154 lines
13 KiB
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154 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.660.11780" ID-GBIF-Dataset="17e2f87e-a5a1-493f-af93-e2b128efdb44" ID-PMC="PMC5549529" ID-Pensoft-Pub="1313-2970-660-43" ID-PubMed="28794673" ID-ZBK="A64E093A34564C5692305C449223F1B8" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-660-43" ModsDocOrigin="ZooKeys 660" ModsDocTitle="Four new species of the millipede genus Eutrichodesmus Silvestri, 1910 from Laos, including two with reduced ozopores (Diplopoda, Polydesmida, Haplodesmidae)" checkinTime="1489056688359" checkinUser="pensoft" docAuthor="Liu, Weixin, Golovatch, Sergei & Wesener, Thomas" docDate="2017" docId="8D456526A318D83F2861F7917042FD11" docLanguage="en" docName="ZooKeys 660: 43-65" docOrigin="ZooKeys 660" docSource="http://dx.doi.org/10.3897/zookeys.660.11780" docTitle="Eutrichodesmus paraster Liu & Wesener, sp. n." docType="treatment" docUuid="7EC7CE3B-1990-49E5-953C-B0B8D5B46DEB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="49" masterDocId="FFA4FFB8FFE7FFC8FFB98A32FFE5B800" masterDocTitle="Four new species of the millipede genus Eutrichodesmus Silvestri, 1910 from Laos, including two with reduced ozopores (Diplopoda, Polydesmida, Haplodesmidae)" masterLastPageNumber="65" masterPageNumber="43" pageNumber="47" updateTime="1668164283224" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Four new species of the millipede genus Eutrichodesmus Silvestri, 1910 from Laos, including two with reduced ozopores (Diplopoda, Polydesmida, Haplodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liu, Weixin</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>660</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>43</mods:start>
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<mods:end>65</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.660.11780</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.660.11780</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-660-43</mods:identifier>
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<mods:identifier type="ZBK">A64E093A34564C5692305C449223F1B8</mods:identifier>
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<mods:identifier type="ZooBank">A64E093A34564C5692305C449223F1B8</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127902352" LSID="urn:lsid:zoobank.org:act:7EC7CE3B-1990-49E5-953C-B0B8D5B46DEB" httpUri="http://treatment.plazi.org/id/8D456526A318D83F2861F7917042FD11" lastPageId="6" lastPageNumber="49" pageId="4" pageNumber="47">
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<subSubSection pageId="4" pageNumber="47" type="nomenclature">
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<paragraph pageId="4" pageNumber="47">
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<taxonomicName LSID="http://zoobank.org/7EC7CE3B-1990-49E5-953C-B0B8D5B46DEB" authority="Liu & Wesener" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus paraster" order="Polydesmida" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="paraster">Eutrichodesmus paraster Liu & Wesener</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="47">sp. n.</taxonomicNameLabel>
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Figs 1C, 8, 9, 10
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="47" type="materials_examined">
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<paragraph pageId="4" pageNumber="47">Material examined.</paragraph>
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<paragraph pageId="4" pageNumber="47">
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Holotype male (SEM), (SMF), Laos, Huaphan Prov., Xop, Cave Tham Long Puang (F 48-123-001),
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.473806">N20°28'25.7"</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="103.362335">E103°21'44.4"</geoCoordinate>
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, 16.I.2009, coll. H. Steiner (101/09-).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="47" type="paratypes">
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<paragraph pageId="4" pageNumber="47">Paratypes.</paragraph>
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<paragraph pageId="4" pageNumber="47">1 female (SMF), same data as holotype; 1 juvenile (ZFMK MYR6131), same data.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="47" type="etymology">
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<paragraph pageId="4" pageNumber="47">Etymology.</paragraph>
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<paragraph pageId="4" pageNumber="47">
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To emphasize the similarity to
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<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus aster" order="Polydesmida" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="aster">Eutrichodesmus aster</taxonomicName>
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Golovatch, Geoffroy,
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<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
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& VandenSpiegel, 2009; adjective.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="47" type="diagnosis">
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<paragraph pageId="4" pageNumber="47">Diagnosis.</paragraph>
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<paragraph pageId="4" pageNumber="47">
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Differs from other species of the genus primarily by the completely reduced ozopores. Superficially very similar to
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<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus aster" order="Polydesmida" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="aster">Eutrichodesmus aster</taxonomicName>
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, but distinguished from the latter through the smaller body, laterally 3-lobulated paraterga, and the relatively complex gonopod showing a large, laterally denticulate, distofemoral process; the acropodite subapically has a very small mesal tooth and an evident, digitiform, dorsal lobule. See also Key above.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="49" pageId="4" pageNumber="47" type="description">
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<paragraph pageId="4" pageNumber="47">Description.</paragraph>
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<paragraph pageId="4" pageNumber="47">Length of adults ca. 8.0 mm (holotype) or 9.0 mm (paratype), width 1.0 mm and 2.5 mm on midbody pro- and metazona, respectively.</paragraph>
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<paragraph pageId="4" pageNumber="47">Coloration uniformly pallid (Fig. 1C).</paragraph>
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<paragraph pageId="4" pageNumber="47">Adults with 20 segments (Fig. 1C), body conglobation complete.</paragraph>
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<paragraph pageId="4" pageNumber="47">Antennae short, but slender; in length, antennomere 6 = 3> 2> 4 = 5> 7> 1 (Fig. 9A).</paragraph>
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<paragraph pageId="4" pageNumber="47">Labrum with three teeth (Fig. 8A).</paragraph>
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<caption pageId="4" pageNumber="47">
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<paragraph pageId="4" pageNumber="47">
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Figure 8.
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<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus paraster" order="Polydesmida" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="paraster">Eutrichodesmus paraster</taxonomicName>
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sp. n., SEM, male holotype from Cave Tham Long Puang. A head (H), subfrontal view B collum (Co) and segments 2-3 (m2-m3 = metaterga 2, 3), dorsal view C collum (Co), dorsal view D segments 4 and 5 (m4-m5 = metaterga 4, 5), dorsal view E segment 4 (m4 = metatergum 4), dorsal view F cross-section of segment 6, caudal view G prozonum 5, dorsal view H segments 17-19 and telson, lateral view (m17-m19 = metaterga 17-19) I telson, ventral view.
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</paragraph>
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</caption>
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<paragraph lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
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Head (Fig. 8A), bacilliform sensilla on antennae (Fig. 9A), gnathochilarium (Fig. 9B), mandibles (Fig. 9D), prozona (Fig. 8G), endoterga (Fig. 9J), metatergal setae
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<pageBreakToken pageId="5" pageNumber="48" start="start">(</pageBreakToken>
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Fig. 9E), sterna (Fig. 8F), pleurosternal keels, stigmata (Fig. 9K), legs (Fig. 9H), gonopod aperture, telson (Fig. 8I) and vulvae (Fig. 9I) all similar to those in
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<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus steineri" order="Polydesmida" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="steineri">Eutrichodesmus steineri</taxonomicName>
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sp. n.
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</paragraph>
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<paragraph pageId="5" pageNumber="48">
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Collum subtrapeziform, with six transverse rows of round microvillose tubercles (Fig. 8
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<normalizedToken originalValue="B–C">B-C</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="5" pageNumber="48">Stricture between pro- and metazona broad and shallow, more finely alveolate-microgranulate than prozona (Fig. 8G). Limbus regularly microcrenulate (Fig. 9G).</paragraph>
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<caption pageId="5" pageNumber="48">
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<paragraph pageId="5" pageNumber="48">
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Figure 9.
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<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus paraster" order="Polydesmida" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="paraster">Eutrichodesmus paraster</taxonomicName>
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sp. n., SEM, male holotype from Cave Tham Long Puang. A right antenna, lateral view (a1-a7 = antennomeres 2-7) B gnathochilarium, ventral view (lp = lateral palpus; ip = inner palpus; st = stipites; ll = lamellae linguales; m = mentum) C paratergum 2 (p2), dorsal view D right mandible, general view (et = external tooth; it = internal tooth; pl = pectinate lamellae; ia = intermediate area; mp = molar plate) E a seta (se) on metatergum 18 F paraterga 17 (p17) and 18 (p18), lateral view G limbus of metatergum 5, dorsal view H midbody leg, frontal view (fe = femur; ta = tarsus) I female paratype, vulvae, general view (op = operculum; b = bursa) J endotergum 5 K stigmata (st), ventral view.
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</paragraph>
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</caption>
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<paragraph lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48">
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Front margin of metaterga 2-4 strongly elevated, each latter with three transverse mixostictic rows of similar tubercles (Fig. 8B,
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<normalizedToken originalValue="D–E">D-E</normalizedToken>
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). Following metaterga with three transverse rows of small, flattened, microvillose tuberculations (Figs 8D, 9E). Metaterga 4-19 each with a very high mid-dorsal projection, slightly smaller on metatergum 4 (Fig. 8
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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, H); tip of projections usually bilobed, always bilobed on each side on
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<pageBreakToken pageId="6" pageNumber="49" start="start">metaterga</pageBreakToken>
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5 and 6 (Fig. 8D). Projections upright, directed slightly caudad only on metatergum 19 (Fig. 8H).
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</paragraph>
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<paragraph pageId="6" pageNumber="49">Paraterga 2 strongly enlarged, vaguely 4-lobulated laterally (Fig. 9C). Following paraterga bi- or trilobate laterally in anterior and posterior parts of body, respectively, each with two small caudal lobulations (Figs 1C, 9F).</paragraph>
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<paragraph pageId="6" pageNumber="49">Ozopores absent.</paragraph>
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<paragraph pageId="6" pageNumber="49">Gonopods (Fig. 10) relatively complex. Coxae large, micropapillate and sparsely setose ventrolaterally, with two small apicolateral lobes (cl). Telopodite slightly longer than coxite, slender throughout, setose in basal half, with a prominent, laterally denticulate, distofemoral process (dp) at about midway. Acropodite twisted, subapically with a very small mesal tooth (t) and an evident digitiform lobule (lo) dorsally; seminal groove terminating subapically in a hairpad.</paragraph>
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<caption pageId="6" pageNumber="49">
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<paragraph pageId="6" pageNumber="49">
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Figure 10.
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<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus paraster" order="Polydesmida" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="paraster">Eutrichodesmus paraster</taxonomicName>
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sp. n., male holotype from Cave Tham Long Puang.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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right gonopod, lateral and mesal views, respectively C distofemoral process, subventral view. Abbreviations: cl = coxal lobe; dp = distofemoral process; lo = lobule; t = tooth.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="49" type="remarks">
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<paragraph pageId="6" pageNumber="49">Remarks.</paragraph>
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<paragraph pageId="6" pageNumber="49">
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As this species was collected in a cave, and has a pallid body it appears to be a real troglobite. The absence of ozopores is unique for the family
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<taxonomicName family="Haplodesmidae" lsidName="" pageId="6" pageNumber="49" rank="family">Haplodesmidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |