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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.881.39535" ID-GBIF-Dataset="2f6463a2-b616-4b94-8708-0851aa4176cc" ID-PMC="PMC6813175" ID-Pensoft-Pub="1313-2970-881-13" ID-Pensoft-UUID="C842A0652FAE520EB140F739AF32A854" ID-PubMed="31662609" ID-ZooBank="175E7C92A110446480D4FA0EAC3473B1" ModsDocID="1313-2970-881-13" checkinTime="1571384713076" checkinUser="pensoft" docAuthor="Choi, Su-Ho, Chandler, Donald S. &amp; Park, Jong-Seok" docDate="2019" docId="931DA0B40EBD5F03B5D8F602DCE57BAE" docLanguage="en" docName="ZooKeys 881: 13-22" docOrigin="ZooKeys 881" docSource="http://dx.doi.org/10.3897/zookeys.881.39535" docTitle="Porongurup clarkei Choi, Chandler &amp; Park, 2019, sp. nov." docType="treatment" docUuid="11556FE3-6FDF-4265-999C-C160B8F6F17E" docUuidSource="ZooBank" docVersion="5" id="C842A0652FAE520EB140F739AF32A854" lastPageNumber="20" masterDocId="C842A0652FAE520EB140F739AF32A854" masterDocTitle="Porongurup, a new genus of pselaphine staphylinid beetles from Western Australia (Coleoptera, Staphylinidae, Pselaphinae, Faronitae)" masterLastPageNumber="22" masterPageNumber="13" pageNumber="18" updateTime="1668167894361" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Porongurup, a new genus of pselaphine staphylinid beetles from Western Australia (Coleoptera, Staphylinidae, Pselaphinae, Faronitae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Choi, Su-Ho</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chandler, Donald S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Park, Jong-Seok</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>881</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>22</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.881.39535</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.881.39535</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-881-13</mods:identifier>
<mods:identifier type="ZooBank">175E7C92A110446480D4FA0EAC3473B1</mods:identifier>
<mods:identifier type="Pensoft-UUID">C842A0652FAE520EB140F739AF32A854</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159754700" LSID="urn:lsid:zoobank.org:act:11556FE3-6FDF-4265-999C-C160B8F6F17E" httpUri="http://treatment.plazi.org/id/931DA0B40EBD5F03B5D8F602DCE57BAE" lastPageId="7" lastPageNumber="20" pageId="5" pageNumber="18">
<subSubSection pageId="5" pageNumber="18" type="nomenclature">
<paragraph pageId="5" pageNumber="18">
<taxonomicName LSID="931DA0B4-0EBD-5F03-B5D8-F602DCE57BAE" class="Insecta" family="Staphylinidae" genus="Porongurup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porongurup clarkei" order="Coleoptera" pageId="5" pageNumber="18" phylum="Arthropoda" rank="species" species="clarkei">
<pageBreakToken pageId="5" pageNumber="18" start="start">Porongurup</pageBreakToken>
clarkei
</taxonomicName>
<taxonomicNameLabel pageId="5" pageNumber="18">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus photos, dorsal view A Porongurup angulatus sp. nov. B P. clarkei sp. nov. C P. tenuis sp. nov. Female habitus, dorsal view D P. angulatus sp. nov. E P. clarkei sp. nov. F P. tenuis sp. nov. Scale bar: 1 mm." figureDoi="10.3897/zookeys.881.39535.figure1" httpUri="https://binary.pensoft.net/fig/348328" pageId="5" pageNumber="18" tableDoi="10.3897/zookeys.881.39535.figure1">Figs 1B</figureCitation>
,
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus photos, dorsal view A Porongurup angulatus sp. nov. B P. clarkei sp. nov. C P. tenuis sp. nov. Female habitus, dorsal view D P. angulatus sp. nov. E P. clarkei sp. nov. F P. tenuis sp. nov. Scale bar: 1 mm." figureDoi="10.3897/zookeys.881.39535.figure1" httpUri="https://binary.pensoft.net/fig/348328" pageId="5" pageNumber="18" tableDoi="10.3897/zookeys.881.39535.figure1">1E</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of the species of Porongurup gen. nov. A P. angulatus sp. nov., ventral view B P. clarkei sp. nov., ventral view C P. clarkei sp. nov., lateral view D P. tenuis sp. nov., ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.881.39535.figure2" httpUri="https://binary.pensoft.net/fig/348329" pageId="5" pageNumber="18" tableDoi="10.3897/zookeys.881.39535.figure2">
2
<normalizedToken originalValue="B–C">B-C</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae of Porongurup angulatus sp. nov. A male B female. Antennae of P. clarkei sp. nov. C male D female. Antennae of P. tenuis sp. nov. E male F female. Male head dorsal view G P. angulatus sp. nov. H P. clarkei sp. nov. I P. tenuis. sp. nov. Female head dorsal view J P. angulatus sp. nov. K P. clarkei sp. nov. L P. tenuis sp. nov. Scale bar: 0.1 mm. * The distinctness and position of the vertexal foveae may be affected by the angle of view and condition of the specimens." figureDoi="10.3897/zookeys.881.39535.figure4" httpUri="https://binary.pensoft.net/fig/348331" pageId="5" pageNumber="18" tableDoi="10.3897/zookeys.881.39535.figure4">
4
<normalizedToken originalValue="C–D">C-D</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae of Porongurup angulatus sp. nov. A male B female. Antennae of P. clarkei sp. nov. C male D female. Antennae of P. tenuis sp. nov. E male F female. Male head dorsal view G P. angulatus sp. nov. H P. clarkei sp. nov. I P. tenuis. sp. nov. Female head dorsal view J P. angulatus sp. nov. K P. clarkei sp. nov. L P. tenuis sp. nov. Scale bar: 0.1 mm. * The distinctness and position of the vertexal foveae may be affected by the angle of view and condition of the specimens." figureDoi="10.3897/zookeys.881.39535.figure4" httpUri="https://binary.pensoft.net/fig/348331" pageId="5" pageNumber="18" tableDoi="10.3897/zookeys.881.39535.figure4">4H</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae of Porongurup angulatus sp. nov. A male B female. Antennae of P. clarkei sp. nov. C male D female. Antennae of P. tenuis sp. nov. E male F female. Male head dorsal view G P. angulatus sp. nov. H P. clarkei sp. nov. I P. tenuis. sp. nov. Female head dorsal view J P. angulatus sp. nov. K P. clarkei sp. nov. L P. tenuis sp. nov. Scale bar: 0.1 mm. * The distinctness and position of the vertexal foveae may be affected by the angle of view and condition of the specimens." figureDoi="10.3897/zookeys.881.39535.figure4" httpUri="https://binary.pensoft.net/fig/348331" pageId="5" pageNumber="18" tableDoi="10.3897/zookeys.881.39535.figure4">4K</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Collection localities of Porongurup gen. nov. Key: P. angulatus sp. nov., circles; P. clarkei sp. nov., triangle; P. tenuis sp. nov., squares." figureDoi="10.3897/zookeys.881.39535.figure5" httpUri="https://binary.pensoft.net/fig/348332" pageId="5" pageNumber="18" tableDoi="10.3897/zookeys.881.39535.figure5">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="20" pageId="6" pageNumber="19" type="type material">
<paragraph pageId="6" pageNumber="19">
<pageBreakToken pageId="6" pageNumber="19" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph lastPageId="7" lastPageNumber="20" pageId="6" pageNumber="19">
<emphasis bold="true" pageId="6" pageNumber="19">
<emphasis italics="true" pageId="6" pageNumber="19">Holotype</emphasis>
. Australia: Western Australia (WA)
</emphasis>
: 1♂, aedeagus dissected and mounted in Euparal on clear plastic card, &quot;
<emphasis bold="true" pageId="6" pageNumber="19">AUSTRALIA: Western Australia</emphasis>
: Stirling Range N.P., Toolbrunup Peak Track, 480-520 m,
<geoCoordinate degrees="34" direction="south" minutes="23.4" orientation="latitude" precision="92" value="-34.39">34°23.4'S</geoCoordinate>
,
<geoCoordinate degrees="118" direction="east" minutes="03.3" orientation="longitude" precision="92" value="118.055">118°03.3'E</geoCoordinate>
, 5 VIII 2004,
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus" order="Myrtales" pageId="6" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="6" pageNumber="19">Eucalyptus</emphasis>
</taxonomicName>
forest &amp; mallee; FMHD#2004-146, Berlese, leaf &amp; log litter,
<pageBreakToken pageId="7" pageNumber="20" start="start">Clarke</pageBreakToken>
&amp; Grimbacher 1115&quot;.
<emphasis bold="true" pageId="7" pageNumber="20">
<emphasis italics="true" pageId="7" pageNumber="20">Paratypes</emphasis>
(
<emphasis italics="true" pageId="7" pageNumber="20">N</emphasis>
= 11; 7 males, 4 females). Australia: Western Australia
</emphasis>
: 1♂2♀ (FMNH, 1♀ slide mounted), Stirling Range N.P., Toolbrunup Peak Track, 430-485m,
<geoCoordinate degrees="34" direction="south" minutes="23.5" orientation="latitude" precision="92" value="-34.391666">34°23.5'S</geoCoordinate>
,
<geoCoordinate degrees="118" direction="east" minutes="03.65" orientation="longitude" precision="9" value="118.06084">118°03.65'E</geoCoordinate>
, 5 VIII 2004, mallee
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus" order="Myrtales" pageId="7" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="20">Eucalyptus</emphasis>
</taxonomicName>
; FMHD#2004-145, Berlese, water-washed soil, 0-18 cm, D. Clarke 1114; 5♂ (UNHC), 1♂ aedeagus dissected and mounted in Euparal on clear plastic card, Porongorup N.P., Bolganup Creek, 12 VI 1980, Berlese, bark &amp; fungi karri tree bases, S. &amp;J. Peck; 1♂ (CBNUIC, aedeagus dissected and mounted in Euparal on clear plastic card, Stirling Range N.P., Toolbrunup Tr. WA, 10 VI 1980, Berlese, rotted logs &amp; moss, S. &amp; J. Peck, SBP47; 1♀(FMNH), Stirling Range N.P., Toolbrunup Tr. WA, 10 VI 1980, Berlese, fungi on Euc. Trunks, S. &amp; J. Peck, SBP45; 1♀(UNHC), Porongorup N.P., Bolganup Creek, 12 VI 1980, Berlese, bark &amp; fungi on karri tree bases, S. &amp; J. Peck, SBP53.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="20" type="diagnosis">
<paragraph pageId="7" pageNumber="20">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="20">
This species can be distinguished from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Porongurup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porongurup angulatus" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis italics="true" pageId="7" pageNumber="20">Porongurup angulatus</emphasis>
</taxonomicName>
sp. nov. by antennomere 3 being longer than wide (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae of Porongurup angulatus sp. nov. A male B female. Antennae of P. clarkei sp. nov. C male D female. Antennae of P. tenuis sp. nov. E male F female. Male head dorsal view G P. angulatus sp. nov. H P. clarkei sp. nov. I P. tenuis. sp. nov. Female head dorsal view J P. angulatus sp. nov. K P. clarkei sp. nov. L P. tenuis sp. nov. Scale bar: 0.1 mm. * The distinctness and position of the vertexal foveae may be affected by the angle of view and condition of the specimens." figureDoi="10.3897/zookeys.881.39535.figure4" httpUri="https://binary.pensoft.net/fig/348331" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure4">Fig. 4C</figureCitation>
), and abdominal segment VI being approximately three times longer than V (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus photos, dorsal view A Porongurup angulatus sp. nov. B P. clarkei sp. nov. C P. tenuis sp. nov. Female habitus, dorsal view D P. angulatus sp. nov. E P. clarkei sp. nov. F P. tenuis sp. nov. Scale bar: 1 mm." figureDoi="10.3897/zookeys.881.39535.figure1" httpUri="https://binary.pensoft.net/fig/348328" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure1">Fig. 1B, E</figureCitation>
). This species is also separated from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Porongurup" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porongurup tenuis" order="Coleoptera" pageId="7" pageNumber="20" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis italics="true" pageId="7" pageNumber="20">Porongurup tenuis</emphasis>
</taxonomicName>
sp. nov. by the median lobe of the male genitalia having a small digit at the apex (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of the species of Porongurup gen. nov. A P. angulatus sp. nov., ventral view B P. clarkei sp. nov., ventral view C P. clarkei sp. nov., lateral view D P. tenuis sp. nov., ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.881.39535.figure2" httpUri="https://binary.pensoft.net/fig/348329" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure2">Fig. 2B</figureCitation>
: arrowed).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="20" type="description">
<paragraph pageId="7" pageNumber="20">Description.</paragraph>
<paragraph pageId="7" pageNumber="20">
Length 1.6-1.9 mm (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus photos, dorsal view A Porongurup angulatus sp. nov. B P. clarkei sp. nov. C P. tenuis sp. nov. Female habitus, dorsal view D P. angulatus sp. nov. E P. clarkei sp. nov. F P. tenuis sp. nov. Scale bar: 1 mm." figureDoi="10.3897/zookeys.881.39535.figure1" httpUri="https://binary.pensoft.net/fig/348328" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure1">Fig. 1B, E</figureCitation>
).
<emphasis italics="true" pageId="7" pageNumber="20">Head.</emphasis>
Head in dorsal view with both shallow frontal sulcus and vertexal foveae. Male and female antennae are almost identical in length (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae of Porongurup angulatus sp. nov. A male B female. Antennae of P. clarkei sp. nov. C male D female. Antennae of P. tenuis sp. nov. E male F female. Male head dorsal view G P. angulatus sp. nov. H P. clarkei sp. nov. I P. tenuis. sp. nov. Female head dorsal view J P. angulatus sp. nov. K P. clarkei sp. nov. L P. tenuis sp. nov. Scale bar: 0.1 mm. * The distinctness and position of the vertexal foveae may be affected by the angle of view and condition of the specimens." figureDoi="10.3897/zookeys.881.39535.figure4" httpUri="https://binary.pensoft.net/fig/348331" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure4">Fig. 4C, D</figureCitation>
). Antennomeres II longer than wide, III subquadrate and smallest,
<normalizedToken originalValue="IV—VIII">IV-VIII</normalizedToken>
longer than wide, IX and X transverse (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae of Porongurup angulatus sp. nov. A male B female. Antennae of P. clarkei sp. nov. C male D female. Antennae of P. tenuis sp. nov. E male F female. Male head dorsal view G P. angulatus sp. nov. H P. clarkei sp. nov. I P. tenuis. sp. nov. Female head dorsal view J P. angulatus sp. nov. K P. clarkei sp. nov. L P. tenuis sp. nov. Scale bar: 0.1 mm. * The distinctness and position of the vertexal foveae may be affected by the angle of view and condition of the specimens." figureDoi="10.3897/zookeys.881.39535.figure4" httpUri="https://binary.pensoft.net/fig/348331" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure4">Fig. 4C, D</figureCitation>
).
<emphasis italics="true" pageId="7" pageNumber="20">Elytra</emphasis>
. Male elytra longer than wide, female elytra shorter, as long as wide (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus photos, dorsal view A Porongurup angulatus sp. nov. B P. clarkei sp. nov. C P. tenuis sp. nov. Female habitus, dorsal view D P. angulatus sp. nov. E P. clarkei sp. nov. F P. tenuis sp. nov. Scale bar: 1 mm." figureDoi="10.3897/zookeys.881.39535.figure1" httpUri="https://binary.pensoft.net/fig/348328" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure1">Fig. 1B, E</figureCitation>
).
<emphasis italics="true" pageId="7" pageNumber="20">Abdomen</emphasis>
. Abdominal segment VI approximately three times longer than V (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Male habitus photos, dorsal view A Porongurup angulatus sp. nov. B P. clarkei sp. nov. C P. tenuis sp. nov. Female habitus, dorsal view D P. angulatus sp. nov. E P. clarkei sp. nov. F P. tenuis sp. nov. Scale bar: 1 mm." figureDoi="10.3897/zookeys.881.39535.figure1" httpUri="https://binary.pensoft.net/fig/348328" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure1">Fig. 1B, E</figureCitation>
).
<emphasis italics="true" pageId="7" pageNumber="20">Aedeagus.</emphasis>
Median lobe of male genitalia as long as parameres with apical articulated extension. Phallobase rounded in dorsal view. Parameres symmetrical (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of the species of Porongurup gen. nov. A P. angulatus sp. nov., ventral view B P. clarkei sp. nov., ventral view C P. clarkei sp. nov., lateral view D P. tenuis sp. nov., ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.881.39535.figure2" httpUri="https://binary.pensoft.net/fig/348329" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure2">Fig. 2B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="20" type="etymology">
<paragraph pageId="7" pageNumber="20">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="20">This species is named for one of the collectors of the holotype, the staphylinid specialist Dave J. Clarke.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="20" type="distribution">
<paragraph pageId="7" pageNumber="20">Distribution.</paragraph>
<paragraph pageId="7" pageNumber="20">
Western Australia (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Collection localities of Porongurup gen. nov. Key: P. angulatus sp. nov., circles; P. clarkei sp. nov., triangle; P. tenuis sp. nov., squares." figureDoi="10.3897/zookeys.881.39535.figure5" httpUri="https://binary.pensoft.net/fig/348332" pageId="7" pageNumber="20" tableDoi="10.3897/zookeys.881.39535.figure5">Fig. 5</figureCitation>
, triangles).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="20" type="habitat">
<paragraph pageId="7" pageNumber="20">Habitat.</paragraph>
<paragraph pageId="7" pageNumber="20">
Specimens of this species were collected by sifting leaf, log, bark, moss or fungus litter in
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus" order="Myrtales" pageId="7" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="20">Eucalyptus</emphasis>
</taxonomicName>
forests, or were taken by Berlese funnel water-washed soil.
</paragraph>
</subSubSection>
</treatment>
</document>
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