160 lines
19 KiB
XML
160 lines
19 KiB
XML
<document id="5B32BAE37405900444FDEB20F5153995" ID-DOI="10.5281/zenodo.276822" ID-GBIF-Dataset="f5489381-8fe8-48a0-9fb6-5addeebaa3f8" ID-ISSN="1175-5326" ID-Zenodo-Dep="276822" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460237113383" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2011" docId="03C687E4FFD1FF9AFF1BEA3711B4FF3D" docLanguage="en" docName="zt02763p033.pdf" docOrigin="Zootaxa 2763" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hlavaciellus miser Jałoszyński, 2011, sp. n." docType="treatment" docVersion="6" lastPageNumber="15" masterDocId="FFFFFF9CFFDFFF95FF8CEB58101EFFCA" masterDocTitle="Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="15" updateTime="1698239820214" updateUser="plazi">
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<mods:titleInfo id="C82F7564FFA73A2A70865F6A1D10A4EE">
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<mods:title id="DF70BF781BFE305F277FBBE42308DB2F">Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="2A1641F3F1C83E0A00224B16C08C7F9E" type="personal">
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<mods:roleTerm id="699BF72B02E7DA9303C23E63D97F42F9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E56CE69A14F5E156C26F7DCE952DC27F">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="D36FE5666BC8B0132D6A4BD9AE2C4B19">text</mods:typeOfResource>
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<mods:title id="2FB71E76BA3D75B935A3569A3663752A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0D3A817059CD5E091EE2E9E351AEDD5F">
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<mods:date id="DD9B61148126B10E05E205983BE036F9">2011</mods:date>
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<mods:detail id="710FE34B21F36994E9C5016419D3CD4D" type="volume">
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<mods:number id="47F298086EA9170D4ECF37927734C68A">2763</mods:number>
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<mods:start id="1868FE03E077E95AD60D8E4B8281882B">1</mods:start>
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<mods:end id="F0080C6BCB77BFBB24703B6778A680D2">33</mods:end>
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<mods:classification id="2F2DE747492CE58F554209BF25502FB4">journal article</mods:classification>
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<mods:identifier id="D97003799F10F0A6045EC7E35751D903" type="DOI">10.5281/zenodo.276822</mods:identifier>
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<mods:identifier id="6253923D38AD37B6A24F0CBF01940B3B" type="GBIF-Dataset">f5489381-8fe8-48a0-9fb6-5addeebaa3f8</mods:identifier>
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<mods:identifier id="5E2705438733D4C217686B2AA7E8230B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="097C6FB97AFD341A951B119FCA61A773" type="Zenodo-Dep">276822</mods:identifier>
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<treatment id="03C687E4FFD1FF9AFF1BEA3711B4FF3D" ID-DOI="http://doi.org/10.5281/zenodo.6192611" ID-GBIF-Taxon="119388736" ID-Zenodo-Dep="6192611" LSID="urn:lsid:plazi:treatment:03C687E4FFD1FF9AFF1BEA3711B4FF3D" httpUri="http://treatment.plazi.org/id/03C687E4FFD1FF9AFF1BEA3711B4FF3D" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="15">
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<subSubSection id="C3756579FFD1FF9BFF1BEA371146FE60" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8BD036F2FFD1FF9BFF1BEA3711DFFE43" blockId="14.[151,449,367,426]" box="[151,449,367,393]" pageId="14" pageNumber="15">
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<heading id="D098819EFFD1FF9BFF1BEA3711DFFE43" bold="true" box="[151,449,367,393]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="B91BEAE0FFD1FF9BFF1BEA3711DFFE43" bold="true" box="[151,449,367,393]" pageId="14" pageNumber="15">
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<taxonomicName id="4C6F4D71FFD1FF9BFF1BEA371166FE43" ID-CoL="3M5PW" box="[151,376,367,393]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="miser" status="sp. nov.">
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<emphasis id="B91BEAE0FFD1FF9BFF1BEA371166FE43" bold="true" box="[151,376,367,393]" italics="true" pageId="14" pageNumber="15">Hlavaciellus miser</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A228579BFFD1FF9BFEF3EA3711DFFE43" box="[383,449,367,393]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD036F2FFD1FF9BFF1BEACA1146FE60" blockId="14.[151,449,367,426]" box="[151,344,402,426]" pageId="14" pageNumber="15">
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(
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<figureCitation id="13542A77FFD1FF9BFF13EACA10E3FE60" box="[159,253,402,426]" captionStart="FIGURES 24 – 26" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[166,1429,853,1827]" captionTargetId="figure@12.[151,1436,844,1840]" captionTargetPageId="12" captionText="FIGURES 24 – 26. Dorsal view of holotypes of Hlavaciellus, males. 24, H. bifoveatus sp. n. 25, H. metrios sp. n. 26, H. miser sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276827/files/figure.png" pageId="14" pageNumber="15">Figs. 26</figureCitation>
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, 35, 36)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD1FF9BFF1BEA841244FDF6" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="8BD036F2FFD1FF9BFF1BEA841244FDF6" blockId="14.[151,1437,475,2012]" pageId="14" pageNumber="15">
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<emphasis id="B91BEAE0FFD1FF9BFF1BEA84126DFE3E" bold="true" box="[151,627,475,500]" pageId="14" pageNumber="15">
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<typeStatus id="54D48850FFD1FF9BFF1BEA8410CCFE3E" box="[151,210,476,500]" pageId="14" pageNumber="15">Type</typeStatus>
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material.
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<typeStatus id="54D48850FFD1FF9BFEDFEA8411DAFE3E" box="[339,452,476,500]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3787662FFD1FF9BFE58EA831270FE3E" box="[468,622,475,500]" name="Malaysia" pageId="14" pageNumber="15">MALAYSIA</collectingCountry>
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:
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</emphasis>
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ɗ, "
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<collectionCode id="ED7EAE37FFD1FF9BFD20EA83130CFE39" box="[684,786,475,499]" pageId="14" pageNumber="15">SABAH</collectionCode>
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: E Mt. Kinabalu \
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<quantity id="4C979B17FFD1FF9BFC7FEA831453FE39" box="[1011,1101,475,499]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="15" unit="m" value="1150.0">1150 m</quantity>
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, rte Ranau–Kota \ Kinabalu,
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<date id="FFD11032FFD1FF9BFF1BEAA71117FDDD" box="[151,265,511,535]" pageId="14" pageNumber="15" value="1987-05-24">24.V.1987</date>
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\ Burckhardt - Löbl" [white, printed], "
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<taxonomicName id="4C6F4D71FFD1FF9BFD6EE95813BAFDD2" box="[738,932,512,536]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
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<emphasis id="B91BEAE0FFD1FF9BFD6EE95813BAFDD2" box="[738,932,512,536]" italics="true" pageId="14" pageNumber="15">HLAVACIELLUS</emphasis>
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</taxonomicName>
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\
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<taxonomicName id="4C6F4D71FFD1FF9BFC4EE959141EFDD2" box="[962,1024,513,536]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="miser">
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<emphasis id="B91BEAE0FFD1FF9BFC4EE959141EFDD2" box="[962,1024,513,536]" italics="true" pageId="14" pageNumber="15">miser</emphasis>
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</taxonomicName>
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m. \ det. P. JAŁOSZYŃSKI, '09 \
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<typeStatus id="54D48850FFD1FF9BFF1BE97C115FFDF6" box="[151,321,548,572]" pageId="14" pageNumber="15" type="holotype">HOLOTYPUS</typeStatus>
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" [red, printed] (
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<collectionCode id="ED7EAE37FFD1FF9BFE7EE97C1250FDF6" box="[498,590,548,572]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="15">MHNG</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD1FF9BFF4BE91F1471FD4E" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="8BD036F2FFD1FF9BFF4BE91F1471FD4E" blockId="14.[151,1437,475,2012]" pageId="14" pageNumber="15">
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<emphasis id="B91BEAE0FFD1FF9BFF4BE91F115FFDAA" bold="true" box="[199,321,583,608]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
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Head of males non-modified; antennomere III subquadrate; antennomere XI very slender, 2.2x as long as broad; aedeagus in ventral view very slender, strongly narrowing in apical third.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD1FF9BFF4BE9D711BCF8E2" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BD036F2FFD1FF9BFF4BE9D710A1FD06" blockId="14.[151,1437,475,2012]" pageId="14" pageNumber="15">
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<emphasis id="B91BEAE0FFD1FF9BFF4BE9D71147FD62" bold="true" box="[199,345,655,680]" pageId="14" pageNumber="15">Description.</emphasis>
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BL 1.83. Body of male (
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<figureCitation id="13542A77FFD1FF9BFDFCE9D712DCFD6D" box="[624,706,655,679]" captionStart="FIGURES 24 – 26" captionStartId="12.[151,264,1860,1883]" captionTargetBox="[166,1429,853,1827]" captionTargetId="figure@12.[151,1436,844,1840]" captionTargetPageId="12" captionText="FIGURES 24 – 26. Dorsal view of holotypes of Hlavaciellus, males. 24, H. bifoveatus sp. n. 25, H. metrios sp. n. 26, H. miser sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276827/files/figure.png" pageId="14" pageNumber="15">Fig. 26</figureCitation>
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) strongly convex, pigmentation reddish-brown, vestiture yellowish.
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</paragraph>
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<paragraph id="8BD036F2FFD1FF9BFF4BE98F1204FB92" blockId="14.[151,1437,475,2012]" pageId="14" pageNumber="15">
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Head large, HL 0.25, HW 0.45; vertex convex, not separated from frons; frons slightly flattened; clypeus convex; supraantennal tubercles small but distinct, clearly but not sharply delimited from median part of frons; eyes large, strongly convex and coarsely faceted. Punctures on dorsal surface of head very fine, unevenly distributed, separated by spaces 1–
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<date id="FFD11032FFD1FF9BFE1FE81C11BCFC96" box="[403,418,836,860]" pageId="14" pageNumber="15">2</date>
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x as wide as puncture diameters; setae short, relatively dense, suberect. Antennae as long as 0.75 BL, strikingly slender, gradually but only slightly thickened up to antennomere X, uniformly covered with thin, long, dense and suberect setae, AnL 1.38, antennomere I 1.5x as long as broad; II distinctly narrower and shorter than I, 1.1x as long as broad; III similar to II, subquadrate; IV slightly broader and much longer than III, 1.5x as long as broad; V slightly broader and distinctly longer than IV, 1.7x as long as broad; VI slightly broader but shorter than V, 1.4x as long as broad; VII–X subequal in length and barely noticeably increasing in width, each slightly broader and longer than VI, 1.4–1.5x as long as broad; XI very slender, slightly narrower than X and distinctly shorter than IX–X together.
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</paragraph>
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<paragraph id="8BD036F2FFD1FF9BFF4BEF3D1498FA9E" blockId="14.[151,1437,475,2012]" pageId="14" pageNumber="15">Pronotum in dorsal view subrectangular with moderately convex anterior margin, broadest at base and in anterior third, PL 0.53, PW 0.73; sides strongly rounded in anterior fourth, distinctly concave between anterior third and slightly sharp, rounded hind angles; posterior margin shallowly bi-emarginate; ante-basal pits very shallow and diffused, each located slightly closer to posterior than to lateral margin of pronotum; lateral edges indistinctly swollen on nearly entire length. Pronotal disc glossy, in middle covered with very fine and shallow punctures separated by spaces 0.5–1.5x as wide as puncture diameters, punctures close to margins of pronotum are larger and denser, those near posterior angles appear coarse. Setae long, moderately dense, suberect to erect.</paragraph>
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<paragraph id="8BD036F2FFD1FF9BFF4BEE381150F99A" blockId="14.[151,1437,475,2012]" pageId="14" pageNumber="15">
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Elytra oval, relatively long, broadest between middle and anterior third, EL 1.05, EW 0.85, EI 1.24; basal pit on each elytron small but distinct, located very close to scutellum; subhumeral lines sharply marked, as long as 0.36x EL, strongly divergent, each developed as border between higher humeral region and lower adsutural area; apices of elytra separately rounded. Punctures on basal third of each elytron only slightly more distinct than those on pronotum, separated by spaces 1–1.5x as wide as puncture diameters, gradually reducing in diameter and depth toward lateral margins and apices; setae similar to those on pronotum but slightly thicker and longer.
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<collectingCountry id="F3787662FFD1FF9BFA95ED4C154FF9E6" box="[1305,1361,1556,1580]" name="India" pageId="14" pageNumber="15">Hind</collectingCountry>
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wings fully developed.
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</paragraph>
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<paragraph id="8BD036F2FFD1FF9BFF4BED0512C2F9BE" blockId="14.[151,1437,475,2012]" box="[199,732,1628,1653]" pageId="14" pageNumber="15">Legs slender and long, all tibiae nearly straight.</paragraph>
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<paragraph id="8BD036F2FFD1FF9BFF4BEDD8148DF952" blockId="14.[151,1437,475,2012]" box="[199,1171,1664,1688]" pageId="14" pageNumber="15">Metaventrite with deep postmesocoxal impressions, occupying about half of its length.</paragraph>
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<paragraph id="8BD036F2FFD1FF9BFF4BEDFC1583F8CE" blockId="14.[151,1437,475,2012]" pageId="14" pageNumber="15">Aedeagus (Figs. 35, 36) slender but relatively small, AeL 0.35; median lobe gradually and in distal third strongly narrowed towards subtriangular, blunt apex; internal armature slightly asymmetrical; parameres in lateral view slender but strongly expanded and distinctly bent dorsally in subapical region, with moderately curved apices.</paragraph>
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<paragraph id="8BD036F2FFD1FF9BFF4BEC4811BCF8E2" blockId="14.[151,1437,475,2012]" box="[199,418,1808,1832]" pageId="14" pageNumber="15">
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<emphasis id="B91BEAE0FFD1FF9BFF4BEC48113BF8E2" bold="true" box="[199,293,1808,1832]" pageId="14" pageNumber="15">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD1FF9BFF4BEC6B12DEF886" box="[199,704,1843,1868]" pageId="14" pageNumber="15" type="distribution">
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<paragraph id="8BD036F2FFD1FF9BFF4BEC6B12DEF886" blockId="14.[151,1437,475,2012]" box="[199,704,1843,1868]" pageId="14" pageNumber="15">
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<emphasis id="B91BEAE0FFD1FF9BFF4BEC6B117EF886" bold="true" box="[199,352,1843,1868]" pageId="14" pageNumber="15">Distribution.</emphasis>
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East
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<collectingCountry id="F3787662FFD1FF9BFE12EC6C1217F886" box="[414,521,1844,1868]" name="Malaysia" pageId="14" pageNumber="15">Malaysia</collectingCountry>
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: Borneo, Sabah.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD1FF9BFF4BEC0013F1F85E" pageId="14" pageNumber="15" type="etymology">
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<paragraph id="8BD036F2FFD1FF9BFF4BEC0013F1F85E" blockId="14.[151,1437,475,2012]" pageId="14" pageNumber="15">
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<emphasis id="B91BEAE0FFD1FF9BFF4BEC001150F8BA" bold="true" box="[199,334,1880,1904]" pageId="14" pageNumber="15">Etymology.</emphasis>
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The Latin adjective "
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<taxonomicName id="4C6F4D71FFD1FF9BFDC8EC01129CF8BA" box="[580,642,1881,1904]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="miser">
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<emphasis id="B91BEAE0FFD1FF9BFDC8EC01129CF8BA" box="[580,642,1881,1904]" italics="true" pageId="14" pageNumber="15">miser</emphasis>
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</taxonomicName>
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", meaning (among others) "poor, unfortunate, miserable", was chosen to reflect lack of any secondary sexual characters in the male of this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756579FFD1FF9AFF4BECF811B4FF3D" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
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<paragraph id="8BD036F2FFD1FF9AFF4BECF811B4FF3D" blockId="14.[151,1437,475,2012]" lastBlockId="15.[151,1436,151,248]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<emphasis id="B91BEAE0FFD1FF9BFF4BECF81125F872" bold="true" box="[199,315,1952,1976]" pageId="14" pageNumber="15">Remarks.</emphasis>
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This unremarkable species can be distinguished from similar congeners on the basis of the slender aedeagus, which in ventral view is strongly narrowing towards the apex from apical third of its length. Additionally,
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<taxonomicName id="4C6F4D71FFD0FF9AFF40EBC11133FF7A" box="[204,301,153,176]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="miser">
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<emphasis id="B91BEAE0FFD0FF9AFF40EBC11133FF7A" box="[204,301,153,176]" italics="true" pageId="15" pageNumber="16">H. miser</emphasis>
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</taxonomicName>
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differs from the most similar
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<taxonomicName id="4C6F4D71FFD0FF9AFDFBEBC112F3FF7A" box="[631,749,153,176]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="metrios">
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<emphasis id="B91BEAE0FFD0FF9AFDFBEBC112F3FF7A" box="[631,749,153,176]" italics="true" pageId="15" pageNumber="16">H. metrios</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C6F4D71FFD0FF9AFCAEEBC113D1FF7A" box="[802,975,152,176]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="clandestinus">
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<emphasis id="B91BEAE0FFD0FF9AFCAEEBC113D1FF7A" box="[802,975,152,176]" italics="true" pageId="15" pageNumber="16">H. clandestinus</emphasis>
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</taxonomicName>
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in more slender antennomere XI (2.2x as long as broad vs. 1.8 and 1.9x, respectively), and the antennomere IV 1.5x as long as III (1.1x in
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<taxonomicName id="4C6F4D71FFD0FF9AFB7FEBE51575FF1E" box="[1267,1387,189,212]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="metrios">
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<emphasis id="B91BEAE0FFD0FF9AFB7FEBE51575FF1E" box="[1267,1387,189,212]" italics="true" pageId="15" pageNumber="16">H. metrios</emphasis>
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</taxonomicName>
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and 1.2x in
|
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<taxonomicName id="4C6F4D71FFD0FF9AFF60EBB91184FF32" box="[236,410,224,248]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="clandestinus">
|
||
<emphasis id="B91BEAE0FFD0FF9AFF60EBB91184FF32" box="[236,410,224,248]" italics="true" pageId="15" pageNumber="16">H. clandestinus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |