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<taxonomicName id="4C7E3589FF1EFF38FF48FE48330DFACE" authority="(Latreille, 1813)" baseAuthorityName="Latreille" baseAuthorityYear="1813" box="[192,625,1318,1342]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
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<emphasis id="B90A9218FF1EFF38FF48FE4830EDFACE" bold="true" box="[192,401,1318,1342]" italics="true" pageId="213" pageNumber="212">Tomarus fossor</emphasis>
(
<bibRefCitation id="EFEF33FBFF1EFF38FE2AFE483315FACE" author="Latreille" box="[418,617,1318,1342]" firstAuthor="Latreille" pageId="213" pageNumber="212" pagination="1813 - 1817" refId="ref212262" refString="Latreille 1813&gt; Latreille 1833 (1813 - 1817)" type="book chapter" year="1813">Latreille, 1813</bibRefCitation>
)
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<treatmentCitation id="0ADF681BFF1EFF38FF60FE223304FA92" author="Latreille" box="[232,632,1356,1379]" page="11" pageId="213" pageNumber="212" year="1813">
<taxonomicName id="4C7E3589FF1EFF38FF60FE223304FA92" ID-CoL="4V239" authority="Latreille 1813: 11" authorityName="Latreille" authorityPageNumber="11" authorityYear="1813" box="[232,632,1356,1379]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FF60FE2230CCFA91" box="[232,432,1356,1377]" italics="true" pageId="213" pageNumber="212">Scarabaeus fossor</emphasis>
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(equinoctial America)
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<emphasis id="B90A9218FF1EFF38FF48FE1D332CFA7B" bold="true" box="[192,592,1395,1419]" pageId="213" pageNumber="212">
Distribution.
<collectingCountry id="F3690E9AFF1EFF38FEF1FE1D3336FA7B" box="[377,586,1395,1419]" name="French Guiana" pageId="213" pageNumber="212">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FDDEFE1D3216FA7B" author="Blackwelder, R. E." box="[598,874,1395,1419]" journalOrPublisher="United States National Museum Bulletin" pageId="213" pageNumber="212" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FC13FE1D353AFA7B" box="[923,1094,1395,1419]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FC13FE1D353AFA7B" box="[923,1094,1395,1419]" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
</taxonomicName>
).
<emphasis id="B90A9218FF1EFF38FBD5FE1D35C4FA7B" bold="true" box="[1117,1208,1395,1419]" pageId="213" pageNumber="212">
<collectingCountry id="F3690E9AFF1EFF38FBD5FE1D35CFFA7B" box="[1117,1203,1395,1419]" name="Brazil" pageId="213" pageNumber="212">Brazil</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FB37FE1D3190FA5B" author="Blackwelder, R. E." journalOrPublisher="United States National Museum Bulletin" pageId="213" pageNumber="212" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FE93FEFD30BAFA5B" box="[283,454,1427,1451]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FE93FEFD30BAFA5B" box="[283,454,1427,1451]" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FE53FEFD33CBFA5B" author="Endrodi, S." box="[475,695,1427,1451]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="213" pageNumber="212" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 54</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FD6FFEFD357AFA5B" box="[743,1030,1427,1451]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor" subGenus="Ligyrus">
<emphasis id="B90A9218FF1EFF38FD6FFEFD3239FA5A" box="[743,837,1427,1450]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FCDBFEFD32CDFA5A" box="[851,945,1427,1450]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FC37FEFD357AFA5B" box="[959,1030,1427,1451]" italics="true" pageId="213" pageNumber="212">fossor</emphasis>
</taxonomicName>
), 1985a: 255 (as
<taxonomicName id="4C7E3589FF1EFF38FB45FEFD307BFA3B" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor" subGenus="Ligyrus">
<emphasis id="B90A9218FF1EFF38FB45FEFD3457FA5A" box="[1229,1323,1427,1450]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FAB1FEFD34EBFA5A" box="[1337,1431,1427,1450]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FF48FEDD307BFA3B" box="[192,263,1459,1483]" italics="true" pageId="213" pageNumber="212">fossor</emphasis>
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);
<bibRefCitation id="EFEF33FBFF1EFF38FE95FEDD3082FA3B" author="Ratcliffe" box="[285,510,1459,1483]" firstAuthor="Ratcliffe" pageId="213" pageNumber="212" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 276</bibRefCitation>
(?);
<bibRefCitation id="EFEF33FBFF1EFF38FDA6FEDD32FFFA3B" author="Escalona, H. E. &amp; L. J. Joly" box="[558,899,1459,1483]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="213" pageNumber="212" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 118</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FC3BFEDD3522FA3B" box="[947,1118,1459,1483]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FC3BFEDD3522FA3B" box="[947,1118,1459,1483]" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
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);
<bibRefCitation id="EFEF33FBFF1EFF38FBFBFEDD3193FA1B" author="Lopez-Garcia" etAl="et al." firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="451 - 492" refId="ref212996" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, and G. Amat-Garcia. 2015. The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa 4048 (4): 451 - 492." type="journal article" year="2015">López-García et al. 2015: 482</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF1EFF38FF70FEBD3013FA1B" author="Lopez-Garcia" box="[248,367,1491,1515]" firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">2016: 502</bibRefCitation>
.
<emphasis id="B90A9218FF1EFF38FEF0FEBD332CFA1B" bold="true" box="[376,592,1491,1515]" pageId="213" pageNumber="212">
<collectingCountry id="F3690E9AFF1EFF38FEF0FEBD3379FA1B" box="[376,517,1491,1515]" name="Venezuela" pageId="213" pageNumber="212">Venezuela</collectingCountry>
(EO):
</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FDDDFEBD3266FA1B" author="Arrow, G. J." bookContentInfo="124 p." box="[597,794,1491,1515]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="213" pageNumber="212" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 37</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FCC0FEBD328CFA1B" box="[840,1008,1491,1515]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FCC0FEBD328CFA1B" box="[840,1008,1491,1515]" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FB8DFEBD3469FA1B" author="Blackwelder, R. E." box="[1029,1301,1491,1515]" journalOrPublisher="United States National Museum Bulletin" pageId="213" pageNumber="212" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FACBFEBD307BF9FB" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FACBFEBD307BF9FB" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FE97FE9D337CF9FB" author="Endrodi, S." box="[287,512,1523,1547]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="213" pageNumber="212" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 54</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FDBDFE9D3225F9FB" box="[565,857,1523,1547]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor" subGenus="Ligyrus">
<emphasis id="B90A9218FF1EFF38FDBDFE9D33EFF9FA" box="[565,659,1523,1546]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FD2BFE9D327DF9FA" box="[675,769,1523,1546]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FC9AFE9D3225F9FB" box="[786,857,1523,1547]" italics="true" pageId="213" pageNumber="212">fossor</emphasis>
</taxonomicName>
) (EO), 1985a: 255 (as
<taxonomicName id="4C7E3589FF1EFF38FBE4FE9D34ECF9FB" box="[1132,1424,1523,1547]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor" subGenus="Ligyrus">
<emphasis id="B90A9218FF1EFF38FBE4FE9D35B6F9FA" box="[1132,1226,1523,1546]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FB53FE9D3445F9FA" box="[1243,1337,1523,1546]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FAC1FE9D34ECF9FB" box="[1353,1424,1523,1547]" italics="true" pageId="213" pageNumber="212">fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FF48FD7D30D9F9DB" author="Ratcliffe" box="[192,421,1555,1579]" firstAuthor="Ratcliffe" pageId="213" pageNumber="212" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 276</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF1EFF38FE38FD7D327BF9DB" author="Escalona, H. E. &amp; L. J. Joly" box="[432,775,1555,1579]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="213" pageNumber="212" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 118</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF1EFF38FCBFFD7D329EF9DB" box="[823,994,1555,1579]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FCBFFD7D329EF9DB" box="[823,994,1555,1579]" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
</taxonomicName>
) (EO);
<bibRefCitation id="EFEF33FBFF1EFF38FBB0FD7D34E7F9DB" author="Lopez-Garcia" box="[1080,1435,1555,1579]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="451 - 492" refId="ref212996" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, and G. Amat-Garcia. 2015. The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa 4048 (4): 451 - 492." type="journal article" year="2015">López-García et al. 2015: 482</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF1EFF38FF48FD5D3040F9BB" author="Lopez-Garcia" box="[192,316,1587,1611]" firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="491 - 509" refId="ref213049" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, R. D. Cave, and G. Amat-Garcia. 2016. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Zootaxa 4170 (3): 491 - 509." type="journal article" year="2016">2016: 502</bibRefCitation>
.
<emphasis id="B90A9218FF1EFF38FEC2FD5D30DFF9BB" bold="true" box="[330,419,1587,1611]" pageId="213" pageNumber="212">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF1EFF38FE24FD5D33F6F9BB" author="Latreille" box="[428,650,1587,1611]" firstAuthor="Latreille" pageId="213" pageNumber="212" pagination="1813 - 1817" refId="ref212262" refString="Latreille 1813&gt; Latreille 1833 (1813 - 1817)" type="book chapter" year="1813">Latreille 1813: 11</bibRefCitation>
(equinoctial America - as
<taxonomicName id="4C7E3589FF1EFF38FC50FD5D35CBF9BB" authorityName="Latreille" authorityYear="1813" box="[984,1207,1587,1611]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FC50FD5D35CBF9BB" box="[984,1207,1587,1611]" italics="true" pageId="213" pageNumber="212">Scarabaeus fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FB58FD5D34DCF9BB" author="Arrow, G. J." bookContentInfo="124 p." box="[1232,1440,1587,1611]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="213" pageNumber="212" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 37</bibRefCitation>
(Colombia, Panama - as
<taxonomicName id="4C7E3589FF1EFF38FE64FD3D33E4F99B" box="[492,664,1619,1643]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FE64FD3D33E4F99B" box="[492,664,1619,1643]" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FD27FD3D32B9F99B" author="Blackwelder, R. E." box="[687,965,1619,1643]" journalOrPublisher="United States National Museum Bulletin" pageId="213" pageNumber="212" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 254</bibRefCitation>
(Argentina, Colombia, Cuba, Panama, Puerto Rico - as
<taxonomicName id="4C7E3589FF1EFF38FE0CFD1D3353F97B" box="[388,559,1651,1675]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FE0CFD1D3353F97B" box="[388,559,1651,1675]" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FDCDFD1D325FF97B" author="Endrodi, S." box="[581,803,1651,1675]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="213" pageNumber="212" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi 1969a: 54</bibRefCitation>
(Bolivia, Colombia, Cuba, Curaçao, Jamaica, Puerto Rico - as
<taxonomicName id="4C7E3589FF1EFF38FEA3FDFD3330F95B" box="[299,588,1683,1707]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor" subGenus="Ligyrus">
<emphasis id="B90A9218FF1EFF38FEA3FDFD30F5F95A" box="[299,393,1683,1706]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FE10FDFD308AF95A" box="[408,502,1683,1706]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FD8DFDFD3330F95B" box="[517,588,1683,1707]" italics="true" pageId="213" pageNumber="212">fossor</emphasis>
</taxonomicName>
), 1985a: 255 (Bolivia, Colombia, Cuba, Jamaica, Puerto Rico - as
<taxonomicName id="4C7E3589FF1EFF38FAE9FDFD30D0F93B" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor" subGenus="Ligyrus">
<emphasis id="B90A9218FF1EFF38FAE9FDFD3194F93B" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
(
<emphasis id="B90A9218FF1EFF38FF7FFDDD3029F93A" box="[247,341,1715,1738]" italics="true" pageId="213" pageNumber="212">Ligyrus</emphasis>
)
<emphasis id="B90A9218FF1EFF38FEEDFDDD30D0F93B" box="[357,428,1715,1739]" italics="true" pageId="213" pageNumber="212">fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FE4AFDDD33D9F93B" author="Ratcliffe" box="[450,677,1715,1739]" firstAuthor="Ratcliffe" pageId="213" pageNumber="212" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 276</bibRefCitation>
(Bolivia?, Colombia, Cuba?, Jamaica?, Panama, Puerto Rico?);
<bibRefCitation id="EFEF33FBFF1EFF38FF48FDBD3373F91B" author="Escalona, H. E. &amp; L. J. Joly" box="[192,527,1747,1771]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="213" pageNumber="212" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly 2006: 118</bibRefCitation>
(Bolivia, Colombia, Cuba, Jamaica, Panama, Puerto Rico - as
<taxonomicName id="4C7E3589FF1EFF38FB61FDBD34ECF91B" box="[1257,1424,1747,1771]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FB61FDBD34ECF91B" box="[1257,1424,1747,1771]" italics="true" pageId="213" pageNumber="212">Ligyrus fossor</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF1EFF38FF48FD9D3023F8FB" author="Ponchel" box="[192,351,1779,1803]" firstAuthor="Ponchel" pageId="213" pageNumber="212" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF1EFF38FDA7FD9D32F4F8FB" author="Lopez-Garcia" box="[559,904,1779,1803]" etAl="et al." firstAuthor="Lopez-Garcia" pageId="213" pageNumber="212" pagination="451 - 492" refId="ref212996" refString="Lopez-Garcia, M. M., H. J. Gasca-Alvarez, and G. Amat-Garcia. 2015. The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution. Zootaxa 4048 (4): 451 - 492." type="journal article" year="2015">López-García et al. 2015: 482</bibRefCitation>
(Bolivia, Colombia, Cuba, Curaçao, Ecuador, El Salvador, Jamaica, Puerto Rico), 2016: 502 (Bolivia, Colombia, Ecuador, El Salvador, Panama, West Indies);
<bibRefCitation id="EFEF33FBFF1EFF38FEACFC5D30B1F8BB" author="Dupuis, F." box="[292,461,1843,1867]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="213" pageNumber="212" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b</bibRefCitation>
(not mentioned).
</paragraph>
<paragraph id="8BC14E0AFF1EFF3BFF48FC3533DAFE81" blockId="213.[192,1300,1883,1908]" lastBlockId="214.[192,1440,217,369]" lastPageId="214" lastPageNumber="213" pageId="213" pageNumber="212">
<emphasis id="B90A9218FF1EFF38FF48FC353063F883" bold="true" box="[192,287,1883,1907]" pageId="213" pageNumber="212">Note 1.</emphasis>
The year of publication of
<taxonomicName id="4C7E3589FF1EFF38FDEEFC32325EF883" baseAuthorityName="Latreille" baseAuthorityYear="1813" box="[614,802,1883,1907]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="212" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1EFF38FDEEFC32325EF883" box="[614,802,1883,1907]" italics="true" pageId="213" pageNumber="212">Tomarus fossor</emphasis>
</taxonomicName>
is sometimes erroneously given
<bibRefCitation id="EFEF33FBFF1EFF38FB27FC32346CF883" author="as" box="[1199,1296,1883,1907]" firstAuthor="as" pageId="213" pageNumber="212" refId="ref212272" refString="Latreille, P. A. 1833 (1813 - 1817). Insectes de l'Amerique equinoxiale, recueillis pendant le voyage de MM. de Humboldt et Bonpland. Seconde partie. p. 9 - 138. In: F. W. H. A. von Humboldt and A. J. A. Bonpland. Voyage de Humboldt et Bonpland. Deuxieme partie. Recueil d'observations de zoologie et d'anatomie comparee, faites dans l'Ocean Atlantique, dans l'interieur du nouveau continent et dans la mer du sud pendant les annees 1799, 1800, 1801, 1802 et 1803. Deuxieme volume. J. Smith et Gide; Paris. 352 p. [Note. This publication is thought by many to contain new species descriptions, but as Sherborn (1899: 428) already pointed out, it is in fact a republication of three papers by Latreille published from 1813 to 1817. We have been unable to find copies of the original papers]." type="book" year="1833">as 1833</bibRefCitation>
.
<emphasis id="B90A9218FF1DFF3BFF48FBB7305CFF01" bold="true" box="[192,288,217,241]" pageId="214" pageNumber="213">Note 2.</emphasis>
The only record of
<taxonomicName id="4C7E3589FF1DFF3BFD86FBB433B7FF01" baseAuthorityName="Latreille" baseAuthorityYear="1813" box="[526,715,217,241]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="214" pageNumber="213" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B90A9218FF1DFF3BFD86FBB433B7FF01" box="[526,715,217,241]" italics="true" pageId="214" pageNumber="213">Tomarus fossor</emphasis>
</taxonomicName>
from the Guianas that we are aware of is one from
<collectingCountry id="F3690E9AFF1DFF3BFAC1FBB73067FEE1" name="French Guiana" pageId="214" pageNumber="213">French Guiana</collectingCountry>
given in
<bibRefCitation id="EFEF33FBFF1DFF3BFE01FB9733CDFEE1" author="Blackwelder, R. E." box="[393,689,249,274]" journalOrPublisher="United States National Museum Bulletin" pageId="214" pageNumber="213" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder (1944: 254)</bibRefCitation>
. Based on the lack of recent records, we suspect Blackwelders record is based on a mislabeled or misidentified specimen. A record from the state of
<collectingRegion id="49BA80E8FF1DFF3BFB2EFA773583FEC1" box="[1190,1279,281,305]" country="Venezuela" name="Bolivar" pageId="214" pageNumber="213">Bolívar</collectingRegion>
in
<collectingCountry id="F3690E9AFF1DFF3BFAAAFA7734DCFEC1" box="[1314,1440,281,305]" name="Venezuela" pageId="214" pageNumber="213">Venezuela</collectingCountry>
was given in
<bibRefCitation id="EFEF33FBFF1DFF3BFED6FA573331FEA1" author="Endrodi, S." box="[350,589,313,337]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="214" pageNumber="213" pagination="1 - 145" part="37" refId="ref200804" refString="Endrodi, S. 1969 a. Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 37 (1): 1 - 145." title="Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). 1. Amerikanische Pentodontini" type="journal article" year="1969">Endrödi (1969a: 54)</bibRefCitation>
and repeated in
<bibRefCitation id="EFEF33FBFF1DFF3BFC91FA5735F9FEA2" author="Escalona, H. E. &amp; L. J. Joly" box="[793,1157,313,338]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="214" pageNumber="213" pagination="111 - 137" part="39" refId="ref201573" refString="Escalona, H. E., and L. J. Joly. 2006. El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini). Boletin de la Sociedad Entomologica Aragonesa 39: 111 - 137." title="El genero Ligyrus Burmeister, 1847 en Venezuela (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini)" type="journal article" year="2006">Escalona and Joly (2006: 118)</bibRefCitation>
. We regard this species as potentially occurring in the Guianas.
</paragraph>
</subSubSection>
</treatment>
</document>