128 lines
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128 lines
13 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/fr.25.83781" ID-Pensoft-Pub="2193-0074-1-99" ID-Pensoft-UUID="2F5D6AE2EEB55A17ACF1623B06B4EA8D" ID-ZooBank="7A16698D4F1848D29D9651A6E0CC15AC" ModsDocID="2193-0074-25-1-99" checkinTime="1667366289781" checkinUser="pensoft" docAuthor="Vasilyan, Davit, Cernansky, Andrej, Szyndlar, Zbigniew & Moers, Thomas" docDate="2022" docId="003739346CB55964AA298E739A26F7CC" docLanguage="en" docName="FossilRecord 25(1): 99-145" docOrigin="Fossil Record 25 (1)" docPubDate="2022-05-10" docSource="http://dx.doi.org/10.3897/fr.25.83781" docTitle="Rana sp." docType="treatment" docVersion="1" id="2F5D6AE2EEB55A17ACF1623B06B4EA8D" lastPageNumber="99" masterDocId="2F5D6AE2EEB55A17ACF1623B06B4EA8D" masterDocTitle="Amphibian and reptilian fauna from the early Miocene of Echzell, Germany" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1667366289781" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Amphibian and reptilian fauna from the early Miocene of Echzell, Germany</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vasilyan, Davit</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8712-0678</mods:nameIdentifier>
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<mods:affiliation>JURASSICA Museum, Route de Fontenais 21. 2900 Porrentruy, Switzerland & Department of Geosciences, University of Fribourg, Chemin du musee 6, 1700, Fribourg, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">davit.vasilyan@jurassica.ch</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cernansky, Andrej</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8920-2503</mods:nameIdentifier>
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<mods:affiliation>Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina, Ilkovicova 6, 842 15 Bratislava, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szyndlar, Zbigniew</mods:namePart>
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<mods:affiliation>Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31 - 016 Krakow, Poland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moers, Thomas</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2268-5824</mods:nameIdentifier>
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<mods:affiliation>Department of Palaeobiology, Swedish Museum of Natural History, P. O. Box 50007, 104 05 Stockholm, Sweden & Bolin Centre for Climate Research, Stockholm University, Stockholm, Sweden</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Fossil Record</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-05-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>25</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>99</mods:start>
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<mods:end>145</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/fr.25.83781</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/fr.25.83781</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2193-0074-1-99</mods:identifier>
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<mods:identifier type="ZooBank">7A16698D4F1848D29D9651A6E0CC15AC</mods:identifier>
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<mods:identifier type="Pensoft-UUID">2F5D6AE2EEB55A17ACF1623B06B4EA8D</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:003739346CB55964AA298E739A26F7CC" httpUri="http://treatment.plazi.org/id/003739346CB55964AA298E739A26F7CC" lastPageNumber="99" pageId="0" pageNumber="99">
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<subSubSection pageId="0" pageNumber="99" type="nomenclature">
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName LSID="00373934-6CB5-5964-AA29-8E739A26F7CC" authority="sp." authorityName="sp." class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">Rana sp.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="description">
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<paragraph pageId="0" pageNumber="99">
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Frogs from Echzell locality. Ilia of Pelophylax sp. HLMD-Ez 2089 (A, B) and HLMD-Ez 2090 (C, D). Ilia of Rana sp. HLMD-Ez 2093 (E, F) and HLMD-Ez 2094 (G, H). Palaeobatrachus robustus (I-W). Frontoparietals HLMD-Ez 2077 (I, J), HLMD-Ez 2078 (K) and HLMD-Ez 2076 (L, M). Jaw bone HLMD-Ez 2084 (N, O). Angulars HLMD-Ez 2083 (P) and HLMD-Ez 2082 (Q). Ilia HLMD-Ez 2079 (R-T) and HLMD-Ez 2080 (V, W). Bones are figured in (A, D, E, G, M, R, V) lateral, (B, C, F, H, S) posterior, (I, K, L, P, Q) dorsal, (J, O) ventral, (N) labial and (T, W) medial views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure9" httpUri="https://binary.pensoft.net/fig/682229" pageId="0" pageNumber="99">Fig. 9E-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="materials_examined">
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<paragraph pageId="0" pageNumber="99">Material.</paragraph>
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<paragraph pageId="0" pageNumber="99">Three ilia HLMD-Ez 2092-2094.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="description">
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<paragraph pageId="0" pageNumber="99">Description.</paragraph>
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<paragraph pageId="0" pageNumber="99">
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The best-preserved ilium (HLMD-Ez 2094, Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Frogs from Echzell locality. Ilia of Pelophylax sp. HLMD-Ez 2089 (A, B) and HLMD-Ez 2090 (C, D). Ilia of Rana sp. HLMD-Ez 2093 (E, F) and HLMD-Ez 2094 (G, H). Palaeobatrachus robustus (I-W). Frontoparietals HLMD-Ez 2077 (I, J), HLMD-Ez 2078 (K) and HLMD-Ez 2076 (L, M). Jaw bone HLMD-Ez 2084 (N, O). Angulars HLMD-Ez 2083 (P) and HLMD-Ez 2082 (Q). Ilia HLMD-Ez 2079 (R-T) and HLMD-Ez 2080 (V, W). Bones are figured in (A, D, E, G, M, R, V) lateral, (B, C, F, H, S) posterior, (I, K, L, P, Q) dorsal, (J, O) ventral, (N) labial and (T, W) medial views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure9" httpUri="https://binary.pensoft.net/fig/682229" pageId="0" pageNumber="99">9G, H</figureCitation>
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) shows a well-developed (rather narrow and high) dorsal acetabular expansion and a reduced (broad and low) ventral acetabular expansion. The dorsal prominence is not very high. It is oriented rather anteriorly than anterodorsally. The dorsal protuberance has an oval outline. Its surface is either smooth (HLMD-Ez 2094, Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Frogs from Echzell locality. Ilia of Pelophylax sp. HLMD-Ez 2089 (A, B) and HLMD-Ez 2090 (C, D). Ilia of Rana sp. HLMD-Ez 2093 (E, F) and HLMD-Ez 2094 (G, H). Palaeobatrachus robustus (I-W). Frontoparietals HLMD-Ez 2077 (I, J), HLMD-Ez 2078 (K) and HLMD-Ez 2076 (L, M). Jaw bone HLMD-Ez 2084 (N, O). Angulars HLMD-Ez 2083 (P) and HLMD-Ez 2082 (Q). Ilia HLMD-Ez 2079 (R-T) and HLMD-Ez 2080 (V, W). Bones are figured in (A, D, E, G, M, R, V) lateral, (B, C, F, H, S) posterior, (I, K, L, P, Q) dorsal, (J, O) ventral, (N) labial and (T, W) medial views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure9" httpUri="https://binary.pensoft.net/fig/682229" pageId="0" pageNumber="99">9G</figureCitation>
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) or possesses muscle scars (HLMD-Ez 2093, Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Frogs from Echzell locality. Ilia of Pelophylax sp. HLMD-Ez 2089 (A, B) and HLMD-Ez 2090 (C, D). Ilia of Rana sp. HLMD-Ez 2093 (E, F) and HLMD-Ez 2094 (G, H). Palaeobatrachus robustus (I-W). Frontoparietals HLMD-Ez 2077 (I, J), HLMD-Ez 2078 (K) and HLMD-Ez 2076 (L, M). Jaw bone HLMD-Ez 2084 (N, O). Angulars HLMD-Ez 2083 (P) and HLMD-Ez 2082 (Q). Ilia HLMD-Ez 2079 (R-T) and HLMD-Ez 2080 (V, W). Bones are figured in (A, D, E, G, M, R, V) lateral, (B, C, F, H, S) posterior, (I, K, L, P, Q) dorsal, (J, O) ventral, (N) labial and (T, W) medial views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure9" httpUri="https://binary.pensoft.net/fig/682229" pageId="0" pageNumber="99">9E</figureCitation>
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). The iliac shaft is rather thin and round in cross-section. A low and thin bone lamina connects the iliac shaft and the dorsal prominence. In posterior view, the dorsal prominence projects laterodorsally. The ilioischiatic juncture is narrow (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Frogs from Echzell locality. Ilia of Pelophylax sp. HLMD-Ez 2089 (A, B) and HLMD-Ez 2090 (C, D). Ilia of Rana sp. HLMD-Ez 2093 (E, F) and HLMD-Ez 2094 (G, H). Palaeobatrachus robustus (I-W). Frontoparietals HLMD-Ez 2077 (I, J), HLMD-Ez 2078 (K) and HLMD-Ez 2076 (L, M). Jaw bone HLMD-Ez 2084 (N, O). Angulars HLMD-Ez 2083 (P) and HLMD-Ez 2082 (Q). Ilia HLMD-Ez 2079 (R-T) and HLMD-Ez 2080 (V, W). Bones are figured in (A, D, E, G, M, R, V) lateral, (B, C, F, H, S) posterior, (I, K, L, P, Q) dorsal, (J, O) ventral, (N) labial and (T, W) medial views. Scale bars: 1 mm." figureDoi="10.3897/fr.25.83781.figure9" httpUri="https://binary.pensoft.net/fig/682229" pageId="0" pageNumber="99">9H</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="remarks">
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<paragraph pageId="0" pageNumber="99">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="99">
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Echzell frog ilia can be clearly assigned to green
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<taxonomicName class="Amphibia" family="Ranidae" genus="Pelophylax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelophylax" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Pelophylax</emphasis>
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</taxonomicName>
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and brown
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<taxonomicName class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Rana</emphasis>
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</taxonomicName>
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frogs based on the following characters. The genus
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<taxonomicName class="Amphibia" family="Ranidae" genus="Pelophylax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelophylax" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Pelophylax</emphasis>
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</taxonomicName>
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is characterised by: anterodorsally oriented, high, large and dorsomedially curved dorsal prominence and protuberance; the dorsal protuberance has a smooth surface; whereas the genus
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<taxonomicName class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana" order="Anura" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="99">Rana</emphasis>
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</taxonomicName>
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by in lateral view anterodorsally and in posterior view laterodorsally oriented, rather low and reduced dorsal prominence and protuberance, the dorsal protuberance has smooth or irregular surface (
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<bibRefCitation author="Boehme, G" journalOrPublisher="Wissenschaftliche Zeitschrift der Humboldt-Universitaet zu Berlin, Mathematisch-Naturwissenschaftliche Reihe" pageId="0" pageNumber="99" pagination="283 - 299" refId="B5" refString="Boehme, G, 1977. Zur Bestimmung quartaerer Anuren Europas an Hand von Skelettelementen. Wissenschaftliche Zeitschrift der Humboldt-Universitaet zu Berlin, Mathematisch-Naturwissenschaftliche Reihe 26: 283 - 299" title="Zur Bestimmung quartaerer Anuren Europas an Hand von Skelettelementen." volume="26" year="1977">
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<normalizedToken originalValue="Böhme">Boehme</normalizedToken>
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1977
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</bibRefCitation>
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;
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<bibRefCitation author="Blain, H-A" journalOrPublisher="Geodiversitas" pageId="0" pageNumber="99" pagination="269 - 295" refId="B3" refString="Blain, H-A, Bailon, S, Agusti, J, 2007. Anurans and squamate reptiles from the latest early Pleistocene of Almenara-Casablanca-3 (Castellon, East of Spain). Systematic, climatic and environmental considerations. Geodiversitas 29: 269 - 295" title="Anurans and squamate reptiles from the latest early Pleistocene of Almenara-Casablanca- 3 (Castellon, East of Spain). Systematic, climatic and environmental considerations." volume="29" year="2007">Blain et al. 2007</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |