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<document id="4BA66370FD3BE47037FF5D1181481CE9" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier &amp; Patrick Arnaud" docDate="2016" docId="03C68782F054FFCFFF42FA11FB25F980" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Onthoecus) amyntas Lacordaire 1856" docType="treatment" docVersion="10" lastPageNumber="57" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="55" updateTime="1698234910999" updateUser="plazi">
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<mods:title id="C37B663F4BF781B3216F12FD201616CF">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
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<mods:namePart id="CE68D1C367E836C952858F5B2F6C542F">François Génier</mods:namePart>
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<mods:namePart id="D1DECFD41D7B2342D8AF627F39B2102B">Patrick Arnaud</mods:namePart>
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<treatment id="03C68782F054FFCFFF42FA11FB25F980" ID-DOI="http://doi.org/10.5281/zenodo.5780322" ID-GBIF-Taxon="127631968" ID-Zenodo-Dep="5780322" LSID="urn:lsid:plazi:treatment:03C68782F054FFCFFF42FA11FB25F980" httpUri="http://treatment.plazi.org/id/03C68782F054FFCFFF42FA11FB25F980" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55">
<subSubSection id="C375651FF054FFC1FF42FA11FC85F9C6" box="[151,867,1532,1558]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="8BD03694F054FFC1FF42FA11FC85F9C6" blockId="54.[151,867,1532,1591]" box="[151,867,1532,1558]" pageId="54" pageNumber="55">
<heading id="D09881F8F054FFC1FF42FA11FC85F9C6" bold="true" box="[151,867,1532,1558]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<emphasis id="B91BEA86F054FFC1FF42FA11FC85F9C6" bold="true" box="[151,867,1532,1558]" pageId="54" pageNumber="55">
27.
<taxonomicName id="4C6F4D17F054FFC1FF16FA11FC85F9C6" ID-CoL="34R2K" authority="Lacordaire, 1856" authorityName="Lacordaire" authorityYear="1856" box="[195,867,1532,1558]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="amyntas" subGenus="Onthoecus">
<emphasis id="B91BEA86F054FFC1FF16FA11FD61F9C6" bold="true" box="[195,647,1532,1558]" italics="true" pageId="54" pageNumber="55">Dendropaemon (Onthoecus) amyntas</emphasis>
Lacordaire, 1856
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C375651FF054FFC1FF42F9F3FCB7F967" pageId="54" pageNumber="55" type="description">
<paragraph id="8BD03694F054FFC1FF42F9F3FE21F9E7" blockId="54.[151,867,1532,1591]" box="[151,455,1566,1591]" pageId="54" pageNumber="55">
(
<figureCitation id="13542A11F054FFC1FF4AF9F3FF1AF9E7" box="[159,252,1566,1591]" captionStart="FIGURES 25 30" captionStartId="103.[151,264,1962,1985]" captionTargetBox="[269,1325,201,1926]" captionTargetId="figure@103.[251,1336,193,1941]" captionTargetPageId="103" captionText="FIGURES 25 30. Dorsal habitus. 25. D. inemarginatus, female holotype; 26. D. nigritulus, male holotype; 27. D. amyntas, male; 28. D. attalus, male holotype; 29. D. lydiae, male holotype; 30. D. morettoi, male holotype." httpUri="https://zenodo.org/record/268279/files/figure.png" pageId="54" pageNumber="55">Figs. 27</figureCitation>
, 8485, 122, 159)
</paragraph>
<paragraph id="8BD03694F054FFC1FF42F989FBE7F9AA" blockId="54.[151,1266,1636,2026]" box="[151,1025,1636,1658]" pageId="54" pageNumber="55">
<taxonomicName id="4C6F4D17F054FFC1FF42F989FDD0F9A9" authority="Amyntas Lacordaire 1856" authorityName="Amyntas Lacordaire" authorityYear="1856" box="[151,566,1636,1658]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F054FFC1FF42F989FE6CF9A9" box="[151,394,1636,1658]" italics="true" pageId="54" pageNumber="55">Dendropaemon Amyntas</emphasis>
Lacordaire
<emphasis id="B91BEA86F054FFC1FDD1F989FDD0F9A9" bold="true" box="[516,566,1636,1657]" pageId="54" pageNumber="55">1856</emphasis>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FD96F988FD37F9AA" box="[579,721,1637,1658]" italics="true" pageId="54" pageNumber="55">
Hist.
<taxonomicName id="4C6F4D17F054FFC1FDAFF988FD44F9AA" box="[634,674,1637,1658]" class="Insecta" family="Coccinellidae" genus="Nat" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Nat</taxonomicName>
. Ins.
</emphasis>
III: 103 (original description)
</paragraph>
<paragraph id="8BD03694F054FFC1FF42F96FFBC1F948" blockId="54.[151,1266,1636,2026]" box="[151,1063,1666,1688]" pageId="54" pageNumber="55">
<taxonomicName id="4C6F4D17F054FFC1FF42F96FFE6CF948" authority="Amyntas" authorityName="Amyntas" box="[151,394,1666,1688]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F054FFC1FF42F96FFE6CF948" box="[151,394,1666,1688]" italics="true" pageId="54" pageNumber="55">Dendropaemon Amyntas</emphasis>
</taxonomicName>
: Lacordaire
<emphasis id="B91BEA86F054FFC1FDDEF96FFDDBF947" bold="true" box="[523,573,1666,1687]" pageId="54" pageNumber="55">1856</emphasis>
,
<emphasis id="B91BEA86F054FFC1FD9FF96EFCF4F948" box="[586,786,1666,1688]" italics="true" pageId="54" pageNumber="55">
Hist.
<taxonomicName id="4C6F4D17F054FFC1FD54F96EFD4FF948" box="[641,681,1667,1688]" class="Insecta" family="Coccinellidae" genus="Nat" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Nat</taxonomicName>
. Ins.
<taxonomicName id="4C6F4D17F054FFC1FD0BF96FFCF4F948" authority="Pl." authorityName="Pl." box="[734,786,1666,1688]" class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Atlas</taxonomicName>
</emphasis>
: Pl. 27,
<figureCitation id="13542A11F054FFC1FCB2F96FFC44F948" box="[871,930,1666,1688]" captionStart="FIGURES 1 6" captionStartId="99.[151,264,1952,1975]" captionTargetBox="[245,1323,190,1918]" captionTargetId="figure@99.[230,1338,173,1934]" captionTargetPageId="99" captionText="FIGURES 1 6. Dorsal habitus. 1. D. bluti, male holotype; 2. D. carinifer, male holotype; 3. D. compressipennis, male holotype; 4. D. cribrosus, male holotype; 5. D. furtadoi, male holotype; 6. D. hirticollis, female holotype." httpUri="https://zenodo.org/record/268275/files/figure.png" pageId="54" pageNumber="55">Fig. 5</figureCitation>
(illustration)
</paragraph>
<paragraph id="8BD03694F054FFC1FF42F94CFCB7F967" blockId="54.[151,1266,1636,2026]" box="[151,849,1697,1719]" pageId="54" pageNumber="55">
<taxonomicName id="4C6F4D17F054FFC1FF42F94CFE62F967" authority="Amyntas" authorityName="Amyntas" box="[151,388,1697,1719]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEA86F054FFC1FF42F94CFE62F967" box="[151,388,1697,1719]" italics="true" pageId="54" pageNumber="55">Dendropemon Amyntas</emphasis>
</taxonomicName>
: Harold
<emphasis id="B91BEA86F054FFC1FE0BF94CFDF7F966" bold="true" box="[478,529,1697,1718]" pageId="54" pageNumber="55">1869</emphasis>
,
<emphasis id="B91BEA86F054FFC1FDC8F94FFD9CF967" box="[541,634,1697,1719]" italics="true" pageId="54" pageNumber="55">Cat. Col.</emphasis>
IV: 1020 (catalogue)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF054FFC1FF42F92DFC43F83A" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="8BD03694F054FFC1FF42F92DFBCDF8C3" blockId="54.[151,1266,1636,2026]" pageId="54" pageNumber="55">
<taxonomicName id="4C6F4D17F054FFC1FF42F92DFDB7F905" ID-CoL="34R3R" authority="Olsoufieff 1924" authorityName="Olsoufieff" authorityYear="1924" box="[151,593,1728,1750]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="waterhousi">
<emphasis id="B91BEA86F054FFC1FF42F92DFE4CF906" box="[151,426,1728,1750]" italics="true" pageId="54" pageNumber="55">Dendropaemon waterhousi</emphasis>
<bibRefCitation id="EFFE4B65F054FFC1FE65F92DFDB7F905" author="Olsoufieff" box="[432,593,1728,1750]" pageId="54" pageNumber="55" refString="Olsoufieff, G. d' (1924) Les Phanaeides (Coleoptera-Lamellicornia), Famille Scarabaeidae-Tr. Coprini. Insecta, 13, 4 - 172, 15 pls." type="journal article" year="1924">
Olsoufieff
<emphasis id="B91BEA86F054FFC1FDCBF92DFDB7F905" bold="true" box="[542,593,1728,1749]" pageId="54" pageNumber="55">1924</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FD88F92CFD40F906" box="[605,678,1729,1750]" italics="true" pageId="54" pageNumber="55">Insecta</emphasis>
13: 125 (original description)
<emphasis id="B91BEA86F054FFC1FC09F92DFB9AF905" bold="true" box="[988,1148,1728,1749]" pageId="54" pageNumber="55">new synonymy</emphasis>
<taxonomicName id="4C6F4D17F054FFC1FF42F933FD6EF924" ID-CoL="34R3R" authority="Pessoa &amp; Lane 1936" authorityName="Pessoa &amp; Lane" authorityYear="1936" box="[151,648,1758,1780]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="waterhousi">
<emphasis id="B91BEA86F054FFC1FF42F933FE4CF924" box="[151,426,1758,1780]" italics="true" pageId="54" pageNumber="55">Dendropaemon waterhousi</emphasis>
:
<bibRefCitation id="EFFE4B65F054FFC1FE62F932FD6EF924" author="Pessoa" box="[439,648,1758,1780]" pageId="54" pageNumber="55" refString="Pessoa, S. B. &amp; Lane, F. (1936) Sobre os coleopteros do genero Dendropaemon Perty, de S. Paulo e regioes visinhas. Revista de Biologia e Hygiene, 7 (2), 89 - 93." type="journal article" year="1936">
Pessôa &amp; Lane
<emphasis id="B91BEA86F054FFC1FD80F932FD6EF924" bold="true" box="[597,648,1759,1780]" pageId="54" pageNumber="55">1936</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FD41F90DFCB6F925" box="[660,848,1758,1781]" italics="true" pageId="54" pageNumber="55">Rev. Biol. Hygiene</emphasis>
7: 90 (identification key, comment)
<taxonomicName id="4C6F4D17F054FFC1FF42F910FD65F8C2" ID-CoL="34R3R" authority="Blut 1939" authorityName="Blut" authorityYear="1939" box="[151,643,1789,1811]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="amynthas" subSpecies="waterhousi">
<emphasis id="B91BEA86F054FFC1FF42F910FDF7F8C3" box="[151,529,1789,1811]" italics="true" pageId="54" pageNumber="55">Dendropaemon amynthas waterhousi</emphasis>
:
<bibRefCitation id="EFFE4B65F054FFC1FDCBF910FD65F8C2" author="Blut" box="[542,643,1789,1811]" pageId="54" pageNumber="55" refString="Blut, H. (1939) Beitrag zur Verbreitung und Systematik der Gattung Dendropaemon. Archiv fur Naturgeschichte, N. F., 8, 263 - 300." type="journal article" year="1939">
Blut
<emphasis id="B91BEA86F054FFC1FD85F910FD65F8C2" bold="true" box="[592,643,1789,1810]" pageId="54" pageNumber="55">1939</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FD45F910FCBCF8C3" box="[656,858,1789,1811]" italics="true" pageId="54" pageNumber="55">Arch. Naturg. (N.F.)</emphasis>
8: 269 (monograph)
</paragraph>
<paragraph id="8BD03694F054FFC1FF42F8F1FBF1F880" blockId="54.[151,1266,1636,2026]" pageId="54" pageNumber="55">
<taxonomicName id="4C6F4D17F054FFC1FF42F8F1FD75F8E1" authority="Pessoa &amp; Lane 1941" authorityName="Pessoa &amp; Lane" authorityYear="1941" box="[151,659,1820,1842]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="waterhousei">
<emphasis id="B91BEA86F054FFC1FF42F8F1FE53F8E2" box="[151,437,1820,1842]" italics="true" pageId="54" pageNumber="55">Dendropaemon waterhousei</emphasis>
:
<bibRefCitation id="EFFE4B65F054FFC1FE17F8F1FD75F8E1" author="Pessoa" box="[450,659,1820,1842]" pageId="54" pageNumber="55" refString="Pessoa, S. B. &amp; Lane, F. (1941) Coleopteros necrophagos de interesse medico-legal. Ensaio monografico sobre a familia Scarabaeidae de Sao Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo, 2, 389 - 504." type="journal article" year="1941">
Pessôa &amp; Lane
<emphasis id="B91BEA86F054FFC1FDB5F8F1FD75F8E1" bold="true" box="[608,659,1820,1841]" pageId="54" pageNumber="55">1941</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FD75F8F1FC85F8E2" box="[672,867,1820,1842]" italics="true" pageId="54" pageNumber="55">Arq. Zool. S. Paulo</emphasis>
2: 490 (identification key, distribution)
<taxonomicName id="4C6F4D17F054FFC1FF42F8D7FD89F89F" authority="Blackwelder 1944" authorityName="Blackwelder" authorityYear="1944" box="[151,623,1850,1872]" class="Insecta" family="Anthribidae" genus="Dendropemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="waterhousei">
<emphasis id="B91BEA86F054FFC1FF42F8D7FE4EF880" box="[151,424,1850,1872]" italics="true" pageId="54" pageNumber="55">Dendropemon waterhousei</emphasis>
:
<bibRefCitation id="EFFE4B65F054FFC1FE63F8D7FD89F89F" author="Blackwelder" box="[438,623,1850,1872]" pageId="54" pageNumber="55" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 341. http: // dx. doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">
Blackwelder
<emphasis id="B91BEA86F054FFC1FDE9F8D7FD89F89F" bold="true" box="[572,623,1850,1871]" pageId="54" pageNumber="55">1944</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FDAEF8D6FCAEF880" box="[635,840,1850,1872]" italics="true" pageId="54" pageNumber="55">
<collectingCountry id="F3787604F054FFC1FDAEF8D6FD41F880" box="[635,679,1851,1872]" name="United States of America" pageId="54" pageNumber="55">U.S.</collectingCountry>
<taxonomicName id="4C6F4D17F054FFC1FD78F8D6FD33F880" box="[685,725,1851,1872]" class="Insecta" family="Coccinellidae" genus="Nat" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Nat</taxonomicName>
. Mus. Bull.
</emphasis>
185: 211 (checklist)
</paragraph>
<paragraph id="8BD03694F054FFC1FF42F8B4FC43F83A" blockId="54.[151,1266,1636,2026]" pageId="54" pageNumber="55">
<taxonomicName id="4C6F4D17F054FFC1FF42F8B4FD82F8BE" ID-CoL="34R2K" authority="Edmonds 1972" authorityName="Edmonds" authorityYear="1972" box="[151,612,1881,1903]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="amyntas" subGenus="Dendropaemon">
<emphasis id="B91BEA86F054FFC1FF42F8B4FE5BF8BF" box="[151,445,1881,1903]" italics="true" pageId="54" pageNumber="55">Dendropaemon (D.) amyntas</emphasis>
:
<bibRefCitation id="EFFE4B65F054FFC1FE1EF8B4FD82F8BE" author="Edmonds" box="[459,612,1881,1903]" pageId="54" pageNumber="55" refString="Edmonds, W. D. (1972) Comparative Skeletal Morphology, Systematics and Evolution of the Phanaeine Dung Beetles (Coleoptera: Scarabaeidae). University of Kansas Science Bulletin, 49 (11), 731 - 874." type="journal article" year="1972">
Edmonds
<emphasis id="B91BEA86F054FFC1FDE4F8B4FD82F8BE" bold="true" box="[561,612,1881,1902]" pageId="54" pageNumber="55">1972</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FDA5F8B7FCB5F8BF" box="[624,851,1881,1903]" italics="true" pageId="54" pageNumber="55">Univ. Kansas Sc. Bull.</emphasis>
49: 850 (misidentification)
<taxonomicName id="4C6F4D17F054FFC1FF42F895FDDAF85D" ID-CoL="34R3R" authority="Arnaud 1982" authorityName="Arnaud" authorityYear="1982" box="[151,572,1912,1934]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="waterhousi">
<emphasis id="B91BEA86F054FFC1FF42F895FE4CF85E" box="[151,426,1912,1934]" italics="true" pageId="54" pageNumber="55">Dendropaemon waterhousi</emphasis>
:
<bibRefCitation id="EFFE4B65F054FFC1FE62F895FDDAF85D" author="Arnaud" box="[439,572,1912,1934]" pageId="54" pageNumber="55" refString="Arnaud, P. (1982) Liste des types de Phanaeini du Museum national d'Histoire naturelle de Paris, suivi d'une note synonymique pour quatre especes decrite par G. d'Olsoufieff (Coleoptera, Scarabaeidae). Revue Francaise d'Entomologie, Nouvelle Serie, 4 (3), 113 - 118." type="journal article" year="1982">
Arnaud
<emphasis id="B91BEA86F054FFC1FDDCF895FDDAF85D" bold="true" box="[521,572,1912,1933]" pageId="54" pageNumber="55">1982</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FD9CF894FCE1F85E" box="[585,775,1913,1934]" italics="true" pageId="54" pageNumber="55">Rev. Fr. Ent. (N.S.)</emphasis>
4: 117 (
<typeStatus id="54D48836F054FFC1FC8FF895FC5FF85E" box="[858,953,1912,1934]" pageId="54" pageNumber="55" type="lectotype">lectotype</typeStatus>
designation)
<taxonomicName id="4C6F4D17F054FFC1FF42F87BFDF6F87C" box="[151,528,1942,1965]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="amyntas" subSpecies="waterhousei">
<emphasis id="B91BEA86F054FFC1FF42F87BFDF6F87C" box="[151,528,1942,1965]" italics="true" pageId="54" pageNumber="55">Dendropaemon amyntas waterhousei</emphasis>
</taxonomicName>
: Vaz-de-Mello
<emphasis id="B91BEA86F054FFC1FD7BF87AFD07F87C" bold="true" box="[686,737,1943,1964]" pageId="54" pageNumber="55">2000</emphasis>
,
<emphasis id="B91BEA86F054FFC1FD3BF875FC49F87C" box="[750,943,1943,1965]" italics="true" pageId="54" pageNumber="55">Hac. Proy. CYTED</emphasis>
: 192 (faunistic)
<taxonomicName id="4C6F4D17F054FFC1FF42F858FDDDF81A" authority="Vitolo 2000" authorityName="Vitolo" authorityYear="2000" box="[151,571,1973,1995]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="waterhousei">
<emphasis id="B91BEA86F054FFC1FF42F858FE53F81B" box="[151,437,1973,1995]" italics="true" pageId="54" pageNumber="55">Dendropaemon waterhousei</emphasis>
: Vitolo
<emphasis id="B91BEA86F054FFC1FDDDF858FDDDF81A" bold="true" box="[520,571,1973,1994]" pageId="54" pageNumber="55">2000</emphasis>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FD92F85BFCB1F81B" box="[583,855,1973,1995]" italics="true" pageId="54" pageNumber="55">Rev. Acad. Colomb. Cienc.</emphasis>
24: 599 (misidentification)
<taxonomicName id="4C6F4D17F054FFC1FF42F839FDA9F839" ID-CoL="34R2K" authority="Arnaud 2002" authorityName="Arnaud" authorityYear="2002" box="[151,591,2004,2026]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="amyntas" subGenus="Dendropaemon">
<emphasis id="B91BEA86F054FFC1FF42F839FE5BF83A" box="[151,445,2004,2026]" italics="true" pageId="54" pageNumber="55">Dendropaemon (D.) amyntas</emphasis>
:
<bibRefCitation id="EFFE4B65F054FFC1FE1EF839FDA9F839" author="Arnaud" box="[459,591,2004,2026]" pageId="54" pageNumber="55" refString="Arnaud, P. (2002) Phanaeini, Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Les Coleopteres du Monde, 28. Hillside Books, Canterbury, 151 pp." type="book" year="2002">
Arnaud
<emphasis id="B91BEA86F054FFC1FDC8F839FDA9F839" bold="true" box="[541,591,2004,2025]" pageId="54" pageNumber="55">2002</emphasis>
</bibRefCitation>
</taxonomicName>
,
<emphasis id="B91BEA86F054FFC1FD89F839FD35F83A" box="[604,723,2004,2026]" italics="true" pageId="54" pageNumber="55">Col. Monde</emphasis>
28: 16 (monograph)
</paragraph>
</subSubSection>
<subSubSection id="C375651FF055FFC0FF42FF7AFE60FF60" box="[151,390,151,176]" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="8BD03694F055FFC0FF42FF7AFE60FF60" blockId="55.[151,1437,151,2013]" box="[151,390,151,176]" pageId="55" pageNumber="56">
<emphasis id="B91BEA86F055FFC0FF42FF7AFECBFF60" bold="true" box="[151,301,151,176]" pageId="55" pageNumber="56">
<typeStatus id="54D48836F055FFC0FF42FF7AFF34FF60" box="[151,210,151,176]" pageId="55" pageNumber="56">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F3787604F055FFC0FEE9FF7AFE65FF60" box="[316,387,151,176]" name="Brazil" pageId="55" pageNumber="56">Brésil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF055FFC0FF12FF51FCD8FECD" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="8BD03694F055FFC0FF12FF51FCD8FECD" blockId="55.[151,1437,151,2013]" pageId="55" pageNumber="56">
<emphasis id="B91BEA86F055FFC0FF12FF51FEA7FF03" bold="true" box="[199,321,188,213]" pageId="55" pageNumber="56">Diagnosis.</emphasis>
The large size, rather thick body combined with dark metallic blue sheen of the dorsum will separate
<taxonomicName id="4C6F4D17F055FFC0FF2FFF0CFE9FFF28" box="[250,377,225,248]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F055FFC0FF2FFF0CFE9FFF28" box="[250,377,225,248]" italics="true" pageId="55" pageNumber="56">D. amyntas</emphasis>
</taxonomicName>
from all other species. It can easily be separated from its sister species
<taxonomicName id="4C6F4D17F055FFC0FB4FFF0CFAEEFF28" box="[1178,1288,223,248]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="attalus">
<emphasis id="B91BEA86F055FFC0FB4FFF0CFAEEFF28" box="[1178,1288,223,248]" italics="true" pageId="55" pageNumber="56">D. attalus</emphasis>
</taxonomicName>
by the shape of the clypeal teeth (Fig. xx) and the more slender metatibia.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF055FFCFFF12FECAFCEBFEB5" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="description">
<paragraph id="8BD03694F055FFC0FF12FECAFC01F845" blockId="55.[151,1437,151,2013]" pageId="55" pageNumber="56">
<emphasis id="B91BEA86F055FFC0FF12FECAFEB7FE90" bold="true" box="[199,337,295,320]" pageId="55" pageNumber="56">Description</emphasis>
.
<collectingCountry id="F3787604F055FFC0FEB7FECAFE7DFE90" box="[354,411,295,320]" name="Malta" pageId="55" pageNumber="56">Male</collectingCountry>
<typeStatus id="54D48836F055FFC0FE70FECAFDEDFE90" box="[421,523,295,320]" pageId="55" pageNumber="56" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="13542A11F055FFC0FDC8FECAFD95FE90" box="[541,627,295,320]" captionStart="FIGURES 25 30" captionStartId="103.[151,264,1962,1985]" captionTargetBox="[269,1325,201,1926]" captionTargetId="figure@103.[251,1336,193,1941]" captionTargetPageId="103" captionText="FIGURES 25 30. Dorsal habitus. 25. D. inemarginatus, female holotype; 26. D. nigritulus, male holotype; 27. D. amyntas, male; 28. D. attalus, male holotype; 29. D. lydiae, male holotype; 30. D. morettoi, male holotype." httpUri="https://zenodo.org/record/268279/files/figure.png" pageId="55" pageNumber="56">Fig. 27</figureCitation>
).
<emphasis id="B91BEA86F055FFC0FD5CFECAFD20FE90" bold="true" box="[649,710,295,320]" pageId="55" pageNumber="56">Body</emphasis>
. Body large, length 18.0 mm, maximum width 10.0 mm; body subrectangular; dorsum narrowly flat.
<emphasis id="B91BEA86F055FFC0FD87FEA1FD75FEB5" bold="true" box="[594,659,332,357]" pageId="55" pageNumber="56">Color</emphasis>
. Dorsal surface dark reddish brown to black, glossy, with metallic sheen; head with greenish to coppery metallic sheen adjacent to the eyes; pronotum with blue metallic sheen except for anteromedian carina, on anterior portion of disc and surface adjacent to lateral fossae; elytra with rather faint blue to purple metallic sheen; ventrum light to dark brown; pygidium with blue and green metallic sheen; legs light to dark brown.
<emphasis id="B91BEA86F055FFC0FE91FE31FE62FE25" bold="true" box="[324,388,476,501]" pageId="55" pageNumber="56">Head</emphasis>
. Clypeus broadly arcuate, anterior portion upturned; clypeal teeth triangular, lateral edges of each tooth only slightly tapering toward apex; clypeal median emargination broadly v-shaped, clypeal edge emarginate on external side of each clypeal tooth, clypeal teeth ventral surface with a fine v-shaped carina, clypeal margin ill-defined, bordered posteriorly by a more or less regular row of punctures, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with fine punctures laterally, fine tubercles medially and minute tubercles posteriorly, transversely tumescent; clypeofrontal carina as high as wide basally, gradually tapering toward apex, straight in dorsal view, simply carinate, clypeofrontal carina apical edge straight in frontal view; eyes small in dorsal view, interocular ratio 6.0.
<emphasis id="B91BEA86F055FFC0FB75FD3AFAFFFD20" bold="true" box="[1184,1305,727,752]" pageId="55" pageNumber="56">Pronotum</emphasis>
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.6; disc of pronotum finely punctate basally, punctures changing into fine rough rugulae anteriorly, with an ill-defined shallow longitudinal depression on posterior half; pronotal anterior margin slightly wider and concave lateral to eye; anterior portion with a strongly tri-sinuous carina produced into a wide truncated lobe medially; anterior angles surface with fine rugulae, similar to lateral margin along posterior edge of anterior margin; lateral fossae simply rounded, concave; lateral portions strongly explanate; pronotal basal fossae well-defined, concave; posterior margin well-defined, lacking crenulation and setae.
<emphasis id="B91BEA86F055FFC0FF08FC39FECFFC3D" bold="true" box="[221,297,980,1005]" pageId="55" pageNumber="56">Elytra</emphasis>
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.2; elytral base lacking distinct margin, simply convex; elytral striae 14 moderately wide, not bordered by a minute carina laterally, shallowly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures ill-defined, adjacent strial edge encroaching on interval, stria 1 well-defined apically, connecting to marginal stria; interstriae moderately convex, minutely punctate throughout, surface glossy.
<emphasis id="B91BEA86F055FFC0FCABFB89FBDDFBAD" bold="true" box="[894,1083,1124,1149]" pageId="55" pageNumber="56">Thoracic sterna</emphasis>
. Proepisternal carina complete, extending laterally; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, vshaped.
<emphasis id="B91BEA86F055FFC0FF21FB41FECAFB15" bold="true" box="[244,300,1196,1221]" pageId="55" pageNumber="56">Legs</emphasis>
. Profemur posterior surface convex, glabrous and punctate internally, posterointernal margin rather thick, uneven, internal edge wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface more or less irregularly punctate. Protibia with four teeth on lateral edge; internal basal angle bluntly lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with some ill-defined irregular punctures externally to median carina, surface glossy between punctures, with a single interrupted setal row along lateral teeth. Mesofemur unmodified on anterointernal edge apically. Mesotibia rather slender, gradually widening toward apex in anterior view; anteroapical edge finely crenulate, anteroapical row of setae incomplete, irregular; apicoanterior edge circularly indented internally; external edge rounded basally, more or less flat apically, surface with irregular punctures and microsculpture. Mesotarsus similar in shape to metatarsus, 3-segmented, first segment rather elongate, approximately three times as long as wide at apex. Metafemur elongate, internal and lateral edges mostly parallel in ventral view, more than twice as long as wide, depressed anterointernally before apex, apicoposterior edge lobate beyond tibial insertion, surface coarsely microsculptured, anterior surface with a well-defined irregular sulcus on median half. Metatibia robust, short, slightly widening toward apex in anterior view, anterior surface completely covered with irregular sculpturing and punctures, metatibial posterior surface concave between longitudinal row of setae and lateral edge, surface dull and irregularly punctate. Metatarsus 3-segmented (
<figureCitation id="13542A11F055FFC0FABAF92AFF22F8D4" captionStart="FIGURES 118 125" captionStartId="113.[151,264,1924,1947]" captionTargetBox="[217,1363,200,1895]" captionTargetId="figure@113.[211,1376,193,1903]" captionTargetPageId="113" captionText="FIGURES 118 125. Metatarsus, ventral view. 118. D. fredericki, holotype; 119. D. angustipennis; 120. D. bahianus; 121. D. nigritulus, paratype; 122. D. amyntas; 123. D. attalus; 124. D. lydiae, paratype; 125. D. morettoi, paratype." httpUri="https://zenodo.org/record/268283/files/figure.png" pageId="55" pageNumber="56">Fig. 122</figureCitation>
), first segment moderately elongate, approximately three times as long as wide at apex, with anterointernal carina ill-defined.
<emphasis id="B91BEA86F055FFC0FEA5F8E2FD81F8F8" bold="true" box="[368,615,1807,1832]" pageId="55" pageNumber="56">Abdominal sternites</emphasis>
. Sternites 36 longitudinally flat; sternites 46 with more than three unaligned row of setae laterally, glabrous medially; sternite 7 approximately longitudinally flat medially, shorter than segment 6 along midline; pygidium with mixed minute and fine punctures on disc.
<emphasis id="B91BEA86F055FFC0FBADF8BAFAC5F8A0" bold="true" box="[1144,1315,1879,1904]" pageId="55" pageNumber="56">
<collectingCountry id="F3787604F055FFC0FBADF8BAFB53F8A0" box="[1144,1205,1879,1904]" name="Malta" pageId="55" pageNumber="56">Male</collectingCountry>
genitalia
</emphasis>
(Figs. 84 85). Parameres dentate in dorsal view; with minute raspy tubercles apically.
</paragraph>
<paragraph id="8BD03694F055FFC0FF12F84DFA7CF868" blockId="55.[151,1437,151,2013]" box="[199,1434,1952,1977]" pageId="55" pageNumber="56">
<emphasis id="B91BEA86F055FFC0FF12F84DFE90F869" bold="true" box="[199,374,1952,1977]" pageId="55" pageNumber="56">Measurements</emphasis>
(
<specimenCount id="9D69FD1DF055FFC0FE53F84DFE0AF869" box="[390,492,1952,1977]" pageId="55" pageNumber="56" type="male">11 males</specimenCount>
,
<specimenCount id="9D69FD1DF055FFC0FE2DF84DFD92F869" box="[504,628,1952,1977]" pageId="55" pageNumber="56" type="female">10 females</specimenCount>
). Length: male 15.020.5 (17.9±1.8), female 16.521.0 (18.2±1.3) mm.
</paragraph>
<paragraph id="8BD03694F055FFC0FF12F829FE4FF80C" blockId="55.[151,1437,151,2013]" box="[199,425,1988,2013]" pageId="55" pageNumber="56">
<emphasis id="B91BEA86F055FFC0FF12F829FE44F80D" bold="true" box="[199,418,1988,2013]" pageId="55" pageNumber="56">
Primary
<typeStatus id="54D48836F055FFC0FEE6F829FE80F80D" box="[307,358,1988,2013]" pageId="55" pageNumber="56">type</typeStatus>
data
</emphasis>
.
</paragraph>
<paragraph id="8BD03694F05AFFCFFF12FF7AFC36FF60" blockId="56.[151,1437,151,1616]" box="[199,976,151,176]" pageId="56" pageNumber="57">
<taxonomicName id="4C6F4D17F05AFFCFFF12FF7AFD92FF7F" authority="Lacordaire." authorityName="Lacordaire." box="[199,628,151,176]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F05AFFCFFF12FF7AFD8BFF60" bold="true" box="[199,621,151,176]" pageId="56" pageNumber="57">
<emphasis id="B91BEA86F05AFFCFFF12FF7AFE07FF7F" bold="true" box="[199,481,151,176]" italics="true" pageId="56" pageNumber="57">Dendropaemon amyntas</emphasis>
Lacordaire
</emphasis>
.
</taxonomicName>
<collectingCountry id="F3787604F05AFFCFFDAEFF7AFD58FF60" box="[635,702,151,176]" name="Brazil" pageId="56" pageNumber="57">Brésil</collectingCountry>
(from primary citation).
</paragraph>
<paragraph id="8BD03694F05AFFCFFF12FF51FCEBFEB5" blockId="56.[151,1437,151,1616]" pageId="56" pageNumber="57">
<taxonomicName id="4C6F4D17F05AFFCFFF12FF51FD6BFF04" authority="Olsoufieff." authorityName="Olsoufieff." box="[199,653,188,213]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="waterhousi">
<emphasis id="B91BEA86F05AFFCFFF12FF51FD60FF05" bold="true" box="[199,646,188,213]" pageId="56" pageNumber="57">
<emphasis id="B91BEA86F05AFFCFFF12FF51FDE2FF05" bold="true" box="[199,516,188,213]" italics="true" pageId="56" pageNumber="57">Dendropaemon waterhousi</emphasis>
Olsoufieff
</emphasis>
.
</taxonomicName>
<typeStatus id="54D48836F05AFFCFFD4DFF50FCEEFF05" box="[664,776,189,213]" pageId="56" pageNumber="57" type="lectotype">Lectotype</typeStatus>
male (MNHN): [LAFERTÉ./ 4820.] partly handwritten; [Ex.Musaeo/ D.Sharp 1890] black border; [Museum Paris/ ex Coll./ R. Oberthur] green card; [Dendropaem.
<taxonomicName id="4C6F4D17F05AFFCFFF42FEE9FEF4FECD" box="[151,274,260,285]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="waterhousi" status="sp. nov.">waterhousi</taxonomicName>
<taxonomicNameLabel id="A22857FDF05AFFCFFEF4FEE9FEA8FECC" box="[289,334,260,284]" pageId="56" pageNumber="57" rank="species">sp.n</taxonomicNameLabel>
/ det. G. OLSOUFIEFF] partly handwritten; [
<taxonomicName id="4C6F4D17F05AFFCFFC88FEE9FBEAFECD" box="[861,1036,260,285]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
/
<taxonomicName id="4C6F4D17F05AFFCFFBF1FEE9FB3AFECD" authority="Ols." authorityName="Ols." box="[1060,1244,260,285]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="waterhousi">waterhousi Ols.</taxonomicName>
/
<typeStatus id="54D48836F05AFFCFFB21FEE9FA75FECC" box="[1268,1427,260,284]" pageId="56" pageNumber="57" type="lectotype">LECTOTYPE</typeStatus>
/ P.ARNAUD DET 1982] red border; [WORLD/ SCARAB./ DATABASE/ WSD00016617]; [
<taxonomicName id="4C6F4D17F05AFFCFFB10FECAFA93FE90" box="[1221,1397,295,320]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">Dendropaemon</taxonomicName>
♂/
<taxonomicName id="4C6F4D17F05AFFCFFF42FEA1FF12FEB4" box="[151,244,332,356]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="amyntas">amyntas</taxonomicName>
/ Lacordaire, 1856/ dét. Génier &amp; Arnaud,2012]
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05AFFCFFF12FE82FB64FCAF" pageId="56" pageNumber="57" type="materials_examined">
<paragraph id="8BD03694F05AFFCFFF12FE82FCFAFC8C" blockId="56.[151,1437,151,1616]" pageId="56" pageNumber="57">
<emphasis id="B91BEA86F05AFFCFFF12FE82FE4DFE58" bold="true" box="[199,427,367,392]" pageId="56" pageNumber="57">Material examined</emphasis>
.
<emphasis id="B91BEA86F05AFFCFFE6EFE82FDAAFE58" bold="true" box="[443,588,367,392]" pageId="56" pageNumber="57">
[NO
<collectionCode id="ED7EAE51F05AFFCFFE22FE82FDA3FE58" box="[503,581,367,392]" pageId="56" pageNumber="57">DATA</collectionCode>
]
</emphasis>
: -., coll. [anonymous]—
<specimenCount id="9D69FD1DF05AFFCFFCB3FE9DFC31FE58" box="[870,983,367,392]" pageId="56" pageNumber="57" type="female">4 females</specimenCount>
,
<specimenCount id="9D69FD1DF05AFFCFFC31FE9DFBD9FE58" box="[996,1087,367,392]" pageId="56" pageNumber="57" type="male">4 males</specimenCount>
,
<specimenCount id="9D69FD1DF05AFFCFFB99FE9DFB2BFE58" box="[1100,1229,367,392]" pageId="56" pageNumber="57" type="generic">1 specimen</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFB08FE9DFAD0FE58" box="[1245,1334,368,392]" pageId="56" pageNumber="57">BMNH</collectionCode>
,
<collectionCode id="ED7EAE51F05AFFCFFA96FE9DFA7EFE58" box="[1347,1432,368,392]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="56" pageNumber="57">IRSNB</collectionCode>
,
<collectionCode id="ED7EAE51F05AFFCFFF42FE78FF09FE7C" LSID="urn:lsid:biocol.org:col:34988" box="[151,239,405,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="56" pageNumber="57">MNHN</collectionCode>
,
<collectionCode id="ED7EAE51F05AFFCFFED4FE78FEA9FE7C" box="[257,335,405,428]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="56" pageNumber="57">MNRJ</collectionCode>
,
<collectionCode id="ED7EAE51F05AFFCFFEB4FE79FE53FE7C" box="[353,437,404,428]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="56" pageNumber="57">NMPC</collectionCode>
); -., coll. [anonymous]—
<specimenCount id="9D69FD1DF05AFFCFFD3CFE79FCA8FE7D" box="[745,846,404,429]" pageId="56" pageNumber="57" type="female">1 female</specimenCount>
,
<specimenCount id="9D69FD1DF05AFFCFFCB4FE79FC58FE7D" box="[865,958,404,429]" pageId="56" pageNumber="57" type="male">2 males</specimenCount>
(incl.
<typeStatus id="54D48836F05AFFCFFBC1FE79FB9BFE7D" box="[1044,1149,404,429]" pageId="56" pageNumber="57" type="lectotype">lectotype</typeStatus>
, 2
<typeStatus id="54D48836F05AFFCFFB79FE79FAA8FE7D" box="[1196,1358,404,429]" pageId="56" pageNumber="57" type="paralectotype">paralectotypes</typeStatus>
of
<taxonomicName id="4C6F4D17F05AFFCFFA54FE78FEF5FE00" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="waterhousi">
<emphasis id="B91BEA86F05AFFCFFA54FE78FEF5FE00" italics="true" pageId="56" pageNumber="57">D. waterhousi</emphasis>
</taxonomicName>
) (
<collectionCode id="ED7EAE51F05AFFCFFEF8FE55FE6FFE00" LSID="urn:lsid:biocol.org:col:34988" box="[301,393,440,464]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="56" pageNumber="57">MNHN</collectionCode>
);
<collectingCountry id="F3787604F05AFFCFFE4AFE5AFDECFE00" box="[415,522,439,464]" name="Brazil" pageId="56" pageNumber="57">
<emphasis id="B91BEA86F05AFFCFFE4AFE5AFDECFE00" bold="true" box="[415,522,439,464]" pageId="56" pageNumber="57">BRAZIL</emphasis>
</collectingCountry>
: [unspecified locality], [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF05AFFCFFB56FE55FB2DFE00" box="[1155,1227,439,464]" pageId="56" pageNumber="57" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFB09FE55FAC1FE00" box="[1244,1319,440,464]" pageId="56" pageNumber="57">CPFA</collectionCode>
);
<collectionCode id="ED7EAE51F05AFFCFFAE9FE55FA7EFE00" box="[1340,1432,440,464]" pageId="56" pageNumber="57">BAHIA</collectionCode>
, Ilhéus, (
<geoCoordinate id="EE5B5053F05AFFCFFF27FE31FEABFE24" box="[242,333,476,500]" direction="south" orientation="latitude" pageId="56" pageNumber="57" precision="925" value="-14.8">14°48'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F05AFFCFFE8DFE31FE57FE24" box="[344,433,476,500]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="925" value="-39.033333">39°2'W</geoCoordinate>
), [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF05AFFCFFCF1FE31FC66FE25" box="[804,896,476,501]" pageId="56" pageNumber="57" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFC5AFE31FC01FE24" box="[911,999,476,500]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="56" pageNumber="57">IRSNB</collectionCode>
); ESPIRITO
<collectionCode id="ED7EAE51F05AFFCFFBA9FE31FB3AFE24" box="[1148,1244,476,500]" pageId="56" pageNumber="57">SANTO</collectionCode>
, Parque Estadual da Forte Grande, Vitória, (
<geoCoordinate id="EE5B5053F05AFFCFFE6AFDEDFDFCFDC8" box="[447,538,512,536]" direction="south" orientation="latitude" pageId="56" pageNumber="57" precision="925" value="-20.3">20°18'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F05AFFCFFDF3FDEDFD6AFDC8" box="[550,652,512,536]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="925" value="-40.333332">40°20'W</geoCoordinate>
),
<date id="FFD11054F05AFFCFFD4AFDEDFCF5FDC8" box="[671,787,512,536]" pageId="56" pageNumber="57" value="2004-10-30">30.x.2004</date>
, coll. Erwin, Pannagnani &amp; Schiffler—
<specimenCount id="9D69FD1DF05AFFCFFB07FE12FACBFDC8" box="[1234,1325,511,536]" pageId="56" pageNumber="57" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFAE9FDEDFA76FDC8" box="[1340,1424,512,536]" pageId="56" pageNumber="57">CEMT</collectionCode>
); Vitória, (
<geoCoordinate id="EE5B5053F05AFFCFFF2EFDC9FEB0FDEC" box="[251,342,548,572]" direction="south" orientation="latitude" pageId="56" pageNumber="57" precision="925" value="-20.316668">20°19'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F05AFFCFFEB4FDC9FE21FDEC" box="[353,455,548,572]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="925" value="-40.35">40°21'W</geoCoordinate>
), [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF05AFFCFFCECFDC9FC71FDED" box="[825,919,548,573]" pageId="56" pageNumber="57" type="female">1 female</specimenCount>
,
<specimenCount id="9D69FD1DF05AFFCFFC77FDC9FC0BFDED" box="[930,1005,548,573]" pageId="56" pageNumber="57" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFC29FDC9FBB5FDEC" box="[1020,1107,548,572]" pageId="56" pageNumber="57">ZMHB</collectionCode>
);
<collectionCode id="ED7EAE51F05AFFCFFBB3FDC9FB53FDEC" box="[1126,1205,548,572]" pageId="56" pageNumber="57">MATO</collectionCode>
GROSSO DO SUL, Dourados, (
<geoCoordinate id="EE5B5053F05AFFCFFEC9FDA5FE91FDB0" box="[284,375,584,608]" direction="south" orientation="latitude" pageId="56" pageNumber="57" precision="925" value="-22.216667">22°13'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F05AFFCFFE51FDAAFE0CFDB0" box="[388,490,583,608]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="925" value="-54.8">54°48'W</geoCoordinate>
),
<date id="FFD11054F05AFFCFFE2AFDA5FDA8FDB0" box="[511,590,584,608]" pageId="56" pageNumber="57" value="2010-10">x.2010</date>
, coll. V.A. Conrad—
<specimenCount id="9D69FD1DF05AFFCFFC91FDAAFC6DFDB0" box="[836,907,583,608]" pageId="56" pageNumber="57" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFC49FDA5FC16FDB0" box="[924,1008,584,608]" pageId="56" pageNumber="57">CEMT</collectionCode>
); same locality,
<date id="FFD11054F05AFFCFFB7DFDAAFAF2FDB0" box="[1192,1300,583,608]" pageId="56" pageNumber="57" value="2009-11-01">1.xi.2009</date>
, coll. J.F.A. Da Luz—
<specimenCount id="9D69FD1DF05AFFCFFEDEFD81FE81FD55" box="[267,359,620,645]" pageId="56" pageNumber="57" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFEA3FD81FE2CFD54" box="[374,458,620,644]" pageId="56" pageNumber="57">CEMT</collectionCode>
);
<collectionCode id="ED7EAE51F05AFFCFFE08FD81FDD0FD54" box="[477,566,620,644]" pageId="56" pageNumber="57">UNESP</collectionCode>
Farm [=Fazenda Experimental da Universidade Estadual Paulista, câmpus de
<collectingCountry id="F3787604F05AFFCFFF42FD62FF25FD78" box="[151,195,655,680]" name="Iceland" pageId="56" pageNumber="57">Ilha</collectingCountry>
Solteira], Selvíria, (
<geoCoordinate id="EE5B5053F05AFFCFFE7FFD7DFDFBFD78" box="[426,541,656,680]" direction="south" orientation="latitude" pageId="56" pageNumber="57" precision="15" value="-20.335556">20°20'8''S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F05AFFCFFDFEFD62FD51FD78" box="[555,695,655,680]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="15" value="-51.412224">51°24'44''W</geoCoordinate>
),
<date id="FFD11054F05AFFCFFD19FD62FCDEFD78" box="[716,824,655,680]" pageId="56" pageNumber="57" value="2010-11-06">6.xi.2010</date>
, coll. H. Wilson—
<specimenCount id="9D69FD1DF05AFFCFFBC1FD62FB97FD78" box="[1044,1137,655,680]" pageId="56" pageNumber="57" type="female">1 female</specimenCount>
(MEFEIS); same locality,
<date id="FFD11054F05AFFCFFF42FD59FEF7FD1D" box="[151,273,692,717]" pageId="56" pageNumber="57" value="2010-11-20">20.xi.2010</date>
, coll. H. Wilson—
<specimenCount id="9D69FD1DF05AFFCFFE24FD59FDDFFD1D" box="[497,569,692,717]" pageId="56" pageNumber="57" type="male">1 male</specimenCount>
(MEFEIS); same locality,
<date id="FFD11054F05AFFCFFCA4FD59FC30FD1D" box="[881,982,692,717]" pageId="56" pageNumber="57" value="2011-02-04">4.ii.2011</date>
, coll. H. Wilson—
<specimenCount id="9D69FD1DF05AFFCFFB63FD59FAF2FD1D" box="[1206,1300,692,717]" pageId="56" pageNumber="57" type="female">1 female</specimenCount>
(MEFEIS);
<collectionCode id="ED7EAE51F05AFFCFFF42FD35FF17FD20" box="[151,241,728,752]" pageId="56" pageNumber="57">MINAS</collectionCode>
GERAIS, Universidade Federal de Viçosa Campus, Viçosa, (
<geoCoordinate id="EE5B5053F05AFFCFFC71FD3AFC18FD20" box="[932,1022,727,752]" direction="south" orientation="latitude" pageId="56" pageNumber="57" precision="925" value="-20.75">20°45'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F05AFFCFFBDFFD3AFB70FD20" box="[1034,1174,727,752]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="15" value="-42.875">42°52'30''W</geoCoordinate>
),
<date id="FFD11054F05AFFCFFB7FFD3AFAE4FD20" box="[1194,1282,727,752]" pageId="56" pageNumber="57" value="1999-09">ix.1999</date>
, coll. Vaz-de- Mello &amp; Milhomem—
<specimenCount id="9D69FD1DF05AFFCFFE4BFD11FE1CFCC5" box="[414,506,764,789]" pageId="56" pageNumber="57" type="female">1 female</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFDDEFD11FDB9FCC4" box="[523,607,764,788]" pageId="56" pageNumber="57">CEMT</collectionCode>
); same locality,
<date id="FFD11054F05AFFCFFCC3FD11FC77FCC5" box="[790,913,764,789]" pageId="56" pageNumber="57" value="1995-11-18">18.xi.1995</date>
, coll. F. Vaz de Mello—
<specimenCount id="9D69FD1DF05AFFCFFB79FD11FB15FCC5" box="[1196,1267,764,789]" pageId="56" pageNumber="57" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFAD6FD11FAB1FCC4" box="[1283,1367,764,788]" pageId="56" pageNumber="57">CEMT</collectionCode>
); RIO
<collectingCountry id="F3787604F05AFFCFFF42FCCDFF5AFCE7" box="[151,188,800,823]" name="South Georgia and the South Sandwich Islands" pageId="56" pageNumber="57">DE</collectingCountry>
JANEIRO, Cordeiro, (
<geoCoordinate id="EE5B5053F05AFFCFFE01FCCDFDC6FCE8" box="[468,544,800,824]" direction="south" orientation="latitude" pageId="56" pageNumber="57" precision="925" value="-22.016666">22°1'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F05AFFCFFDE7FCCDFD7FFCE8" box="[562,665,800,824]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="925" value="-42.366665">42°22'W</geoCoordinate>
),
<date id="FFD11054F05AFFCFFD66FCF2FCE5FCE8" box="[691,771,799,824]" pageId="56" pageNumber="57" value="1989-02">ii.1989</date>
, coll. R. Salgado—
<specimenCount id="9D69FD1DF05AFFCFFC22FCF2FBA5FCE8" box="[1015,1091,799,824]" pageId="56" pageNumber="57" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFB8CFCCDFB4BFCE8" box="[1113,1197,800,824]" pageId="56" pageNumber="57">CEMT</collectionCode>
); Mendes, (
<geoCoordinate id="EE5B5053F05AFFCFFAE8FCCDFA7EFCE8" box="[1341,1432,800,824]" direction="south" orientation="latitude" pageId="56" pageNumber="57" precision="925" value="-22.516666">22°31'S</geoCoordinate>
,
<geoCoordinate id="EE5B5053F05AFFCFFF42FCA9FF1BFC8D" box="[151,253,836,861]" direction="west" orientation="longitude" pageId="56" pageNumber="57" precision="925" value="-43.75">43°45'W</geoCoordinate>
), [no date], coll. [anonymous]—
<specimenCount id="9D69FD1DF05AFFCFFDA5FCA9FD50FC8D" box="[624,694,836,861]" pageId="56" pageNumber="57" type="male">1 male</specimenCount>
(
<collectionCode id="ED7EAE51F05AFFCFFD10FCA9FCF6FC8C" box="[709,784,836,860]" pageId="56" pageNumber="57">CPFA</collectionCode>
).
</paragraph>
<paragraph id="8BD03694F05AFFCFFF12FC8AFB64FCAF" blockId="56.[151,1437,151,1616]" box="[199,1154,871,896]" pageId="56" pageNumber="57">
<emphasis id="B91BEA86F05AFFCFFF12FC8AFE9BFC50" bold="true" box="[199,381,871,896]" pageId="56" pageNumber="57">Natural history</emphasis>
. Unknown. Three specimens collected using unbaited window traps.
</paragraph>
</subSubSection>
<subSubSection id="C375651FF05AFFCFFF12FC61FB25F980" pageId="56" pageNumber="57" type="discussion">
<paragraph id="8BD03694F05AFFCFFF12FC61FAEDFC18" blockId="56.[151,1437,151,1616]" pageId="56" pageNumber="57">
<emphasis id="B91BEA86F05AFFCFFF12FC61FED2FC75" bold="true" box="[199,308,908,933]" pageId="56" pageNumber="57">Remarks</emphasis>
. Females differs in having the clypeofrontal carina much lower, approximately 4 times as wide as high and by the anterior pronotal carina simply tri-sinuate and only slightly more developed medially.
</paragraph>
<paragraph id="8BD03694F05AFFCFFF12FC39FE44FAE0" blockId="56.[151,1437,151,1616]" pageId="56" pageNumber="57">Variation is seen in the extent and intensity of the metallic sheen, as well as the color varying from greenish to dark purplish-blue. In some individuals the pronotal punctures are slightly larger and coarser. Specimens from the cerrado slightly but consistently differs in the shape of the clypeal teeth which are smaller and more obliquely oriented laterally. Females from the cerrado also have the median portion of the pronotal carina produced into an upward angular projection, the carina is simply arcuate in specimens from the Atlantic forest. The configuration of the male median pronotal projection is also variable, some individuals present a simply dorsoventrally flat projection, and in some other the anterior edge of the projection is thickened and more or less marginate dorsally. The inferior surface of the lobate median projection is produced into a more or less sharp longitudinal carina. This carina, which is very blunt and less developed in the Atlantic forest specimens is dividing the anterior concavity into two equal portions.</paragraph>
<paragraph id="8BD03694F05AFFCFFF12FAD1FE2DFAA7" blockId="56.[151,1437,151,1616]" pageId="56" pageNumber="57">
<emphasis id="B91BEA86F05AFFCFFF12FAD1FDC5FA85" bold="true" box="[199,547,1340,1365]" pageId="56" pageNumber="57">Nomenclature and taxonomy</emphasis>
.
<taxonomicName id="4C6F4D17F05AFFCFFDE6FAD0FC98FA85" authority="Lacordaire, 1856" authorityName="Lacordaire" authorityYear="1856" box="[563,894,1340,1365]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="amyntas">
<emphasis id="B91BEA86F05AFFCFFDE6FAD0FD52FA84" box="[563,692,1341,1364]" italics="true" pageId="56" pageNumber="57">D. amyntas</emphasis>
Lacordaire, 1856
</taxonomicName>
bona species =
<taxonomicName id="4C6F4D17F05AFFCFFBECFAD0FA7AFA84" authority="Olsoufieff, 1924" authorityName="Olsoufieff" authorityYear="1924" box="[1081,1436,1340,1365]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="waterhousi">
<emphasis id="B91BEA86F05AFFCFFBECFAD0FB3FFA85" box="[1081,1241,1340,1365]" italics="true" pageId="56" pageNumber="57">D. waterhousi</emphasis>
<bibRefCitation id="EFFE4B65F05AFFCFFB37FAD1FA7AFA84" author="Olsoufieff" box="[1250,1436,1340,1365]" pageId="56" pageNumber="57" refString="Olsoufieff, G. d' (1924) Les Phanaeides (Coleoptera-Lamellicornia), Famille Scarabaeidae-Tr. Coprini. Insecta, 13, 4 - 172, 15 pls." type="journal article" year="1924">Olsoufieff, 1924</bibRefCitation>
</taxonomicName>
(auctores),
<emphasis id="B91BEA86F05AFFCFFEC0FAB2FE22FAA8" bold="true" box="[277,452,1375,1400]" pageId="56" pageNumber="57">new synonymy</emphasis>
.
</paragraph>
<paragraph id="8BD03694F05AFFCFFF12FA69FB25F980" blockId="56.[151,1437,151,1616]" pageId="56" pageNumber="57">
Lacordaires (1856) description and illustration (
<typeStatus id="54D48836F05AFFCFFD3CFA69FCA8FA4D" box="[745,846,1412,1437]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
) of this species were considered invalid by previous authors. The description alone is too vague to identify the species, however the illustration is very accurate and clearly shows one of the diagnostic characters of this species. The clypeal teeth are clearly defined in the drawing and correspond to the species that is found in the cerrado and Atlantic forest of
<collectingCountry id="F3787604F05AFFCFFBDDFA02FBB6F9D8" box="[1032,1104,1519,1544]" name="Brazil" pageId="56" pageNumber="57">Brazil</collectingCountry>
. This species was redescribed as
<taxonomicName id="4C6F4D17F05AFFCFFF6EF9F8FEB8F9FD" box="[187,350,1556,1581]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="waterhousi">
<emphasis id="B91BEA86F05AFFCFFF6EF9F8FEB8F9FD" box="[187,350,1556,1581]" italics="true" pageId="56" pageNumber="57">D. waterhousi</emphasis>
</taxonomicName>
by dOlsoufieff in 1924. Harold (1869) lists Lacordaires name but failed to recognize the description as valid, which seems to have started the confusion over the identity of this species.
</paragraph>
</subSubSection>
</treatment>
</document>