201 lines
24 KiB
XML
201 lines
24 KiB
XML
<document id="5B3C619D908534A6637F80959180F18F" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier & Patrick Arnaud" docDate="2016" docId="03C68782F051FFC3FF42FC39FB15FA4F" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Glaphyropaemon) inemarginatus Genier & Arnaud, new species" docType="treatment" docVersion="10" lastPageNumber="53" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="52" updateTime="1698234910999" updateUser="plazi">
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<mods:title id="558F6946EF481044CE6BE2E01954D51A">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
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<mods:namePart id="B5F60AABA23469EA646BAC2A6A5F47F3">François Génier</mods:namePart>
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<mods:roleTerm id="CD92BB7D7C2B675B4DC87BD42897709A">Author</mods:roleTerm>
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<mods:namePart id="44390623B5232366E13F3081E1DAECF4">Patrick Arnaud</mods:namePart>
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<mods:typeOfResource id="D140320FDCB5BEF1DEA968C7B0C7A87E">text</mods:typeOfResource>
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<mods:title id="85A2C9B3B57AAFF764E9C203025C9ABC">Zootaxa</mods:title>
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<mods:part id="3F00C28D9EF1C8637CB40684DE17CC9F">
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<mods:date id="CC43A8F4CF8BEC7F2DBAC097247E0204">2016</mods:date>
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<mods:detail id="AB199F3DEBA18384F8DB669E67EDAA24" type="volume">
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<mods:number id="9D1B95B69D3DAEB28DFCA5179435A627">4099</mods:number>
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<mods:detail id="34929EAA47E7A9037565F9F199D30134" type="issue">
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<mods:number id="6C0211B22FC51B536179AECD5A1ACFE9">1</mods:number>
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<mods:classification id="B6E156A4DAFD5287E2B0FD9EDFB71D13">journal volume</mods:classification>
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<mods:identifier id="AB0C28B0A78397A8BD1E6756167DFF95" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
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<treatment id="03C68782F051FFC3FF42FC39FB15FA4F" ID-DOI="http://doi.org/10.5281/zenodo.5780314" ID-GBIF-Taxon="127631965" ID-Zenodo-Dep="5780314" LSID="urn:lsid:plazi:treatment:03C68782F051FFC3FF42FC39FB15FA4F" httpUri="http://treatment.plazi.org/id/03C68782F051FFC3FF42FC39FB15FA4F" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
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<subSubSection id="C375651FF051FFC4FF42FC39FB41FC3E" box="[151,1191,980,1006]" pageId="51" pageNumber="52" type="nomenclature">
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<paragraph id="8BD03694F051FFC4FF42FC39FB41FC3E" blockId="51.[151,1191,980,1039]" box="[151,1191,980,1006]" pageId="51" pageNumber="52">
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<heading id="D09881F8F051FFC4FF42FC39FB41FC3E" bold="true" box="[151,1191,980,1006]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
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<emphasis id="B91BEA86F051FFC4FF42FC39FB41FC3E" bold="true" box="[151,1191,980,1006]" pageId="51" pageNumber="52">
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25.
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<taxonomicName id="4C6F4D17F051FFC4FF16FC39FBECFC3E" ID-CoL="6CLD4" authority="Genier & Arnaud" authorityName="Genier & Arnaud" box="[195,1034,980,1006]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="inemarginatus" status="sp. nov." subGenus="Glaphyropaemon">
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<emphasis id="B91BEA86F051FFC4FF16FC39FCC4FC3E" bold="true" box="[195,802,980,1006]" italics="true" pageId="51" pageNumber="52">Dendropaemon (Glaphyropaemon) inemarginatus</emphasis>
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Génier & Arnaud
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22857FDF051FFC4FBC2FC39FB41FC3E" box="[1047,1191,980,1006]" pageId="51" pageNumber="52" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C375651FF051FFC4FF42FC1BFED6FBDF" box="[151,304,1014,1039]" pageId="51" pageNumber="52" type="description">
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<paragraph id="8BD03694F051FFC4FF42FC1BFED6FBDF" blockId="51.[151,1191,980,1039]" box="[151,304,1014,1039]" pageId="51" pageNumber="52">
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(
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<figureCitation id="13542A11F051FFC4FF4AFC1BFF14FBDF" box="[159,242,1014,1039]" captionStart="FIGURES 25 – 30" captionStartId="103.[151,264,1962,1985]" captionTargetBox="[269,1325,201,1926]" captionTargetId="figure@103.[251,1336,193,1941]" captionTargetPageId="103" captionText="FIGURES 25 – 30. Dorsal habitus. 25. D. inemarginatus, female holotype; 26. D. nigritulus, male holotype; 27. D. amyntas, male; 28. D. attalus, male holotype; 29. D. lydiae, male holotype; 30. D. morettoi, male holotype." httpUri="https://zenodo.org/record/268279/files/figure.png" pageId="51" pageNumber="52">Fig. 25</figureCitation>
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,
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<figureCitation id="13542A11F051FFC4FF28FC1BFECEFBDF" box="[253,296,1014,1039]" captionStart="FIGURE 158" captionStartId="118.[151,250,1248,1270]" captionTargetBox="[150,1436,193,1226]" captionTargetId="figure@118.[150,1436,193,1227]" captionTargetPageId="118" captionText="FIGURE 158. Known distribution of D. convexus, D. fredericki, D. angustipennis, D. bahianus, D. inemarginatus, D. nigritulus." httpUri="https://zenodo.org/record/268288/files/figure.png" pageId="51" pageNumber="52">158</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C375651FF051FFC4FF42FBD3FBA0FB84" box="[151,1094,1086,1109]" pageId="51" pageNumber="52" type="reference_group">
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<paragraph id="8BD03694F051FFC4FF42FBD3FBA0FB84" blockId="51.[151,1094,1086,1109]" box="[151,1094,1086,1109]" pageId="51" pageNumber="52">
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<taxonomicName id="4C6F4D17F051FFC4FF42FBD3FD32FB84" ID-CoL="34R2Y" authority="Martinez & Clavijo 1990" authorityName="Martinez & Clavijo" authorityYear="1990" box="[151,724,1086,1108]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="refulgens" subGenus="Dendropaemon">
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<emphasis id="B91BEA86F051FFC4FF42FBD3FE20FB84" box="[151,454,1086,1108]" italics="true" pageId="51" pageNumber="52">Dendropaemon (D.) refulgens</emphasis>
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:
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<bibRefCitation id="EFFE4B65F051FFC4FE01FBD3FD32FB84" author="Martinez" box="[468,724,1086,1108]" pageId="51" pageNumber="52" refString="Martinez, A. & Clavijo, J. (1990) Notas sobre Phanaeina Venezolanos, con descripcion de una nueva subespecie de Diabroctis (Coleoptera, Scarabaeidae, Coprini). Boletin de Entomologia Venezolana, Nueva Seria, 5 (20) 147 - 157." type="journal article" year="1990">
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Martínez & Clavijo
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<emphasis id="B91BEA86F051FFC4FD74FBD2FD32FB84" bold="true" box="[673,724,1087,1108]" pageId="51" pageNumber="52">1990</emphasis>
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</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="B91BEA86F051FFC4FD35FBD3FC7AFB84" box="[736,924,1086,1109]" italics="true" pageId="51" pageNumber="52">Bol. Ent. Ven. N.S.</emphasis>
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5: 155 (biology)
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</paragraph>
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</subSubSection>
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<subSubSection id="C375651FF051FFC4FF42FB6CFB60FB4A" box="[151,1158,1153,1178]" pageId="51" pageNumber="52" type="materials_examined">
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<paragraph id="8BD03694F051FFC4FF42FB6CFB60FB4A" blockId="51.[151,1436,1153,2006]" box="[151,1158,1153,1178]" pageId="51" pageNumber="52">
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<emphasis id="B91BEA86F051FFC4FF42FB6CFECBFB4A" bold="true" box="[151,301,1153,1178]" pageId="51" pageNumber="52">
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<typeStatus id="54D48836F051FFC4FF42FB6CFF34FB4A" box="[151,210,1153,1178]" pageId="51" pageNumber="52">Type</typeStatus>
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locality
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</emphasis>
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. Coromoto, Departamento Atures, Territorio Federal Amazonas,
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<collectingCountry id="F3787604F051FFC4FBDEFB6CFB64FB4A" box="[1035,1154,1153,1178]" name="Venezuela" pageId="51" pageNumber="52">Venezuela</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C375651FF051FFC4FF12FB48FC93FB32" pageId="51" pageNumber="52" type="diagnosis">
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<paragraph id="8BD03694F051FFC4FF12FB48FC93FB32" blockId="51.[151,1436,1153,2006]" pageId="51" pageNumber="52">
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<emphasis id="B91BEA86F051FFC4FF12FB48FEA7FB6D" bold="true" box="[199,321,1189,1214]" pageId="51" pageNumber="52">Diagnosis.</emphasis>
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The only species in the genus with a vestigial posterior pronotal margin. In some species the margin is reduced but always visible on a distance laterally and medially.
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</paragraph>
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</subSubSection>
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<subSubSection id="C375651FF051FFC3FF12FB00FD19FBAF" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
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<paragraph id="8BD03694F051FFC3FF12FB00FE7BFC18" blockId="51.[151,1436,1153,2006]" lastBlockId="52.[151,1437,151,1440]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
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<emphasis id="B91BEA86F051FFC4FF12FB00FEB7FAD6" bold="true" box="[199,337,1261,1286]" pageId="51" pageNumber="52">Description</emphasis>
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. Female
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<typeStatus id="54D48836F051FFC4FE6EFB00FDFBFAD6" box="[443,541,1261,1286]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
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(
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<figureCitation id="13542A11F051FFC4FDFBFB00FD65FAD6" box="[558,643,1261,1286]" captionStart="FIGURES 25 – 30" captionStartId="103.[151,264,1962,1985]" captionTargetBox="[269,1325,201,1926]" captionTargetId="figure@103.[251,1336,193,1941]" captionTargetPageId="103" captionText="FIGURES 25 – 30. Dorsal habitus. 25. D. inemarginatus, female holotype; 26. D. nigritulus, male holotype; 27. D. amyntas, male; 28. D. attalus, male holotype; 29. D. lydiae, male holotype; 30. D. morettoi, male holotype." httpUri="https://zenodo.org/record/268279/files/figure.png" pageId="51" pageNumber="52">Fig. 25</figureCitation>
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).
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<emphasis id="B91BEA86F051FFC4FD42FB00FD32FAD6" bold="true" box="[663,724,1261,1286]" pageId="51" pageNumber="52">Body</emphasis>
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. Body moderately large, length 13.0 mm, maximum width
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<quantity id="4C979B71F051FFC4FAACFB00FF20FAF9" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="51" pageNumber="52" unit="mm" value="6.5">6.5 mm</quantity>
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; body subrectangular in dorsal view; dorsum slightly convex.
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<emphasis id="B91BEA86F051FFC4FC76FAFCFC02FAFA" bold="true" box="[931,996,1297,1322]" pageId="51" pageNumber="52">Color</emphasis>
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. Dorsal surface dark brown to black, glossy, with green metallic sheen; head with green metallic sheen on posterior portion of clypeus, genae and frons; pronotum with green metallic sheen except for anteromedian carina, on anterior portion of disc and surface adjacent to lateral fossae; elytra with uniform green metallic sheen; ventrum reddish brown to dark brown; pygidium with green metallic sheen; legs reddish brown to dark brown.
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<emphasis id="B91BEA86F051FFC4FC07FA4CFBF4FA6A" bold="true" box="[978,1042,1441,1466]" pageId="51" pageNumber="52">Head</emphasis>
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. Clypeus broadly arcuate, anterior portion upturned; clypeal teeth ogival; clypeal median emargination narrowly v-shaped, clypeal edge acutely notched on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin illdefined, lacking sharp carina posteriorly, clypeal surface with transverse blunt rugulae; clypeogenal suture welldefined, bluntly carinate internally; genal surface irregularly punctate, transversally and shallowly sulcate on posterior half; clypeofrontal carina rather low, approximately 4 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge straight in frontal view; eyes small in dorsal view, interocular ratio 5.8.
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<emphasis id="B91BEA86F051FFC4FE47F970FDECF966" bold="true" box="[402,522,1693,1718]" pageId="51" pageNumber="52">Pronotum</emphasis>
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. Pronotum transverse in dorsal view, pronotal width/length ratio 1.4; disc of pronotum minutely punctate basally, puncture becoming fine anteriorly, with an ill-defined shallow longitudinal depression on posterior half; pronotal anterior margin wider and flat lateral to eyes; anterior portion with a trisinuous carina, carina produced into a tubercle medially; anterior angles surface simply punctate, slightly but distinctly sulcate along posterior edge of anterior margin; lateral fossae simply rounded, concave; lateral portions unmodified; pronotal basal fossae ill-defined, slightly concave; posterior margin vestigial, reduced to a minute interrupted carina throughout.
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<emphasis id="B91BEA86F051FFC4FE3EF898FDD1F85E" bold="true" box="[491,567,1909,1934]" pageId="51" pageNumber="52">Elytra</emphasis>
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. Elytra approximately as long as wide in dorsal view, elytral combined width/ length ratio 1.1; elytral base lacking distinct margin, simply convex; elytral striae 1–4 moderately wide basally and fine apically, evenly impressed throughout, elytral striae 5 atrophied, lacking fine carina on each side on disc, strial punctures fine and well-defined, adjacent strial edge encroaching on interval, stria 1 well-defined apically, connecting to marginal stria; interstriae slightly convex, minutely punctate throughout, surface glossy.
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<emphasis id="B91BEA86F056FFC3FAE4FF51FF04FF28" bold="true" pageId="52" pageNumber="53">Thoracic sterna</emphasis>
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. Proepisternal carina complete, extending laterally; metasternal median lobe angularly produced anteromedially, ventral ridge well-defined, v-shaped.
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<emphasis id="B91BEA86F056FFC3FCDBFEE9FCA0FECD" bold="true" box="[782,838,260,285]" pageId="52" pageNumber="53">Legs</emphasis>
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. Profemur posterior surface slightly but distinctly convex, rather coarsely punctate and glabrous internally, posterointernal margin rather thin, evenly developed, internal edge rather wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with rugose irregular punctures externally to median carina, surface finely and irregularly microsculptured between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge slightly sinuate in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less rounded, with several large elongate setiferous punctures. Mesotarsus similar in shape to metatarsus, 3-segmented, first segment short, approximately as long as wide at apex. Metafemur broadly oval in anterior view, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on more than half the length. Metatibia moderately slender, slightly widening toward apex in anterior view, anterior surface with distinct row of setae, with fine irregular punctures on a glossy surface, metatibial posterior surface flat between longitudinal row of setae and lateral edge, glossy between punctures. Metatarsus 3-segmented, first segment short, approximately as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge.
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<emphasis id="B91BEA86F056FFC3FAC3FCA9FF1AFC50" bold="true" pageId="52" pageNumber="53">Abdominal sternites</emphasis>
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. Sternites 3–6 longitudinally flat; sternites 4–6 with 1–3 unaligned rows of setae laterally, glabrous medially; sternite 7 approximately longitudinally flat medially, shorter than segment 6 along midline; pygidium finely punctate on disc.
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</paragraph>
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<paragraph id="8BD03694F056FFC3FF12FC39FD2EFC3C" blockId="52.[151,1437,151,1440]" box="[199,712,980,1005]" pageId="52" pageNumber="53">
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<emphasis id="B91BEA86F056FFC3FF12FC39FE90FC3D" bold="true" box="[199,374,980,1005]" pageId="52" pageNumber="53">Measurements</emphasis>
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(
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<specimenCount id="9D69FD1DF056FFC3FE5DFC39FE0FFC3D" box="[392,489,980,1005]" pageId="52" pageNumber="53" type="female">1 female</specimenCount>
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). Length: 13.0 mm.
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</paragraph>
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<paragraph id="8BD03694F056FFC3FF12FC17FD19FBAF" blockId="52.[151,1437,151,1440]" pageId="52" pageNumber="53">
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<emphasis id="B91BEA86F056FFC3FF12FC17FE55FBC3" bold="true" box="[199,435,1018,1043]" pageId="52" pageNumber="53">
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Primary
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<typeStatus id="54D48836F056FFC3FEE9FC17FE89FBC3" box="[316,367,1018,1043]" pageId="52" pageNumber="53">type</typeStatus>
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data
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</emphasis>
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. Female
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<typeStatus id="54D48836F056FFC3FDF9FC17FD68FBC3" box="[556,654,1018,1043]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
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(CMNC): [Marz-979/
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<collectingCountry id="F3787604F056FFC3FC7CFC16FBA9FBC3" box="[937,1103,1019,1043]" name="Venezuela" pageId="52" pageNumber="53">VENEZUELA</collectingCountry>
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/ T.F.Amazonas/ D o Atures/ Coromoto/ G.y H.Martínez-leg./ Coll. A. Martínez] handwritten; [H. & A. HOWDEN/ COLLECTION/ ex. A. Martínez coll.]; [WORLD/ SCARAB./ DATABASE/ WSD00016731]; [
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<typeStatus id="54D48836F056FFC3FBDEFBAEFB44FB8B" box="[1035,1186,1091,1115]" pageId="52" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
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♀/
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<taxonomicName id="4C6F4D17F056FFC3FB32FBAFFA72FB8B" box="[1255,1428,1090,1115]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
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<emphasis id="B91BEA86F056FFC3FB32FBAFFA72FB8B" box="[1255,1428,1090,1115]" italics="true" pageId="52" pageNumber="53">Dendropaemon</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C6F4D17F056FFC3FF42FB85FEDEFBAF" box="[151,312,1128,1151]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="inemarginatus" status="sp. nov.">
|
||
<emphasis id="B91BEA86F056FFC3FF42FB85FEDEFBAF" box="[151,312,1128,1151]" italics="true" pageId="52" pageNumber="53">inemarginatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22857FDF056FFC3FEEBFB8AFE97FBAF" box="[318,369,1127,1151]" pageId="52" pageNumber="53" rank="species">n.sp.</taxonomicNameLabel>
|
||
/ Génier & Arnaud, 2014] red card.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF056FFC3FF12FB67FD61FB73" box="[199,647,1162,1187]" pageId="52" pageNumber="53" type="materials_examined">
|
||
<paragraph id="8BD03694F056FFC3FF12FB67FD61FB73" blockId="52.[151,1437,151,1440]" box="[199,647,1162,1187]" pageId="52" pageNumber="53">
|
||
<emphasis id="B91BEA86F056FFC3FF12FB67FE4FFB73" bold="true" box="[199,425,1162,1187]" pageId="52" pageNumber="53">Material examined</emphasis>
|
||
. Primary
|
||
<typeStatus id="54D48836F056FFC3FDCDFB66FDAEFB73" box="[536,584,1163,1187]" pageId="52" pageNumber="53">type</typeStatus>
|
||
only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF056FFC3FF12FB43FDDEFAE3" pageId="52" pageNumber="53" type="etymology">
|
||
<paragraph id="8BD03694F056FFC3FF12FB43FEB4FB3B" blockId="52.[151,1437,151,1440]" pageId="52" pageNumber="53">
|
||
<emphasis id="B91BEA86F056FFC3FF12FB43FEA1FB17" bold="true" box="[199,327,1198,1223]" pageId="52" pageNumber="53">Etymology</emphasis>
|
||
.
|
||
<taxonomicName id="4C6F4D17F056FFC3FE8CFB5DFE1DFB17" box="[345,507,1200,1223]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="119" phylum="Arthropoda" rank="species" species="inemarginatus">
|
||
<emphasis id="B91BEA86F056FFC3FE8CFB5DFE1DFB17" box="[345,507,1200,1223]" italics="true" pageId="52" pageNumber="53">Inemarginatus</emphasis>
|
||
</taxonomicName>
|
||
, without + margin, an adjective referring to the configuration of the posterior pronotal margin.
|
||
</paragraph>
|
||
<paragraph id="8BD03694F056FFC3FF12FB1BFDDEFAE3" blockId="52.[151,1437,151,1440]" pageId="52" pageNumber="53">
|
||
<emphasis id="B91BEA86F056FFC3FF12FB1BFE65FADF" bold="true" box="[199,387,1270,1295]" pageId="52" pageNumber="53">Natural history</emphasis>
|
||
. The
|
||
<typeStatus id="54D48836F056FFC3FE19FB1BFDC8FADF" box="[460,558,1270,1295]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
|
||
was collected in a zone of loose and sandy soil in the early hours of the morning (
|
||
<bibRefCitation id="EFFE4B65F056FFC3FED1FAF6FDCDFAE3" author="Martinez" box="[260,555,1306,1331]" pageId="52" pageNumber="53" refString="Martinez, A. & Clavijo, J. (1990) Notas sobre Phanaeina Venezolanos, con descripcion de una nueva subespecie de Diabroctis (Coleoptera, Scarabaeidae, Coprini). Boletin de Entomologia Venezolana, Nueva Seria, 5 (20) 147 - 157." type="journal article" year="1990">Martínez & Clavijo, 1990</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF056FFC3FF12FAD3FB15FA4F" pageId="52" pageNumber="53" type="discussion">
|
||
<paragraph id="8BD03694F056FFC3FF12FAD3FD6AFA87" blockId="52.[151,1437,151,1440]" box="[199,652,1342,1367]" pageId="52" pageNumber="53">
|
||
<emphasis id="B91BEA86F056FFC3FF12FAD3FED2FA87" bold="true" box="[199,308,1342,1367]" pageId="52" pageNumber="53">Remarks</emphasis>
|
||
.
|
||
<collectingCountry id="F3787604F056FFC3FE97FAD3FE9AFA87" box="[322,380,1342,1367]" name="Malta" pageId="52" pageNumber="53">Male</collectingCountry>
|
||
and variation unknown.
|
||
</paragraph>
|
||
<paragraph id="8BD03694F056FFC3FF12FA8FFB15FA4F" blockId="52.[151,1437,151,1440]" pageId="52" pageNumber="53">
|
||
Superficially resembling species of
|
||
<taxonomicName id="4C6F4D17F056FFC3FDBFFA8FFCD8FAAB" box="[618,830,1378,1403]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91BEA86F056FFC3FDBFFA8FFCD8FAAB" box="[618,830,1378,1403]" italics="true" pageId="52" pageNumber="53">Coprophanaeoides</emphasis>
|
||
</taxonomicName>
|
||
, however, the presence of a complete proepisternal carina place this species within
|
||
<emphasis id="B91BEA86F056FFC3FE2FFA6BFD0FFA4F" box="[506,745,1414,1439]" italics="true" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C6F4D17F056FFC3FE2FFA6BFD41FA4F" box="[506,679,1414,1439]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus" sensu="stricto">Dendropaemon</taxonomicName>
|
||
<taxonomicNameLabel id="A22857FDF056FFC3FD7BFA65FD0FFA4F" box="[686,745,1416,1439]" pageId="52" pageNumber="53" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</emphasis>
|
||
where it seems to occupy an isolated position.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |