136 lines
18 KiB
XML
136 lines
18 KiB
XML
<document id="FFE9E36B3D64698DB8C7610B9BFD5CE3" ENCODING="UTF-8" ID-GBIF-Dataset="63f88698-8d89-46c9-ac22-13d0c2840523" ModsDocID="6755" checkinTime="1243377363463" checkinUser="christiana" docAuthor="Brown, W. L." docDate="1978" docId="25B18CFF1E1673CED443F515CFD13CAF" docLanguage="en" docName="6755" docOrigin="Psyche 84" docSource="http://atbi.biosci.ohio-state.edu/HymOnline/reference-full.html?id=6755" docTitle="Pilotrochus besmerus Brown, 1978, new species" docType="treatment" docVersion="10" lastPageNumber="224" masterDocId="D8E50F678A508EF1B9F922C984C419A7" masterDocTitle="An aberrant new genus of myrmicine ant from Madagascar." masterLastPageNumber="224" masterPageNumber="218" pageNumber="221" updateTime="1701312566000" updateUser="plazi">
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<mods:mods id="ABFF08B37FCD31839E3610BFA3109989" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6046CF43CA8E1880A0FE954C1B4790AE">
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<mods:title id="B6E5C371E99C52D677DF4EC60C275DAD">An aberrant new genus of myrmicine ant from Madagascar.</mods:title>
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</mods:titleInfo>
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<mods:name id="B83AF285CBDD19BAAD801CAA7B347BA1" type="personal">
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<mods:role id="9F36D9A474ED9B28F9626870CB4B55F7">
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<mods:roleTerm id="ADDCF3E0B7E9F35792B32333F5AB374A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="118EB2418F1B407B85EE23FC6ECD9AFF">Brown, W. L.</mods:namePart>
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<mods:typeOfResource id="5B1B77D6A2F463069E4CD744EB6C3B5B">text</mods:typeOfResource>
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<mods:titleInfo id="44AB0CBC8F83DA02CB24FABF2A34AE5D">
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<mods:title id="5E414D318938AEE48651E91B538F35A6">Psyche</mods:title>
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</mods:titleInfo>
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<mods:part id="19E94560599C28DFAA495A2253D58F7F">
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<mods:date id="71B253669E28ABD769B6419E91A616E4">1978</mods:date>
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<mods:detail id="86D1176C2DA8F82AFA598EA1671E266E" type="volume">
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<mods:number id="077435C8E5F756022C34474E8FF06718">84</mods:number>
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</mods:detail>
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<mods:extent id="A0EED27BBBF7EADF061CB032ABBE664D" unit="page">
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<mods:start id="505D314538591EC6EAFF151BE564143E">218</mods:start>
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<mods:end id="B85144DE99EF87B02369DC8581F530B4">224</mods:end>
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<mods:location id="3E16FF1E6C934AFCC731CA1DD4B279E0">
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<mods:url id="9593E31D81D54623EB44B7F8D739A5D0">http://atbi.biosci.ohio-state.edu/HymOnline/reference-full.html?id=6755</mods:url>
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</mods:location>
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<mods:classification id="8EFDB198DF0298E0A7F299606FB2CCA2">journal article</mods:classification>
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<mods:identifier id="6DC9072362415C86044D4D7C40B8EE8D" type="HNS-Pub">6755</mods:identifier>
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</mods:mods>
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<treatment id="25B18CFF1E1673CED443F515CFD13CAF" ID-DOI="http://doi.org/10.5281/zenodo.6284205" ID-GBIF-Taxon="100127128" ID-Zenodo-Dep="6284205" LSID="urn:lsid:plazi:treatment:25B18CFF1E1673CED443F515CFD13CAF" httpUri="http://treatment.plazi.org/id/25B18CFF1E1673CED443F515CFD13CAF" lastPageNumber="224" pageNumber="221">
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<subSubSection id="94BEED61BF8F283FA7CDA0B57F22F18F" type="nomenclature">
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<paragraph id="763C3A903B1CA6698113E1A086081DA6" pageNumber="221">
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<taxonomicName id="CC064EF4A8BC2CFF8658F593400245CF" ID-CoL="4HVMF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34152" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus besmerus Brown" lsidName-HNS="Pilotrochus besmerus Brown" order="Hymenoptera" pageNumber="221" phylum="Arthropoda" rank="species" species="besmerus">Pilotrochus besmerus</taxonomicName>
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<taxonomicNameLabel id="3B9AB89D730250A2774C392429A99B92" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="6EE6F311D6AC66F31F2988F03337F3AF" type="description">
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<paragraph id="8D2AA82CA18BC9B12827E6BA7889D907" pageNumber="221">Figs. 1,2.</paragraph>
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<paragraph id="ED7FF4FBA819B0F8B4D264EA595EF3CF" pageNumber="221">Holotype worker: Total length (TL) 2.9, head length (HL) 0.68, head width (HW) 0.60, mandibles extend beyond median clypeal free margin (ML) 0.11 mm; when head is tilted back a little from full-face view, mandibles may extend beyond clypeus as much as 0.18 mm; trunk length (WL) 0.64, scape L (excluding radicle) 0.32, greatest length of eye 0.07 mm. Petiole length (chord of arc) 0.45, hind femur length 0.43, hind tibia length 0.34, hind tarsus length 0.60 mm, of which metatarsus is half.</paragraph>
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<paragraph id="89492989D483EB21C31A7FBA4F5AF0BA" pageNumber="221">Details of form and sculpture are well shown in the figures. As seen in dorsal view, eyes hidden in full-face view, barely visible when head is tilted back slightly, as in fig. 1. Pronotum slightly broader than long (width 0.38 mm), rounded in front, excised behind; mesonotum subquadratic, about 0.20 mm wide, with feebly convex borders on all four sides. Petiolar node 0.17 mm wide, postpetiolar node with rounded sides, slightly wider behind (width 0.21 mm, length 0.20 mm).</paragraph>
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<paragraph id="C13A088F6E08D4ED0B6FCC1840051897" pageNumber="221">Pronotum smooth and shining, with about 10 costulae running longitudinally at different lengths; mesonotum with 5 longitudinal costulae on rugulae. About 5 longitudinal rugules each on petiolar and postpetiolar nodes, but weaker on the postpetiole, so that its disc is primarily smooth and shining like the gaster.</paragraph>
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<caption id="10253F804888CD27222D3FE11752221F">
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<paragraph id="0484F8C152ADCEC91867F5ABDC746649" pageNumber="222">
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<pageStartToken id="6383594AC9CBD6256F5763FF4D19A00F" pageNumber="222">Fig</pageStartToken>
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1.
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<taxonomicName id="025933E2AF0290FD799864A3EEB8BD6B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34152" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus besmerus Brown" lsidName-HNS="Pilotrochus besmerus Brown" order="Hymenoptera" pageNumber="222" phylum="Arthropoda" rank="species" species="besmerus">Pilotrochus besmerus</taxonomicName>
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,
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<taxonomicNameLabel id="EEAEA6D26EE194F260B284B9F98752EC" rank="genus">new genus</taxonomicNameLabel>
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and species, holotype worker, head in dorsal view, tilted back slightly from the full-face position so as to show the mandibular dentition better, X 67 Drawing by Susan Poulakis
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</paragraph>
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</caption>
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<paragraph id="EA6996A77FD7654503A775C294260D4F" pageNumber="222">Hairs mostly a little less than 0.1 mm long on anterior head and scapes, a little more than 0.1 mm on posterior vertex; 0.15 - 0.25 on trunk, petiole, postpetiole and gaster, becoming shorter again at gastric apex; about 0.1 to about 0.3 mm on legs; flagelliform, many with tips reflexed or even looped back.</paragraph>
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</subSubSection>
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<subSubSection id="09762711B8F4783381D24F9B6CF23CFF" type="materials_examined">
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<paragraph id="FC5EB6C3B6DC4C991B500CB0A17F8E57" pageNumber="222">Holotype (Museum of Comparative Zoology at Harvard University) a unique worker taken in a Berlese sample of forest humus and litter from along the road to Anosibe, 33 km south of Mora-manga, in east central Madagascar, 4 - 12 April 1975 (A. Peyrieras).</paragraph>
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</subSubSection>
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<subSubSection id="9B2F9C14C1F1B980601BABFCE80A62DB" type="discussion">
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<paragraph id="156D1CAFCE8FD1254BF14469F7B5D679" lastPageNumber="224" pageNumber="222">
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The relationships of
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<taxonomicName id="CCD58CB157B7269EF3268F18D1BB2955" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24875" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus Brown" lsidName-HNS="Pilotrochus Brown" order="Hymenoptera" pageNumber="222" phylum="Arthropoda" rank="genus">Pilotrochus</taxonomicName>
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are obscure. The shape of the head, with its broad and deep scrobes and small ventrolateral eyes, recalls that of
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<taxonomicName id="C9DAC67A02023BE72EB6221F2D076F2D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24909" class="Insecta" family="Formicidae" genus="Dacetinops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dacetinops Brown & Wilson" lsidName-HNS="Dacetinops Brown & Wilson" order="Hymenoptera" pageNumber="222" phylum="Arthropoda" rank="genus">Dacetinops</taxonomicName>
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, or the
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<taxonomicName id="55DF65DB0606373BD4394E18BBBD685A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2439" class="Insecta" family="Formicidae" genus="Codiomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Codiomyrmex Wheeler" lsidName-HNS="Codiomyrmex Wheeler" order="Hymenoptera" pageNumber="222" phylum="Arthropoda" rank="genus">Codiomyrmex</taxonomicName>
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group of
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<taxonomicName id="F4B2C1614673C5FEB1C63F7562AE02ED" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2280" lsidName-HNS="Dacetini Forel" pageNumber="222" rank="tribe">Dacetini</taxonomicName>
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, or even
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<taxonomicName id="D2A7EE33EE2C05FA59E99CFFAF18831E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2372" class="Insecta" family="Formicidae" genus="Tatuidris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tatuidris Brown & Kempf" lsidName-HNS="Tatuidris Brown & Kempf" order="Hymenoptera" pageNumber="222" phylum="Arthropoda" rank="genus">Tatuidris</taxonomicName>
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, but
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<taxonomicName id="A99AD75CE77B7CC57F19A7E279730D97" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24875" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus Brown" lsidName-HNS="Pilotrochus Brown" order="Hymenoptera" pageNumber="222" phylum="Arthropoda" rank="genus">Pilotrochus</taxonomicName>
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differs strongly from all of these in its 8 - segmented antennae, in the form of its mandibles and their teeth, and in the form of its trunk and petiole. The lateral " hairwheel " organs are also striking, and so far as I am aware are unmatched among the ants, although the region of the ventral furrow between pro- and mesothorax is sometimes modified and apparently glandular in some dacetines and a few other myrmicines.
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</paragraph>
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<caption id="82A00DED6F5B51662157E5FCB98B2709">
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<paragraph id="8C645D5B7EBCBACD37FAE934FF6ECC79" pageNumber="223">
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Fig. 2.
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<taxonomicName id="78B1F2E9C4455DF9D339E3014067D185" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34152" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus besmerus Brown" lsidName-HNS="Pilotrochus besmerus Brown" order="Hymenoptera" pageNumber="223" phylum="Arthropoda" rank="species" species="besmerus">Pilotrochus besmerus</taxonomicName>
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,
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<taxonomicNameLabel id="32442FD21432055913CC62F666641DCB" rank="genus">new genus</taxonomicNameLabel>
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and species, holotype worker in side view, X 67. Drawing by Susan Poulakis.
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</paragraph>
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</caption>
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<paragraph id="69B9BF70EBE9A3E1325C108E9E830DCE" pageNumber="224">
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The 2 - segmented antennal club, together with some points of habitus (especially coarse sculpture and long, flexuous pilosity, plus the antennal scrobes), recalls the neotropical genus
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<taxonomicName id="7BDCAE870DBFD481E055C28E53793AEC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24912" class="Insecta" family="Formicidae" genus="Lachnomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachnomyrmex Wheeler" lsidName-HNS="Lachnomyrmex Wheeler" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Lachnomyrmex</taxonomicName>
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, but the shape of the trunk in
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<taxonomicName id="FA56BAA382F13F740361560709ED9D1C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24875" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus Brown" lsidName-HNS="Pilotrochus Brown" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Pilotrochus</taxonomicName>
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is completely different, the propodeal teeth so prominent in
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<taxonomicName id="C765A9626A67381DF509DD20CEC30FAB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24912" class="Insecta" family="Formicidae" genus="Lachnomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lachnomyrmex Wheeler" lsidName-HNS="Lachnomyrmex Wheeler" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Lachnomyrmex</taxonomicName>
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are completely absent, and the mandibles in the two genera are very different.
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</paragraph>
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<paragraph id="48AD68E5949F92B90E8457238CAA4210" pageNumber="224">
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At the moment, all one can say without seeing the winged forms and larvae of
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<taxonomicName id="FE2DAC265816DE17809E6422FE99FB00" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24875" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus Brown" lsidName-HNS="Pilotrochus Brown" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Pilotrochus</taxonomicName>
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, and without knowing something of its lifeway, is that it is a member of subfamily
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<taxonomicName id="075D9C239212BA543A4761DD7731950B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2258" lsidName-HNS="Myrmicinae Lepeletier" pageNumber="224" rank="subFamily">Myrmicinae</taxonomicName>
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, but one not belonging to any of the well-circumscribed " higher " tribes (
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<taxonomicName id="676E56C2B4D7959CCEC6CAE57DFA1DC4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2278" lsidName-HNS="Crematogastrini Forel" pageNumber="224" rank="tribe">Crematogastrini</taxonomicName>
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,
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<taxonomicName id="19AE347A043B57EB8C8EF10E838FC060" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2280" lsidName-HNS="Dacetini Forel" pageNumber="224" rank="tribe">Dacetini</taxonomicName>
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,
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<taxonomicName id="470F0DC1619CC6D735B66C1B2DA764E3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2270" lsidName-HNS="Basicerotini Brown" pageNumber="224" rank="tribe">Basicerotini</taxonomicName>
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,
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<taxonomicName id="2DD4F9444841AA580B1DD5B9FC05ACCC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2274" lsidName-HNS="Cataulacini Emery" pageNumber="224" rank="tribe">Cataulacini</taxonomicName>
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,
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<taxonomicName id="8C613BD5DB1AFB9E1B26ABAE474AB648" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2269" lsidName-HNS="Attini Smith" pageNumber="224" rank="tribe">Attini</taxonomicName>
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, etc.). Thus, it falls among the mass of generic complexes related to
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<taxonomicName id="A02F632D24E3A6DB05227160DAAD4EE5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24868" class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmica Latreille" lsidName-HNS="Myrmica Latreille" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Myrmica</taxonomicName>
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,
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<taxonomicName id="09E3C468EBB0A926A256698DA9957ED1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole Westwood" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
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,
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<taxonomicName id="278ED746B8D8934C45AF8155621A9590" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24818" class="Insecta" family="Formicidae" genus="Myrmecina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmecina Curtis" lsidName-HNS="Myrmecina Curtis" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Myrmecina</taxonomicName>
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,
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<taxonomicName id="B6062734F647BAB6830D4580DD652144" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24915" class="Insecta" family="Formicidae" genus="Rogeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rogeria Emery" lsidName-HNS="Rogeria Emery" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Rogeria</taxonomicName>
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, etc., among which tribal boundaries are impossible to define for the present, or at least are in dispute.
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</paragraph>
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<paragraph id="CA71D3154233A14AAF41FB0BAEFA796E" pageNumber="224">
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Considering the revisionary work that must be done before we have a rational tribal classification of
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<taxonomicName id="74E9D4D05A345D56899E33FA53E2C63A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2258" lsidName-HNS="Myrmicinae Lepeletier" pageNumber="224" rank="subFamily">Myrmicinae</taxonomicName>
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, it seems to me that nothing would be gained by erecting a new tribe for
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<taxonomicName id="9D4A89A53EC20A75C03F796AC2B9AD4F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24875" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus Brown" lsidName-HNS="Pilotrochus Brown" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="genus">Pilotrochus</taxonomicName>
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, even though for the time being I am able to fit it comfortably into any existing myrmicine tribe.
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</paragraph>
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</subSubSection>
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<subSubSection id="F135DA40C31274F57A754DDA37FCEE3A" type="etymology">
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<paragraph id="C9C3719F65762D958080BB2187246FA9" pageNumber="224">
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The generic name is derived from the Greek ' pilos' (hair) + ' tro-chos' (wheel), while the specific name
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<taxonomicName id="D97DD9979FC9DFE53F5275975AD41E03" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34152" class="Insecta" family="Formicidae" genus="Pilotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilotrochus besmerus Brown" lsidName-HNS="Pilotrochus besmerus Brown" order="Hymenoptera" pageNumber="224" phylum="Arthropoda" rank="species" species="besmerus">besmerus</taxonomicName>
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combines the Latin ' bes' (eight of twelve) with the Greek ' meros' (part), in reference to the 8 - segmented antennae as compared to the primitive myrmicine (and formicid) 12 - merous condition.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |