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<document id="13CC83F003E773DA9EE6B8BDC5DE6A7F" ID-ISSN="0253-6730" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649690268132" checkinUser="carolina" docAuthor="Sassaroli, Stefano &amp; Venturi, Federico" docDate="2012" docId="03DA011AFFDEFFE30A99FDE9FEC208FE" docLanguage="en" docName="RevPaleobiol.31.1.85-114.pdf" docOrigin="Revue de Paléobiologie 31 (1)" docStyle="DocumentStyle:C172DF54475A4E3C75B7590733AD2922.3:RevPaleobiol.2012-.journal_article.1cover" docStyleId="C172DF54475A4E3C75B7590733AD2922" docStyleName="RevPaleobiol.2012-.journal_article.1cover" docStyleVersion="3" docTitle="Hildaites wrighti SPATH 1913" docType="treatment" docVersion="2" lastPageNumber="96" masterDocId="FFE37962FFD5FFEF0953FF85FFA50C7C" masterDocTitle="Early Toarcian (post-OAE) Hildoceratinae (Ammonitina) fauna from the Marchean Apennines (Italy)" masterLastPageNumber="114" masterPageNumber="85" pageNumber="95" updateTime="1699312896216" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="6AA657A45800A13991195EA5DACB87D8">Early Toarcian (post-OAE) Hildoceratinae (Ammonitina) fauna from the Marchean Apennines (Italy)</mods:title>
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<mods:namePart id="CD13FEA01633E9143C80FC2C30754A49">Sassaroli, Stefano</mods:namePart>
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<treatment id="03DA011AFFDEFFE30A99FDE9FEC208FE" LSID="urn:lsid:plazi:treatment:03DA011AFFDEFFE30A99FDE9FEC208FE" httpUri="http://treatment.plazi.org/id/03DA011AFFDEFFE30A99FDE9FEC208FE" lastPageId="12" lastPageNumber="96" pageId="11" pageNumber="95">
<subSubSection id="C369E387FFDEFFE40A99FDE9FB5A0ED8" pageId="11" pageNumber="95" type="nomenclature">
<paragraph id="8BCCB00CFFDEFFE40A99FDE9FAB70EF8" blockId="11.[970,1298,620,676]" box="[970,1298,620,644]" pageId="11" pageNumber="95">
<heading id="D0840760FFDEFFE40A99FDE9FAB70EF8" box="[970,1298,620,644]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="95" reason="4">
<taxonomicName id="4C73CB8FFFDEFFE40A99FDE9FAB70EF8" ID-CoL="3LWWB" authority="SPATH, 1913" authorityName="SPATH" authorityYear="1913" box="[970,1298,620,644]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFDEFFE40A99FDE9FAB70EF8" bold="true" box="[970,1298,620,644]" pageId="11" pageNumber="95">
<emphasis id="B9076C1EFFDEFFE40A99FDE9FB270EF8" bold="true" box="[970,1154,620,644]" italics="true" pageId="11" pageNumber="95">Hildaites wrighti</emphasis>
SPATH, 1913
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A8EFD09FB5A0ED8" blockId="11.[970,1298,620,676]" box="[989,1279,652,676]" pageId="11" pageNumber="95">
<emphasis id="B9076C1EFFDEFFE40A8EFD09FB5A0ED8" bold="true" box="[989,1279,652,676]" pageId="11" pageNumber="95">
Pl. I, fig. 1a-b;
<figureCitation id="1348AC89FFDEFFE40DD6FD09FB630ED8" box="[1157,1222,652,676]" captionStart="Fig" captionStartId="13.[151,186,1402,1424]" captionTargetBox="[151,1436,257,1349]" captionTargetId="figure-321@13.[151,1436,257,1359]" captionTargetPageId="13" captionText="Fig. 6: A-B: Hildaites wrighti, specimen MSA60, whorl section (x1) and suture (x5). C-D: Hildaites subserpentinus, specimen MSA254, whorl section (x1) and suture (x3). E-F: Hildaites striatus, specimen MSA30, whorl section (x1) and suture (x3). G-I: Hildaites evolutus n. sp., holotype MSA426, shell draw (x1), whorl section (x1) and suture (x3). J-L: Hildaites exilis, specimen MSA102, shell draw (x1), whorl section (x1) and suture (x3)." pageId="11" pageNumber="95">Fig. 6</figureCitation>
, A-B
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C369E387FFDEFFE40A6CFD4EFAC9082A" pageId="11" pageNumber="95" type="description">
<paragraph id="8BCCB00CFFDEFFE40A6CFD4EFC690E81" blockId="11.[831,1437,715,1112]" pageId="11" pageNumber="95">
?1870.
<emphasis id="B9076C1EFFDEFFE40AC6FD4EFBF70E9C" box="[917,1106,715,736]" italics="true" pageId="11" pageNumber="95">Ammonites radians</emphasis>
REINECKE. - MENEGHINI, pl. 11, fig. 7.
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A6CFC80FBDC0F4B" blockId="11.[831,1437,715,1112]" pageId="11" pageNumber="95">
1884.
<taxonomicName id="4C73CB8FFFDEFFE40AC6FC80FAB50F66" authority="(D' ORBIGNY)" baseAuthorityName="D' ORBIGNY" box="[917,1296,773,794]" class="Cephalopoda" family="Hildoceratidae" genus="Harpoceras" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="normaniaum">
<emphasis id="B9076C1EFFDEFFE40AC6FC80FB2D0F66" box="[917,1160,773,794]" italics="true" pageId="11" pageNumber="95">Harpoceras normaniaum</emphasis>
(D ORBIGNY)
</taxonomicName>
. - WRIGHT, pl. 83, figs. 1-2 (
<typeStatus id="54C80EAEFFDEFFE40D47FCA4FBCB0F4B" box="[1044,1134,801,823]" pageId="11" pageNumber="95" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A6CFCBBFC5D0F0D" blockId="11.[831,1437,715,1112]" pageId="11" pageNumber="95">
1913.
<taxonomicName id="4C73CB8FFFDEFFE40AC6FCBAFB450F2F" authority="SPATH" authorityName="SPATH" box="[917,1248,830,852]" class="Cephalopoda" family="Hildoceratidae" genus="Protogrammoceras" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFDEFFE40AC6FCBAFB390F2F" box="[917,1180,830,852]" italics="true" pageId="11" pageNumber="95">Protogrammoceras wrighti</emphasis>
SPATH
</taxonomicName>
, p. 553 (
<emphasis id="B9076C1EFFDEFFE40C6FFCBAFA320F28" box="[1340,1431,831,852]" italics="true" pageId="11" pageNumber="95">nom. nov</emphasis>
.
<typeStatus id="54C80EAEFFDEFFE40AC6FCDEFC480F0D" box="[917,1005,859,881]" pageId="11" pageNumber="95" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A6CFCFDFA8B0FF2" blockId="11.[831,1437,715,1112]" box="[831,1326,888,910]" pageId="11" pageNumber="95">
?1969.
<taxonomicName id="4C73CB8FFFDEFFE40AC6FCFDFBD70FF1" box="[917,1138,888,909]" class="Cephalopoda" family="Hildoceratidae" genus="Protogrammoceras" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B9076C1EFFDEFFE40AC6FCFDFBEA0FF1" box="[917,1103,888,909]" italics="true" pageId="11" pageNumber="95">Protogrammoceras</emphasis>
sp.
</taxonomicName>
, PINNA, pl. 4, fig.6.
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A6CFC10FC1E0FBB" blockId="11.[831,1437,715,1112]" pageId="11" pageNumber="95">
?1980.
<taxonomicName id="4C73CB8FFFDEFFE40AC6FC11FBDA0FD6" authority="GUEX." authorityName="GUEX." box="[917,1151,916,938]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="striatus">
<emphasis id="B9076C1EFFDEFFE40AC6FC11FB9E0FD6" box="[917,1083,916,938]" italics="true" pageId="11" pageNumber="95">Hildaites striatus</emphasis>
GUEX.
</taxonomicName>
- WIEDENMAYER, pl. 32, figs. 1-5.
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A6CFC4BFBA5087D" blockId="11.[831,1437,715,1112]" pageId="11" pageNumber="95">
1992.
<taxonomicName id="4C73CB8FFFDEFFE40AC6FC4BFB240F9F" authority="SPATH." authorityName="SPATH." box="[917,1153,974,995]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFDEFFE40AC6FC4BFB9D0F9F" box="[917,1080,974,995]" italics="true" pageId="11" pageNumber="95">Hildaites wrighti</emphasis>
SPATH.
</taxonomicName>
- HOWARTH, pl.30, figs. 7a-b (
<typeStatus id="54C80EAEFFDEFFE40ACFFC6EFC53087D" box="[924,1014,1003,1025]" pageId="11" pageNumber="95" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A6CFB8DFA2B0861" blockId="11.[831,1437,715,1112]" box="[831,1422,1032,1054]" pageId="11" pageNumber="95">
?2001.
<taxonomicName id="4C73CB8FFFDEFFE40AC6FB8DFB250861" authority="SPATH." authorityName="SPATH." box="[917,1152,1032,1053]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFDEFFE40AC6FB8DFB9D0861" box="[917,1080,1032,1053]" italics="true" pageId="11" pageNumber="95">Hildaites wrighti</emphasis>
SPATH.
</taxonomicName>
- MACCHIONI, pl. 6, fig. 14.
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A6CFBA0FAF80846" blockId="11.[831,1437,715,1112]" box="[831,1373,1060,1082]" pageId="11" pageNumber="95">
?2002a.
<taxonomicName id="4C73CB8FFFDEFFE40AC6FBA1FB250846" authority="SPATH." authorityName="SPATH." box="[917,1152,1060,1082]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFDEFFE40AC6FBA1FB9D0845" box="[917,1080,1060,1081]" italics="true" pageId="11" pageNumber="95">Hildaites wrighti</emphasis>
SPATH.
</taxonomicName>
- MACCHIONI, fig. 4.9.
</paragraph>
<paragraph id="8BCCB00CFFDEFFE40A6CFBC4FAC9082A" blockId="11.[831,1437,715,1112]" box="[831,1388,1089,1112]" pageId="11" pageNumber="95">
?2006.
<taxonomicName id="4C73CB8FFFDEFFE40AC6FBC4FB25082B" authority="SPATH." authorityName="SPATH." box="[917,1152,1089,1111]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="11" pageNumber="95" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFDEFFE40AC6FBC4FB9D082A" box="[917,1080,1089,1110]" italics="true" pageId="11" pageNumber="95">Hildaites wrighti</emphasis>
SPATH.
</taxonomicName>
- BÉCAUD, pl. 32, fig. 1.
</paragraph>
</subSubSection>
<subSubSection id="C369E387FFDEFFE40A6CFBFAFC2E0884" pageId="11" pageNumber="95" type="materials_examined">
<paragraph id="8BCCB00CFFDEFFE40A6CFBFAFC2E0884" blockId="11.[831,1438,1151,2071]" pageId="11" pageNumber="95">
<emphasis id="B9076C1EFFDEFFE40A6CFBFAFC0908EB" bold="true" box="[831,940,1151,1175]" pageId="11" pageNumber="95">Material:</emphasis>
Only one medium sized specimen
<collectionCode id="ED6228C9FFDEFFE40C1BFBFAFA2708EB" box="[1352,1410,1151,1175]" country="Ireland" httpUri="http://grbio.org/cool/vvha-6rck" name="Museum of Science and Art" pageId="11" pageNumber="95">MSA</collectionCode>
60 with the body chamber partly preserved and the shell strongly encrusted by pyrite, coming from Marconessa quarry.
</paragraph>
</subSubSection>
<subSubSection id="C369E387FFDEFFE40A6CFB7AFACB098B" pageId="11" pageNumber="95" type="diagnosis">
<paragraph id="8BCCB00CFFDEFFE40A6CFB7AFACB098B" blockId="11.[831,1438,1151,2071]" pageId="11" pageNumber="95">
<emphasis id="B9076C1EFFDEFFE40A6CFB7AFC13096B" bold="true" box="[831,950,1279,1303]" pageId="11" pageNumber="95">Diagnosis:</emphasis>
Moderately evolute and compressed shell, almost elliptical whorl section, narrow and gently bisulcate or tabulate keeled venter, high almost vertical umbilical wall in the outer whorls and rounded umbilical edge. The venter is gently bisulcate with a strong and raised keel. The fine, dense and gently sigmoidal ribs start gently backward from the umbilical wall and they end moderately projected on the ventro-lateral edge.
</paragraph>
</subSubSection>
<subSubSection id="C369E387FFDEFFE40A6CFA7AFA9A0BCB" pageId="11" pageNumber="95" type="description">
<paragraph id="8BCCB00CFFDEFFE40A6CFA7AFA9A0BCB" blockId="11.[831,1438,1151,2071]" pageId="11" pageNumber="95">
<emphasis id="B9076C1EFFDEFFE40A6CFA7AFC680A6B" bold="true" box="[831,973,1535,1559]" pageId="11" pageNumber="95">Description:</emphasis>
Compared to the
<typeStatus id="54C80EAEFFDEFFE40DFBFA7AFAA00A6B" box="[1192,1285,1535,1559]" pageId="11" pageNumber="95" type="holotype">holotype</typeStatus>
refigured by HOWARTH (1992, pl. 30, figs. 7a-7b) our specimen seems to have a more compressed whorl section and lesser falcate ribs. HOWARTH (1992) states that the
<typeStatus id="54C80EAEFFDEFFE40C45F9DAFAD60A0B" box="[1302,1395,1631,1655]" pageId="11" pageNumber="95" type="holotype">holotype</typeStatus>
has «
<emphasis id="B9076C1EFFDEFFE40A1EF905FB4E0AEB" box="[845,1259,1663,1687]" italics="true" pageId="11" pageNumber="95">a narrow tricarinate-bisulcate venter</emphasis>
», but we think that this statement overemphasizes the same HOWARTH s figure (7b), which like our specimen really shows the weak sulci jointed with a strong and raised keel, which also shows a narrow and gently bisulcate venter in the fragmocone and a narrow and flattened venter in the body chamber. HOWARTH has not provided a draw or description of the suture line of SPATH s
<typeStatus id="54C80EAEFFDEFFE40C56F8DAFAC30B0B" box="[1285,1382,1887,1911]" pageId="11" pageNumber="95" type="holotype">holotype</typeStatus>
. Our specimen has a simple not jagged suture, with a wide L lobe and moderately developed umbilical lobes.
</paragraph>
</subSubSection>
<subSubSection id="C369E387FFDEFFE30A13F83AFEC208FE" lastPageId="12" lastPageNumber="96" pageId="11" pageNumber="95" type="discussion">
<paragraph id="8BCCB00CFFDEFFE30A13F83AFE7D0F9C" blockId="11.[831,1438,1151,2071]" lastBlockId="12.[151,757,257,1155]" lastPageId="12" lastPageNumber="96" pageId="11" pageNumber="95">
<emphasis id="B9076C1EFFDEFFE40A13F83AFAB90BAB" bold="true" box="[832,1308,1983,2007]" pageId="11" pageNumber="95">Discussion and comparative diagnosis:</emphasis>
HOWARTH (1992) inferred that WRIGHT s
<typeStatus id="54C80EAEFFDEFFE40DF3F85AFAA50B8B" box="[1184,1280,2015,2039]" pageId="11" pageNumber="95" type="holotype">holotype</typeStatus>
, which is the only sample collected in Britain, comes from
<collectingRegion id="49B77EEEFFDEFFE40C6AF87AFA39046B" box="[1337,1436,2047,2071]" country="United Kingdom" name="Somerset" pageId="11" pageNumber="95">Somerset</collectingRegion>
(Ilmister, Barrington) and probably it was found in the Fish and Insect Bed, which is Exaratum subzone, Falciferum zone in age. Contrary to HOWARTH s (1992) opinion we think that
<taxonomicName id="4C73CB8FFFD9FFE308C0FEE6FDA00D06" box="[403,517,354,378]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFD9FFE308C0FEE6FDA00D06" box="[403,517,354,378]" italics="true" pageId="12" pageNumber="96">H. wrighti</emphasis>
</taxonomicName>
is not a synonym of
<taxonomicName id="4C73CB8FFFD9FFE309C4FE06FE450DE7" authority="GUEX, 1973" authorityName="GUEX" authorityYear="1973" box="[151,480,387,411]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="striatus">
<emphasis id="B9076C1EFFD9FFE309C4FE06FEF40DE6" box="[151,337,387,411]" italics="true" pageId="12" pageNumber="96">Hildaites striatus</emphasis>
GUEX, 1973
</taxonomicName>
, because the latter shows lesser sigmoidal, finer and denser ribs, bundled or bifurcating near the umbilical wall with typical streaking appearance as well as a narrower and not sulcated venter with a thin keel. On the other side,
<taxonomicName id="4C73CB8FFFD9FFE30B70FD80FD360E60" box="[547,659,516,540]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFD9FFE30B70FD80FD930E60" box="[547,566,517,540]" italics="true" pageId="12" pageNumber="96">H</emphasis>
.
<emphasis id="B9076C1EFFD9FFE30B15FD81FD360E60" box="[582,659,516,540]" italics="true" pageId="12" pageNumber="96">wrighti</emphasis>
</taxonomicName>
shows a bisulcate venter with strong keel and usually single ribs. Besides,
<taxonomicName id="4C73CB8FFFD9FFE309AAFDC3FECE0E21" box="[249,363,581,605]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="striatus">
<emphasis id="B9076C1EFFD9FFE309AAFDC3FEA90E21" box="[249,268,582,605]" italics="true" pageId="12" pageNumber="96">H</emphasis>
.
<emphasis id="B9076C1EFFD9FFE30848FDC3FECE0E21" box="[283,363,582,605]" italics="true" pageId="12" pageNumber="96">striatus</emphasis>
</taxonomicName>
has a more compressed and evolute shell than
<taxonomicName id="4C73CB8FFFD9FFE3085AFDE3FEDC0E01" box="[265,377,613,637]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFD9FFE3085AFDE3FEDC0E01" box="[265,377,613,637]" italics="true" pageId="12" pageNumber="96">H. wrighti</emphasis>
</taxonomicName>
, whose umbilical wall is high and rounded, not sloped and flattened as that of
<taxonomicName id="4C73CB8FFFD9FFE30BCEFD03FD480EE1" box="[669,749,646,669]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="striatus">
<emphasis id="B9076C1EFFD9FFE30BCEFD03FD480EE1" box="[669,749,646,669]" italics="true" pageId="12" pageNumber="96">striatus</emphasis>
</taxonomicName>
. Consequently, we cannot share HOWARTH s opinion. The Italian specimens figured by MENEGHINI (1870, pl. 11, fig. 7), ZANZUCCHI (1963, pl. 18, figs. 1-6) and PINNA (1969, pl. 1, fig. 16) have an uncertain taxonomy, as well as the little and badly preserved small sized fragment of the GEMELLARO s collection, recently revisioned by MACCHIONI (2002b, p. 147). However, they have not stratigraphical recording. We also doubt whether to ascribe the specimens figured by WIEDENMAYER (1980, pl. 32, figs. 3-5) to
<taxonomicName id="4C73CB8FFFD9FFE30837FC4FFE740F9D" box="[356,465,969,993]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="wrighti">
<emphasis id="B9076C1EFFD9FFE30837FC4FFED20F9D" box="[356,375,970,993]" italics="true" pageId="12" pageNumber="96">H</emphasis>
.
<emphasis id="B9076C1EFFD9FFE308D7FC4CFE740F9D" box="[388,465,969,993]" italics="true" pageId="12" pageNumber="96">wrighti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCCB00CFFD9FFE309C4FC6CFEC208FE" blockId="12.[151,757,257,1155]" pageId="12" pageNumber="96">
<emphasis id="B9076C1EFFD9FFE309C4FC6CFE80087D" bold="true" box="[151,293,1001,1025]" pageId="12" pageNumber="96">Occurrence:</emphasis>
HOWARTH (1992) believes that the
<typeStatus id="54C80EAEFFD9FFE30BC4FC6CFD51087D" box="[663,756,1001,1025]" pageId="12" pageNumber="96" type="holotype">holotype</typeStatus>
occurs in the
<taxonomicName id="4C73CB8FFFD9FFE30872FB8FFE2F085D" baseAuthorityName="Young &amp; Bird" baseAuthorityYear="1828" box="[289,394,1034,1057]" class="Cephalopoda" family="Hildoceratidae" genus="Cleviceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="exaratum">
<emphasis id="B9076C1EFFD9FFE30872FB8FFE2F085D" box="[289,394,1034,1057]" italics="true" pageId="12" pageNumber="96">Exaratum</emphasis>
</taxonomicName>
subzone (lower
<taxonomicName id="4C73CB8FFFD9FFE30B65FB8FFD08085E" baseAuthorityName="SOWERBY" baseAuthorityYear="1820" box="[566,685,1034,1058]" class="Cephalopoda" family="Hildoceratidae" genus="Harpoceras" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="falciferum">
<emphasis id="B9076C1EFFD9FFE30B65FB8FFD08085E" box="[566,685,1034,1058]" italics="true" pageId="12" pageNumber="96">Falciferum</emphasis>
</taxonomicName>
zone), thus it should be the earliest occurrence of
<taxonomicName id="4C73CB8FFFD9FFE30B27FBAFFD73083E" authorityName="BUCKMAN" authorityYear="1921" box="[628,726,1066,1090]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="genus">
<emphasis id="B9076C1EFFD9FFE30B27FBAFFD73083E" box="[628,726,1066,1090]" italics="true" pageId="12" pageNumber="96">Hildaites</emphasis>
</taxonomicName>
in Britain. However, the Apennines specimen occurs in the early
<taxonomicName id="4C73CB8FFFD9FFE30980FBEEFE8308FE" box="[211,294,1131,1154]" class="Cephalopoda" family="Hildoceratidae" genus="Hildaites" kingdom="Animalia" order="Ammonoidea" pageId="12" pageNumber="96" phylum="Mollusca" rank="species" species="striatus">
<emphasis id="B9076C1EFFD9FFE30980FBEEFE8308FE" box="[211,294,1131,1154]" italics="true" pageId="12" pageNumber="96">Striatus</emphasis>
</taxonomicName>
zone.
</paragraph>
</subSubSection>
</treatment>
</document>