3143 / singled /
<collectorName id="26D712AEFFD39653F41FF8D7FC11F8CB" box="[803,900,1824,1849]" pageId="7" pageNumber="478">Endrödy</collectorName>
&amp;
<collectorName id="26D712AEFFD39653F49BF8D7FB99F8CB" box="[935,1036,1824,1849]" pageId="7" pageNumber="478">Bellamy</collectorName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33824F3FFD39653F7FBF8B3FB5BF8AF" box="[199,1230,1860,1885]" pageId="7" pageNumber="478" type="etymology">
<paragraph id="8B9D7778FFD39653F7FBF8B3FB5BF8AF" blockId="7.[151,1436,1716,2029]" box="[199,1230,1860,1885]" pageId="7" pageNumber="478">
<emphasis id="B956AB6AFFD39653F7FBF8B3FEDBF8AF" bold="true" box="[199,334,1860,1885]" pageId="7" pageNumber="478">Etymology.</emphasis>
The name refers to the
<typeStatus id="5499C9DAFFD39653F56BF8B2FD12F8AF" box="[599,647,1861,1885]" pageId="7" pageNumber="478">type</typeStatus>
locality, Meletse Reserve (
<collectingRegion id="49E6B99AFFD39653F48BF8B3FBB7F8AF" box="[951,1058,1860,1885]" country="South Africa" name="Limpopo" pageId="7" pageNumber="478">Limpopo</collectingRegion>
,
<collectingCountry id="F33537E8FFD39653F311F8B3FB56F8AF" box="[1069,1219,1860,1885]" name="South Africa" pageId="7" pageNumber="478">
<collectingRegion id="49E6B99AFFD39653F311F8B3FBFBF8AF" box="[1069,1134,1860,1885]" country="Tanzania" name="Kusini Unguja" pageId="7" pageNumber="478">South</collectingRegion>
Africa
</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C33824F3FFD3965CF7FBF89FFC78FF26" lastPageId="8" lastPageNumber="479" pageId="7" pageNumber="478" type="diagnosis">
<paragraph id="8B9D7778FFD3965CF7FBF89FFC78FF26" blockId="7.[151,1436,1716,2029]" lastBlockId="8.[151,1437,151,1688]" lastPageId="8" lastPageNumber="479" pageId="7" pageNumber="478">
<emphasis id="B956AB6AFFD39653F7FBF89FFED4F873" bold="true" box="[199,321,1896,1921]" pageId="7" pageNumber="478">Diagnosis.</emphasis>
This species is similar to
<taxonomicName id="4C220CFBFFD39653F555F89EFD40F872" authorityName="Fahraeus" authorityYear="1870" box="[617,725,1896,1920]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="478" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956AB6AFFD39653F555F89EFD40F872" box="[617,725,1896,1920]" italics="true" pageId="7" pageNumber="478">A. obesus</emphasis>
</taxonomicName>
by having,
<emphasis id="B956AB6AFFD39653F467F89EFC51F872" box="[859,964,1896,1920]" italics="true" pageId="7" pageNumber="478">inter alia</emphasis>
, protibia with deep metian cavity (visible laterally) (
<figureCitation id="13196BFDFFD39653F631F87BFEFCF857" box="[269,361,1932,1957]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[151,1435,681,1847]" captionTargetId="figure@5.[151,1435,681,1847]" captionTargetPageId="5" captionText="FIGURE 2. Diagnostic characters proposed for Anaxius: mentum (A), proleg (B, G), mesoleg (C, F), metaleg (D), metaventrite (H, I). Anaxius bloubergensis (E, F), A. campbellae (A, H), A. meletsensis (G, I), and A. prozeskyi (B, C, D). Abbreviations: d—denticles, ld—surface between labial palpi, lk—longitudinal keel, lw—lateral wing, p—palpifer, t—apical denticle, tub—tubercles." httpUri="https://zenodo.org/record/1457471/files/figure.png" pageId="7" pageNumber="478">Fig. 2G</figureCitation>
), widened protarsus, with subsequent tarsomeres slightly narrower, and not apically depressed intercoxal process of prostenum (lateral view).
<taxonomicName id="4C220CFBFFD39653F58EF846FC01F83A" authorityName="Kamiński &amp; Schoeman" authorityYear="2018" box="[690,916,1968,1992]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="478" phylum="Arthropoda" rank="species" species="meletsensis">
<emphasis id="B956AB6AFFD39653F58EF846FC01F83A" box="[690,916,1968,1992]" italics="true" pageId="7" pageNumber="478">Anaxius meletsensis</emphasis>
</taxonomicName>
can be easily distinguished by having simple mesofemora (with preapical denticle in
<taxonomicName id="4C220CFBFFD39653F562F822FD3FF81F" authorityName="Fahraeus" authorityYear="1870" box="[606,682,2005,2029]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="478" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956AB6AFFD39653F562F822FD3FF81F" box="[606,682,2005,2029]" italics="true" pageId="7" pageNumber="478">obesus</emphasis>
</taxonomicName>
) and metafemora (with fringe of bristles on underside in
<taxonomicName id="4C220CFBFFD39653F203F822FA1EF81F" authorityName="Fahraeus" authorityYear="1870" box="[1343,1419,2005,2029]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="478" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956AB6AFFD39653F203F822FA1EF81F" box="[1343,1419,2005,2029]" italics="true" pageId="7" pageNumber="478">obesus</emphasis>
</taxonomicName>
). Moreover, some differences might be observed in the structure of the metatibae of both species, which are curved in the middle in
<taxonomicName id="4C220CFBFFDC965CF672FF4BFE58FF26" authorityName="Kamiński &amp; Schoeman" authorityYear="2018" box="[334,461,188,212]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="meletsensis">
<emphasis id="B956AB6AFFDC965CF672FF4BFE58FF26" box="[334,461,188,212]" italics="true" pageId="8" pageNumber="479">meletsensis</emphasis>
</taxonomicName>
(
<figureCitation id="13196BFDFFDC965CF6E1FF4BFDA0FF26" box="[477,565,188,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="8" pageNumber="479">Fig. 1D</figureCitation>
), and more basally in
<taxonomicName id="4C220CFBFFDC965CF412FF4BFCEFFF26" authorityName="Fahraeus" authorityYear="1870" box="[814,890,188,212]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956AB6AFFDC965CF412FF4BFCEFFF26" box="[814,890,188,212]" italics="true" pageId="8" pageNumber="479">obesus</emphasis>
</taxonomicName>
(
<figureCitation id="13196BFDFFDC965CF4B6FF4BFC4AFF26" box="[906,991,188,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="8" pageNumber="479">Fig. 1E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33824F3FFDC965CF7FBFF28FD22F9A2" pageId="8" pageNumber="479" type="description">
<paragraph id="8B9D7778FFDC965CF7FBFF28FC33FF0A" blockId="8.[151,1437,151,1688]" box="[199,934,223,248]" pageId="8" pageNumber="479">
<emphasis id="B956AB6AFFDC965CF7FBFF28FECCFF0A" bold="true" box="[199,345,223,248]" pageId="8" pageNumber="479">Description.</emphasis>
Body—length = 7.0–
<quantity id="4CDADA9DFFDC965CF572FF28FD33FF05" box="[590,678,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="479" unit="mm" value="7.5">7.5 mm</quantity>
. Habitus as in
<figureCitation id="13196BFDFFDC965CF475FF28FC37FF0A" box="[841,930,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="8" pageNumber="479">Fig. 1D</figureCitation>
.
</paragraph>
<paragraph id="8B9D7778FFDC965CF7FBFEF3FB01FD92" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="479">Dorsal side of head covered with confluent punctures. Frontoclypeal suture smooth. Canthus rounded at base; narrower than head at level of eyes. Clypeal emargination shallow (clypeal emargination width: depth ratio ca. 9.0). Labrum wide (width: length ratio ca. 1.9); slightly emarginate medially; with basal membrane exposed. Ligula exposed, not covered by mentum. Mentum widest in apical part; with well-developed medial keel; with exposed lateral lobes; not covering cardo or basistipes; apically with median indentation. Submentum pentagonal; short. Apical segment of maxillary palpus trapezoidal; with sensory field occupying whole proximal edge. Second segment of maxillary palpus elongate (length: width ratio ca. 2.6). Palpifer located laterally on basistipes. Anterior tentorial pit circular. Eye not dorsally shielded by any keel. Margin between maxillary fossa and antenna insertion well marked. Antenna relatively long (antenna: pronotum length ratio ca. 1.7); third antennomere short (3rd: 2nd antennomere length ratio ca. 3.0); fifth antennomere elongated (length: width ratio ca. 2.0).</paragraph>
<paragraph id="8B9D7778FFDC965CF7FBFD9AFEE6FC72" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="479">Pronotum slightly narrower than elytra (elytra: pronotum width ratio ca. 1.2); relatively narrow (length: width ratio of pronotal disc ca. 0.9); disc shiny, covered with fine punctures (0.5–1.0 diameters apart). Lateral sides of pronotal disc straight at basal half. Disc without apophyseal depressions. Basal, anterior, and lateral emargination of pronotal disc present. Base of pronotum straight. Posterior angles not protruding beyond base. Pronotal hypomeron dull; covered with very fine longitudinal rugosities; without submarginal indentation. Intercoxal process of prosternum not depressed apically (lateral view); apically with oval depression on ventral surface; slightly protruding towards first abdominal segment. Procoxae narrowly separated (procoxa: intercoxal process width ratio ca. 2.0).</paragraph>
<paragraph id="8B9D7778FFDC965CF7FBFC7BFD6EFBE2" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="479">Five elytral intervals finely marked; remaining ones indistinguishable. Elytron covered by fine punctures (0.1 diameters apart; sometimes confluent). Elytral base straight; not emarginate. Elytral humerus rounded. Epipleuron slightly narrowing apically, then of constant width; basally covering all elytral intervals (ventral view). Scutellum relatively large; triangular. Metathoracic wings absent.</paragraph>
<paragraph id="8B9D7778FFDC965CF7FBFBEBFCB7FB52" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="479">
Metaventrite extremely short (metacoxal cavity: metaventrite (between insertions of meso- and metacoxae) length ratio ca. 10.0); with longitudinal process at middle in males (
<figureCitation id="13196BFDFFDC965CF495FBC8FC6FFBAA" box="[937,1018,1087,1112]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[151,1435,681,1847]" captionTargetId="figure@5.[151,1435,681,1847]" captionTargetPageId="5" captionText="FIGURE 2. Diagnostic characters proposed for Anaxius: mentum (A), proleg (B, G), mesoleg (C, F), metaleg (D), metaventrite (H, I). Anaxius bloubergensis (E, F), A. campbellae (A, H), A. meletsensis (G, I), and A. prozeskyi (B, C, D). Abbreviations: d—denticles, ld—surface between labial palpi, lk—longitudinal keel, lw—lateral wing, p—palpifer, t—apical denticle, tub—tubercles." httpUri="https://zenodo.org/record/1457471/files/figure.png" pageId="8" pageNumber="479">
Fig.
<date id="FF9C51B8FFDC965CF4DCFBB7FC6FFBAA" box="[992,1018,1088,1112]" pageId="8" pageNumber="479">2I</date>
</figureCitation>
). Process of first abdominal ventrite wide (distances between mesocoxae: metacoxae ratio ca. 0.9). Fifth abdominal ventrite without submarginal sulcus; covered with fine punctures (4–5 diameters apart).
</paragraph>
<paragraph id="8B9D7778FFDC965CF7FBFB5BFEC9FAC2" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="479">
Male protarsus widened, with subsequent tarsomeres slightly narrower. Protibia with deep longitudinal cavity. Mesotibiae, profemora, mesofemora, and metafemora simple. Metafemora similar to that of
<taxonomicName id="4C220CFBFFDC965CF3A1FB27FA9DFB1A" authorityName="Fahraeus" authorityYear="1870" box="[1181,1288,1232,1256]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956AB6AFFDC965CF3A1FB27FA9DFB1A" box="[1181,1288,1232,1256]" italics="true" pageId="8" pageNumber="479">A. obesus</emphasis>
</taxonomicName>
. Female legs simple. First tarsomere of metatarsi elongated (1.7× longer than 4th one). Length of metatarsus equal to 0.75 of metatibial length.
</paragraph>
<paragraph id="8B9D7778FFDC965CF7FBFACBFA93FA6F" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="479">Aedeagal tegmen widest in middle; unipartite; with wide basal gap. Apical part slightly divided (up to 0.10 of whole tegmen length). Clava straight. Penis of even width along most of its length; with apical part not covered by tegmen, exposed dorsally. Basal apophyses of penis relatively short (ca. 0.4 of whole tegmen length).</paragraph>
<paragraph id="8B9D7778FFDC965CF7FBFA50FD22F9A2" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="479">Ovipositor relatively short (body: ovipositor length ratio ca. 3.0). Paraproct much longer than coxites (paraproct: coxites length ratio ca. 1.8); triangular, not shielding valvifer or any other lobes of coxites. Valvifer and second lobe slightly transverse, while third one triangular. Fourth lobe rounded, situated dorsally. Gonostylus located on dorsal side of fourth lobe. Vagina and bursa copulatrix without sclerites. Spermatheca sac-like. Proctiger covering nearly whole ventral side of ovipositor.</paragraph>
</subSubSection>
<subSubSection id="C33824F3FFDC965CF7FBF9ABFEC7F96A" pageId="8" pageNumber="479" type="distribution">
<paragraph id="8B9D7778FFDC965CF7FBF9ABFEC7F96A" blockId="8.[151,1437,151,1688]" pageId="8" pageNumber="479">
<emphasis id="B956AB6AFFDC965CF7FBF9ABFEF5F987" bold="true" box="[199,352,1628,1653]" pageId="8" pageNumber="479">Distribution.</emphasis>
This species has been collected in the following ecoregion of
<collectingCountry id="F33537E8FFDC965CF308F9ABFB53F987" box="[1076,1222,1628,1653]" name="South Africa" pageId="8" pageNumber="479">
<collectingRegion id="49E6B99AFFDC965CF308F9ABFBE3F987" box="[1076,1142,1628,1653]" country="Tanzania" name="Kusini Unguja" pageId="8" pageNumber="479">South</collectingRegion>
Africa
</collectingCountry>
(
<figureCitation id="13196BFDFFDC965CF3E4F9ABFA8AF987" box="[1240,1311,1628,1653]" captionStart="FIGURE 5" captionStartId="11.[151,250,1731,1753]" captionTargetBox="[161,1437,844,1704]" captionTargetId="figure@11.[151,1437,844,1710]" captionTargetPageId="11" captionText="FIGURE 5. Distribution of the species representing Anaxius." httpUri="https://zenodo.org/record/1457477/files/figure.png" pageId="8" pageNumber="479">Fig. 5</figureCitation>
): Southern Africa bushveld.
</paragraph>
</subSubSection>
</treatment>
</document>
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