treatments-xml/data/03/A8/D7/03A8D74FFF88FFE5C9C2FE4E40F3F8CD.xml

<document id="8A7CAC8AE50F9788B5136F9BA99656E1" ID-CLB-Dataset="29235" ID-DOI="10.11646/zootaxa.4497.1.1" ID-GBIF-Dataset="d5b94fdb-db02-4530-85aa-12d9757e3313" ID-ISSN="1175-5326" ID-Zenodo-Dep="1451056" ID-ZooBank="890E26F1-1B03-4F13-9299-A5E8BED8439D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538973324849" checkinUser="plazi" docAuthor="Grehan, John R. &amp; Mielke, Os. G. C." docDate="2018" docId="03A8D74FFF88FFE5C9C2FE4E40F3F8CD" docLanguage="en" docName="zootaxa.4497.1.1.pdf" docOrigin="Zootaxa 4497 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pfitzneriella yuliyakovalevae ♂, Grehan &amp; C. Mielke 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FF91AF37FF98FFF4C955FF924474FFBB" masterDocTitle="New species of Dugdaleiella, gen. nov., Kozloviella, gen. nov., and Pfitzneriella Viette from upper elevation Andes of Ecuador and Peru (Lepidoptera: Hepialidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="17" updateTime="1698567105610" updateUser="ExternalLinkService">
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<mods:title id="7C0A6EC7DF182BD9B3E1F42A05B1BBF1">New species of Dugdaleiella, gen. nov., Kozloviella, gen. nov., and Pfitzneriella Viette from upper elevation Andes of Ecuador and Peru (Lepidoptera: Hepialidae)</mods:title>
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<mods:namePart id="387A07B6FF7B72FD41434B9C3CB6DF5F">Grehan, John R.</mods:namePart>
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<mods:namePart id="974176C0428B478B296879A0B856340A">Mielke, Os. G. C.</mods:namePart>
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<mods:date id="C8E97A670E430604CFAB45EBBEE06DC4">2018</mods:date>
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<mods:number id="A378A7B677AD685D9E2E874C2981C810">2018-10-08</mods:number>
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<subSubSection id="C31B35D2FF88FFE4C9C2FE4E4021FDAC" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BBE6659FF88FFE4C9C2FE4E4648FE4D" blockId="16.[151,1109,476,535]" box="[151,572,476,502]" pageId="16" pageNumber="17">
<heading id="D0F6D135FF88FFE4C9C2FE4E4648FE4D" bold="true" box="[151,572,476,502]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B975BA4BFF88FFE4C9C2FE4E4648FE4D" bold="true" box="[151,572,476,502]" pageId="16" pageNumber="17">
<taxonomicName id="4C011DDAFF88FFE4C9C2FE4E4586FE4D" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[151,498,476,502]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="yuliyakovalevae" status="sp. nov.">
<emphasis id="B975BA4BFF88FFE4C9C2FE4E4586FE4D" bold="true" box="[151,498,476,502]" italics="true" pageId="16" pageNumber="17">Pfitzneriella yuliyakovalevae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF88FFE4C8AFFE4E4648FE4D" box="[506,572,476,502]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
</paragraph>
<paragraph id="8BBE6659FF88FFE4C9C2FE6C4021FDAC" blockId="16.[151,1109,476,535]" box="[151,1109,510,535]" pageId="16" pageNumber="17">
(
<figureCitation id="133A7ADCFF88FFE4C9CAFE6C4505FDAC" box="[159,369,510,535]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="16" pageNumber="17">Figs 7a, 7b, 7c, 7d</figureCitation>
, 
<figureCitation id="133A7ADCFF88FFE4C829FE6D45CBFDAC" box="[380,447,510,535]" captionStart="FIGURES 8–9" captionStartId="5.[151,264,1506,1529]" captionTargetBox="[151,1435,193,1486]" captionTargetId="figure@5.[151,1435,193,1486]" captionTargetPageId="5" captionText="FIGURES 8–9. Voucher specimens. 8, Dugdaleiella monticola male. 8a, LT (ZMHB); 8b, Cotopaxi, Ecuador (CMNH); 8c, Volcán Chiles, Ecuador (CMNH); 8d, male, Volcán Chiles, Ecuador (CMNH). 9, Labial palpi (male unless otherwise specified), 9a, Kozloviella viazmenskyi sp. n.; 9b, Pfitzneriella antonkozlovi sp. n.; 9c, P. rawlinsi sp. n.; 9d, P. titarenkoi sp. n.; 9e, P. yuliyakovalevae sp. n.; 9f, P. yuliyakovalevae sp. n. (female); 9g, Dugdaleiella monticola. Photos: Jane Hyland (Fig. 8), John Grehan (Figs 9a–b, d–h), and Carlos Mielke (Fig. 9c). Scale 1 mm (when present)." httpUri="https://zenodo.org/record/1451060/files/figure.png" pageId="16" pageNumber="17">9e, 9f</figureCitation>
, 
<figureCitation id="133A7ADCFF88FFE4C89FFE6D465BFDAC" box="[458,559,510,535]" captionStart="FIGURE 10" captionStartId="6.[151,250,1649,1671]" captionTargetBox="[152,1437,195,1610]" captionTargetId="figure@6.[152,1437,195,1610]" captionTargetPageId="6" captionText="FIGURE 10. Wing venation (male unless otherwise specified). 10a, Kozloviella viazmenskyi sp. n.; 10b, P. antonkozlovi sp. n.; 10c, P. olafi sp. n.; 10d, P. rawlinsi sp. n.; 10e, P. remota (from photo only, venation incomplete due to damaged condition of HT); 10f, P. yuliyakovalevae sp. n.; 10g, P. yuliyakovalevae sp. n. (female); 10h, Dugdaleiella monticola." httpUri="https://zenodo.org/record/1451062/files/figure.png" pageId="16" pageNumber="17">
<quantity id="4CF9CBBCFF88FFE4C89FFE6D4582FDAC" box="[458,502,511,535]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="16" pageNumber="17" unit="g" value="10.0">10g</quantity>
, 10h
</figureCitation>
, 
<figureCitation id="133A7ADCFF88FFE4CB6FFE6C4614FDAC" box="[570,608,510,535]" captionStart="FIGURE 11" captionStartId="7.[151,250,1429,1451]" captionTargetBox="[150,1436,810,1407]" captionTargetId="figure@7.[150,1436,810,1407]" captionTargetPageId="7" captionText="FIGURE 11. Wing markings. 11a; Kozloviella viazmenskyi sp. n., 11b; Pfitzneriella antonkozlovi sp. n.; 11c, P. olafi sp. n.; 11d, P. rawlinsi sp. n.; 11e, P. remota; 11f, P. yuliyakovalevae sp. n. Blue line—edge of white banding." httpUri="https://zenodo.org/record/1451064/files/figure.png" pageId="16" pageNumber="17">11f</figureCitation>
, 
<figureCitation id="133A7ADCFF88FFE4CB39FE6D46BBFDAC" box="[620,719,510,535]" captionStart="FIGURES 12–14. 12" captionStartId="9.[151,264,1172,1195]" captionTargetBox="[150,1436,193,1151]" captionTargetId="figure@9.[150,1436,193,1151]" captionTargetPageId="9" captionText="FIGURES 12–14. 12, Antenna. Pfitzneriella rawlinsi sp. n. (unscaled). 13, Legs. 13a, Kozloviella viazmenskyi sp. n.; 13b, P. olafi sp. n.; 13c, P. rawlinsi sp. n.; 13d, P. titarenkoi sp. n.; 13e, P. yuliyakovalevae sp. n. 14, Metatibia. 14a, P. antonkozlovi sp. n.; 14b, P. rawlinsi sp. n.; 14c, P. titarenkoi; 14d, P. yuliyakovalevae sp. n.; 14e, P. antonkozlovi sp. n.; 14f, Dugdaleiella monticola. Photos: John Grehan (Figs 12, 14e, 14f), Carlos Mielke (Figs 13,14a–d). Scale 1 mm." httpUri="https://zenodo.org/record/1451066/files/figure.png" pageId="16" pageNumber="17">13e, 14d</figureCitation>
, 
<figureCitation id="133A7ADCFF88FFE4CB8EFE6C4705FDAC" box="[731,881,510,535]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[197,1386,199,1851]" captionTargetId="figure@10.[197,1386,199,1851]" captionTargetPageId="10" captionText="FIGURES 15–16. Tergosternal connection. 15a, Kozloviella viazmenskyi sp. n.; 15b, Pfitzneriella yuliyakovalevae sp. n.; 15c, P. rawlinsi sp. n.; 15d, Dugdaleiella monticola. 16, Abdomen (males unless otherwise specified). 16a, K. viazmenskyi sp. n.; 16b, P. antonkozlovi sp. n.; 16c, P. olafi sp. n.; 16d, P. rawlinsi sp. n.; 16e, P. titarenkoi sp. n.; 16f, P. yuliyakovalevae sp. n.; 16g, P. yuliyakovalevae sp. n. (female); 16h–16j, Dugdaleiella monticola. 16h, tergum II; 16i, sternum II; 16j, tergum and sternum VIII. Photos: John Grehan (Figs 15d, 16h–16j), Carlos Mielke (Figs 15a–15c, 16a–16g)." httpUri="https://zenodo.org/record/1451068/files/figure.png" pageId="16" pageNumber="17">
15c, 16f, 
<quantity id="4CF9CBBCFF88FFE4CA10FE6C4705FDAC" box="[837,881,510,535]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="16" pageNumber="17" unit="g" value="16.0">16g</quantity>
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, 
<figureCitation id="133A7ADCFF88FFE4CA28FE6D4038FDAC" box="[893,1100,510,535]" captionStart="FIGURES 17–25" captionStartId="11.[151,264,1644,1667]" captionTargetBox="[151,1435,193,1629]" captionTargetId="figure@11.[151,1435,193,1629]" captionTargetPageId="11" captionText="FIGURES 17–25. Genitalia (male unless otherwise specified). 17, Kozloviella viazmenskyi sp. n. HT. 17a, ventral; 17b, lateral. 18, Pfitzneriella antonkozlovi sp. n. HT, ventral. 19, P. olafi sp. n. HT, ventral. 20, P. rawlinsi sp. n. HT, ventral. 21, P. remota HT (redrawn from Viette (1951a)). 22, P. titarenkoi sp. n. HT, ventral. 23, P. yuliyakovalevae sp. n. 23a, ventral; 23b, lateral. 24, P. yuliyakovalevae sp. n. PT female. 24a, posterior; 24b, lateral. 25, Dugdaleiella monticola. 25a, LT (ZMHB); 25b, Cotopaxi, Parque Nacional Cotopaxi. Dissection JRG 136 (CMNH). sa = saccus, fs = fultura superior, fi = fultura inferior, as = apodemal suture. Scale 1 mm (when present). Photos: Carlos Mielke (Figs 17–20, 22–25b)." httpUri="https://zenodo.org/record/1451070/files/figure.png" pageId="16" pageNumber="17">23a, 23b, 24a, 24b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF88FFE4C9C2FDD446A8FC3E" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BBE6659FF88FFE4C9C2FDD446A8FC3E" blockId="16.[151,1437,582,2022]" pageId="16" pageNumber="17">
<emphasis id="B975BA4BFF88FFE4C9C2FDD4457CFDE4" bold="true" box="[151,264,582,607]" pageId="16" pageNumber="17">Diagnosis</emphasis>
. Male FW pattern of pale markings on grayish brown similar to 
<taxonomicName id="4C011DDAFF88FFE4CAB0FDD540E5FDE4" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[997,1169,582,607]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="antonkozlovi" status="sp. nov.">
<emphasis id="B975BA4BFF88FFE4CAB0FDD540E5FDE4" box="[997,1169,582,607]" italics="true" pageId="16" pageNumber="17">P. antonkozlovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF88FFE4CDCCFDD440ACFDE5" box="[1177,1240,582,607]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B975BA4BFF88FFE4CDCCFDD440C7FDE4" bold="true" box="[1177,1203,582,607]" pageId="16" pageNumber="17">sp</emphasis>
. 
<emphasis id="B975BA4BFF88FFE4CD97FDD440A5FDE4" bold="true" box="[1218,1233,582,607]" pageId="16" pageNumber="17">n</emphasis>
.
</taxonomicNameLabel>
, but basal region with more extensive dark area enclosing yellowish white spot along CuA2, and adjacent white spot with narrow brown edge between CuA2 and CuP. The darker grayish brown FW color tone differs from the yellowish brown of 
<taxonomicName id="4C011DDAFF88FFE4C9C2FD2B4492FD6A" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[151,230,696,721]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="olafi" status="sp. nov.">
<emphasis id="B975BA4BFF88FFE4C9C2FD2B4492FD6A" box="[151,230,696,721]" italics="true" pageId="16" pageNumber="17">P. olafi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF88FFE4C9BEFD2A455DFD6B" box="[235,297,696,721]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B975BA4BFF88FFE4C9BEFD2A4571FD6A" bold="true" box="[235,261,696,721]" pageId="16" pageNumber="17">sp</emphasis>
. 
<emphasis id="B975BA4BFF88FFE4C846FD2A4556FD6A" bold="true" box="[275,290,696,721]" pageId="16" pageNumber="17">n</emphasis>
.
</taxonomicNameLabel>
, 
<taxonomicName id="4C011DDAFF88FFE4C862FD2B45DAFD6A" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[311,430,696,721]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rawlinsi" status="sp. nov.">
<emphasis id="B975BA4BFF88FFE4C862FD2B45DAFD6A" box="[311,430,696,721]" italics="true" pageId="16" pageNumber="17">P. rawlinsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF88FFE4C8E1FD2A4586FD6B" box="[436,498,696,721]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B975BA4BFF88FFE4C8E1FD2A45BAFD6A" bold="true" box="[436,462,696,721]" pageId="16" pageNumber="17">sp</emphasis>
. 
<emphasis id="B975BA4BFF88FFE4C889FD2A459FFD6A" bold="true" box="[476,491,696,721]" pageId="16" pageNumber="17">n</emphasis>
.
</taxonomicNameLabel>
, and 
<taxonomicName id="4C011DDAFF88FFE4CB7AFD2B46E1FD6B" authorityName="Pfitzner" authorityYear="1906" box="[559,661,697,720]" class="Insecta" family="Hepialidae" genus="Triodia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="B975BA4BFF88FFE4CB7AFD2B46E1FD6B" box="[559,661,697,720]" italics="true" pageId="16" pageNumber="17">P. remota</emphasis>
</taxonomicName>
, from the indistinct markings on a dark grayish brown background in 
<taxonomicName id="4C011DDAFF88FFE4C9C2FD4F456AFD4E" authority="Grehan &amp; Mielke, 2018" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[151,286,732,757]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="titarenkoi" status="sp. nov.">
<emphasis id="B975BA4BFF88FFE4C9C2FD4F456AFD4E" box="[151,286,732,757]" italics="true" pageId="16" pageNumber="17">P. titarenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF88FFE4C870FD4E4517FD4F" box="[293,355,732,757]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B975BA4BFF88FFE4C870FD4E454BFD4E" bold="true" box="[293,319,732,757]" pageId="16" pageNumber="17">sp</emphasis>
. 
<emphasis id="B975BA4BFF88FFE4C818FD4E4528FD4E" bold="true" box="[333,348,732,757]" pageId="16" pageNumber="17">n</emphasis>
.
</taxonomicNameLabel>
, and the light grayish brown with pale to indistinct grayish white markings in 
<taxonomicName id="4C011DDAFF88FFE4CD87FD4E4110FD4E" authority="Grehan &amp; Mielke, 2018" authorityName="Grehan &amp; Mielke" authorityYear="2018" box="[1234,1380,732,757]" class="Insecta" family="Hepialidae" genus="Dugdaleiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B975BA4BFF88FFE4CD87FD4E4110FD4E" box="[1234,1380,732,757]" italics="true" pageId="16" pageNumber="17">Dugdaleiella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2460730FF88FFE4CC3EFD4E44DAFCA2" pageId="16" pageNumber="17" rank="genus">
<emphasis id="B975BA4BFF88FFE4CC3EFD4E44DAFCA2" bold="true" pageId="16" pageNumber="17">gen. n.</emphasis>
</taxonomicNameLabel>
Apices of the pseudotegumen are widely separated in contrast to proximate or adjacent apices of other 
<taxonomicName id="4C011DDAFF88FFE4C9C2FCB64556FC86" authorityName="Viette" authorityYear="1951" box="[151,290,804,829]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B975BA4BFF88FFE4C9C2FCB64556FC86" box="[151,290,804,829]" italics="true" pageId="16" pageNumber="17">Pfitzneriella</emphasis>
</taxonomicName>
species. The female is externally distinguished from males of 
<taxonomicName id="4C011DDAFF88FFE4CABFFCB64001FC86" authorityName="Viette" authorityYear="1951" box="[1002,1141,804,829]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B975BA4BFF88FFE4CABFFCB64001FC86" box="[1002,1141,804,829]" italics="true" pageId="16" pageNumber="17">Pfitzneriella</emphasis>
</taxonomicName>
by the broad pale greyish transverse FW band. The external female genitalia is distinguished from all other 
<taxonomicName id="4C011DDAFF88FFE4CD7CFCDA40D6FCDA" authorityName="Stephens" authorityYear="1829" box="[1065,1186,840,865]" class="Insecta" family="Hepialidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
we have examined by the narrow, strongly setose, bar-shaped dorsal plate.
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF88FFE5C992FC02448EFA52" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BBE6659FF88FFE4C992FC02467FFC76" blockId="16.[151,1437,582,2022]" pageId="16" pageNumber="17">
<emphasis id="B975BA4BFF88FFE4C992FC024525FC12" bold="true" box="[199,337,912,937]" pageId="16" pageNumber="17">Description</emphasis>
. Male (
<figureCitation id="133A7ADCFF88FFE4C8F2FC024653FC12" box="[423,551,912,937]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="16" pageNumber="17">Figs 7a, 7b</figureCitation>
): Wingspan 
<quantity id="4CF9CBBCFF88FFE4CBE7FC024777FC13" box="[690,771,912,937]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="mm" value="50.0">50 mm</quantity>
; FW length: 
<quantity id="4CF9CBBCFF88FFE4CAC6FC024790FC13" box="[915,996,912,937]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="17" unit="mm" value="25.0">25 mm</quantity>
, width: 
<quantity id="4CF9CBBCFF88FFE4CD6EFC0240FEFC13" box="[1083,1162,912,936]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="17" unit="mm" value="11.0">11 mm</quantity>
, ratio 2.3: 1; HW length: 
<quantity id="4CF9CBBCFF88FFE4C9C2FC27449CFC77" box="[151,232,949,973]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="17" unit="mm" value="20.0">20 mm</quantity>
, width: 
<quantity id="4CF9CBBCFF88FFE4C817FC2745E7FC77" box="[322,403,949,973]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="mm" value="10.0">10 mm</quantity>
, ratio 2: 1.
</paragraph>
<paragraph id="8BBE6659FF88FFE4C992FC4A47E6FB1E" blockId="16.[151,1437,582,2022]" pageId="16" pageNumber="17">
<emphasis id="B975BA4BFF88FFE4C992FC4A4570FC4A" box="[199,260,984,1009]" italics="true" pageId="16" pageNumber="17">Head</emphasis>
. Frons and vertex covered with yellowish brown piliform scales tipped with dark brown, two tubercles, each dorsal and slightly medial to each antenna. Interocular-antennal scales piliform, overlapping eye. Antenna filiform, blackish-brown, flagellum laterally compressed, with numerous sensillae chaetica, terminal segments missing [antennae broken]; scape rounded, covered with lamellar scales. Clypeus unscaled, projecting anteriorly as a transverse ridge. Labium subsquare. Labial palpus two segmented, proximal palpomere short, oval, distal palpomere long and narrow, about five times long as basal (
<figureCitation id="133A7ADCFF88FFE4CA66FB1E47F1FB1E" box="[819,901,1164,1189]" captionStart="FIGURES 8–9" captionStartId="5.[151,264,1506,1529]" captionTargetBox="[151,1435,193,1486]" captionTargetId="figure@5.[151,1435,193,1486]" captionTargetPageId="5" captionText="FIGURES 8–9. Voucher specimens. 8, Dugdaleiella monticola male. 8a, LT (ZMHB); 8b, Cotopaxi, Ecuador (CMNH); 8c, Volcán Chiles, Ecuador (CMNH); 8d, male, Volcán Chiles, Ecuador (CMNH). 9, Labial palpi (male unless otherwise specified), 9a, Kozloviella viazmenskyi sp. n.; 9b, Pfitzneriella antonkozlovi sp. n.; 9c, P. rawlinsi sp. n.; 9d, P. titarenkoi sp. n.; 9e, P. yuliyakovalevae sp. n.; 9f, P. yuliyakovalevae sp. n. (female); 9g, Dugdaleiella monticola. Photos: Jane Hyland (Fig. 8), John Grehan (Figs 9a–b, d–h), and Carlos Mielke (Fig. 9c). Scale 1 mm (when present)." httpUri="https://zenodo.org/record/1451060/files/figure.png" pageId="16" pageNumber="17">Fig. 9e</figureCitation>
).
</paragraph>
<paragraph id="8BBE6659FF88FFE4C992FB22414DF91A" blockId="16.[151,1437,582,2022]" pageId="16" pageNumber="17">
<emphasis id="B975BA4BFF88FFE4C992FB22456CFB72" box="[199,280,1200,1225]" italics="true" pageId="16" pageNumber="17">Thorax</emphasis>
. Dorsally covered with yellowish brown scales, scutum III pale greyish brown, free of scales other than posterior and medial regions, obscured from direct view by overlapping mesothoracic scales. Ventral thorax yellowish to greyish brown. Legs (
<figureCitation id="133A7ADCFF88FFE4CB74FB6A460BFAAB" box="[545,639,1272,1297]" captionStart="FIGURES 12–14. 12" captionStartId="9.[151,264,1172,1195]" captionTargetBox="[150,1436,193,1151]" captionTargetId="figure@9.[150,1436,193,1151]" captionTargetPageId="9" captionText="FIGURES 12–14. 12, Antenna. Pfitzneriella rawlinsi sp. n. (unscaled). 13, Legs. 13a, Kozloviella viazmenskyi sp. n.; 13b, P. olafi sp. n.; 13c, P. rawlinsi sp. n.; 13d, P. titarenkoi sp. n.; 13e, P. yuliyakovalevae sp. n. 14, Metatibia. 14a, P. antonkozlovi sp. n.; 14b, P. rawlinsi sp. n.; 14c, P. titarenkoi; 14d, P. yuliyakovalevae sp. n.; 14e, P. antonkozlovi sp. n.; 14f, Dugdaleiella monticola. Photos: John Grehan (Figs 12, 14e, 14f), Carlos Mielke (Figs 13,14a–d). Scale 1 mm." httpUri="https://zenodo.org/record/1451066/files/figure.png" pageId="16" pageNumber="17">Fig. 13e</figureCitation>
) coloured as the thorax; length ratio (pro-, meso-, meta- legs) 1: 1.12: 0.78; hind leg pale orange-brown, basal tibia with slight swelling of scent gland with androconia (
<figureCitation id="133A7ADCFF88FFE4CC7DFA8E41FAFA8E" box="[1320,1422,1308,1333]" captionStart="FIGURES 12–14. 12" captionStartId="9.[151,264,1172,1195]" captionTargetBox="[150,1436,193,1151]" captionTargetId="figure@9.[150,1436,193,1151]" captionTargetPageId="9" captionText="FIGURES 12–14. 12, Antenna. Pfitzneriella rawlinsi sp. n. (unscaled). 13, Legs. 13a, Kozloviella viazmenskyi sp. n.; 13b, P. olafi sp. n.; 13c, P. rawlinsi sp. n.; 13d, P. titarenkoi sp. n.; 13e, P. yuliyakovalevae sp. n. 14, Metatibia. 14a, P. antonkozlovi sp. n.; 14b, P. rawlinsi sp. n.; 14c, P. titarenkoi; 14d, P. yuliyakovalevae sp. n.; 14e, P. antonkozlovi sp. n.; 14f, Dugdaleiella monticola. Photos: John Grehan (Figs 12, 14e, 14f), Carlos Mielke (Figs 13,14a–d). Scale 1 mm." httpUri="https://zenodo.org/record/1451066/files/figure.png" pageId="16" pageNumber="17">Fig. 14d</figureCitation>
); epiphysis present. FW costal margin slightly convex, outer margin weakly convex with extended shallow curve to anal margin. Venation hepialine (
<bibRefCitation id="EF901BA8FF88FFE4CB48FAF64691FAC6" box="[541,741,1380,1405]" pageId="16" pageNumber="17" refId="ref14701" refString="Dumbleton, L. J. (1966) Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera). New Zealand Journal of Science, 9, 920 - 981." type="journal article">Dumbleton 1966</bibRefCitation>
), base of FW anal vein strongly curved anteriorly, distally curved posteriorly (
<figureCitation id="133A7ADCFF88FFE4C823FA1A45AEFA1B" box="[374,474,1416,1441]" captionStart="FIGURE 10" captionStartId="6.[151,250,1649,1671]" captionTargetBox="[152,1437,195,1610]" captionTargetId="figure@6.[152,1437,195,1610]" captionTargetPageId="6" captionText="FIGURE 10. Wing venation (male unless otherwise specified). 10a, Kozloviella viazmenskyi sp. n.; 10b, P. antonkozlovi sp. n.; 10c, P. olafi sp. n.; 10d, P. rawlinsi sp. n.; 10e, P. remota (from photo only, venation incomplete due to damaged condition of HT); 10f, P. yuliyakovalevae sp. n.; 10g, P. yuliyakovalevae sp. n. (female); 10h, Dugdaleiella monticola." httpUri="https://zenodo.org/record/1451062/files/figure.png" pageId="16" pageNumber="17">
Fig. 
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</figureCitation>
). FW dorsal ground colour a mix of diffuse dark brown and black interspersed with pale yellowish brown; cell between 1A and CuA2 with large black patch enclosing yellowish white spot at edge of CuA2, and with curved posterior edge, posterior edge bordered by yellowish brown margin that extends anteriorly across FW as a broken post discal line, base of cell with prominent oval white spot ended with dark brown (
<figureCitation id="133A7ADCFF88FFE4CC3AFA6B44CBF98D" captionStart="FIGURE 11" captionStartId="7.[151,250,1429,1451]" captionTargetBox="[150,1436,810,1407]" captionTargetId="figure@7.[150,1436,810,1407]" captionTargetPageId="7" captionText="FIGURE 11. Wing markings. 11a; Kozloviella viazmenskyi sp. n., 11b; Pfitzneriella antonkozlovi sp. n.; 11c, P. olafi sp. n.; 11d, P. rawlinsi sp. n.; 11e, P. remota; 11f, P. yuliyakovalevae sp. n. Blue line—edge of white banding." httpUri="https://zenodo.org/record/1451064/files/figure.png" pageId="16" pageNumber="17">Fig. 11f</figureCitation>
); outer margin with shallow yellowish brown semicircles centered on each vein; fringe yellowish brown with black patches opposite terminus of each vein. FW ventral ground colour greyish brown, fringe as for dorsal; costal pocket present. HW dorsal greyish brown with pale orange-brown shading near apex and along outer fringe. HW ventral greyish brown except for orange-brown scales between Sc and costal margin, and fringe as in FW.
</paragraph>
<paragraph id="8BBE6659FF88FFE4C992F93F41F3F85D" blockId="16.[151,1437,582,2022]" pageId="16" pageNumber="17">
<emphasis id="B975BA4BFF88FFE4C992F93F4545F97D" box="[199,305,1709,1734]" italics="true" pageId="16" pageNumber="17">Abdomen</emphasis>
. Greyish brown. Tergites and sternites moderately sclerotized. Tergosternal connection with elongate tergosternal bar, elongate lateral tergal brace and short curved dorsal brace not connected to anterior ridge of tergum II; tergal knob absent (
<figureCitation id="133A7ADCFF88FFE4C8A6F9674621F8B5" box="[499,597,1781,1806]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[197,1386,199,1851]" captionTargetId="figure@10.[197,1386,199,1851]" captionTargetPageId="10" captionText="FIGURES 15–16. Tergosternal connection. 15a, Kozloviella viazmenskyi sp. n.; 15b, Pfitzneriella yuliyakovalevae sp. n.; 15c, P. rawlinsi sp. n.; 15d, Dugdaleiella monticola. 16, Abdomen (males unless otherwise specified). 16a, K. viazmenskyi sp. n.; 16b, P. antonkozlovi sp. n.; 16c, P. olafi sp. n.; 16d, P. rawlinsi sp. n.; 16e, P. titarenkoi sp. n.; 16f, P. yuliyakovalevae sp. n.; 16g, P. yuliyakovalevae sp. n. (female); 16h–16j, Dugdaleiella monticola. 16h, tergum II; 16i, sternum II; 16j, tergum and sternum VIII. Photos: John Grehan (Figs 15d, 16h–16j), Carlos Mielke (Figs 15a–15c, 16a–16g)." httpUri="https://zenodo.org/record/1451068/files/figure.png" pageId="16" pageNumber="17">Fig. 15c</figureCitation>
). Tergum II with anterior ridge across median, a thin lateral ridge, with a rounded curve at the outer edge of the lateral tuberculate plate, not extending to anterior margin; sternum II subrectangular, anterior margin broad, curved V-shape to lateral margin, anterior lateral arms short, dorsal edge of tergosternal attachment with short longitudinal ridge extending posteriorly. Tergum VII rectangular, wider than long, but slightly longer than preceding tergites. Tergum VIII rectangular, narrowing slightly to posterior margin; sternum VIII crescent shaped, anterior margin straight, lateral margins almost straight and parallel, posterior margin convex either side of median, medially rounded either side minute and narrow V-shaped notch (
<figureCitation id="133A7ADCFF88FFE4CC49F85F410EF85D" box="[1308,1402,1997,2022]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[197,1386,199,1851]" captionTargetId="figure@10.[197,1386,199,1851]" captionTargetPageId="10" captionText="FIGURES 15–16. Tergosternal connection. 15a, Kozloviella viazmenskyi sp. n.; 15b, Pfitzneriella yuliyakovalevae sp. n.; 15c, P. rawlinsi sp. n.; 15d, Dugdaleiella monticola. 16, Abdomen (males unless otherwise specified). 16a, K. viazmenskyi sp. n.; 16b, P. antonkozlovi sp. n.; 16c, P. olafi sp. n.; 16d, P. rawlinsi sp. n.; 16e, P. titarenkoi sp. n.; 16f, P. yuliyakovalevae sp. n.; 16g, P. yuliyakovalevae sp. n. (female); 16h–16j, Dugdaleiella monticola. 16h, tergum II; 16i, sternum II; 16j, tergum and sternum VIII. Photos: John Grehan (Figs 15d, 16h–16j), Carlos Mielke (Figs 15a–15c, 16a–16g)." httpUri="https://zenodo.org/record/1451068/files/figure.png" pageId="16" pageNumber="17">Fig. 16f</figureCitation>
).
</paragraph>
<paragraph id="8BBE6659FF89FFE5C992FF054765FE17" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
<emphasis id="B975BA4BFF89FFE5C992FF05451FFF0B" box="[199,363,151,176]" italics="true" pageId="17" pageNumber="18">Male genitalia</emphasis>
(
<figureCitation id="133A7ADCFF89FFE5C82CFF054663FF0B" box="[377,535,151,176]" captionStart="FIGURES 17–25" captionStartId="11.[151,264,1644,1667]" captionTargetBox="[151,1435,193,1629]" captionTargetId="figure@11.[151,1435,193,1629]" captionTargetPageId="11" captionText="FIGURES 17–25. Genitalia (male unless otherwise specified). 17, Kozloviella viazmenskyi sp. n. HT. 17a, ventral; 17b, lateral. 18, Pfitzneriella antonkozlovi sp. n. HT, ventral. 19, P. olafi sp. n. HT, ventral. 20, P. rawlinsi sp. n. HT, ventral. 21, P. remota HT (redrawn from Viette (1951a)). 22, P. titarenkoi sp. n. HT, ventral. 23, P. yuliyakovalevae sp. n. 23a, ventral; 23b, lateral. 24, P. yuliyakovalevae sp. n. PT female. 24a, posterior; 24b, lateral. 25, Dugdaleiella monticola. 25a, LT (ZMHB); 25b, Cotopaxi, Parque Nacional Cotopaxi. Dissection JRG 136 (CMNH). sa = saccus, fs = fultura superior, fi = fultura inferior, as = apodemal suture. Scale 1 mm (when present). Photos: Carlos Mielke (Figs 17–20, 22–25b)." httpUri="https://zenodo.org/record/1451070/files/figure.png" pageId="17" pageNumber="18">Figs 23a, 23b</figureCitation>
). Tegumen not distinct from pseudotegumen. Saccus very short, curving parallel to fultura inferior, apodemal suture with short posterior-ventral shelf laterally, each with a short, central and posteriorly oriented spur. Tergal lobes absent. Pseudotegumen as a pair of triangular plates, wide dorsally, narrowing ventrally to a blunt apex, slightly angled laterally and with small dorsally oriented spine on the posterior edge, convex adjacent to phallus, not fused medially, ventral apex widely separated medially. Valva broad, spatulate with rounded distal margin, inner and distal surfaces setose. Fultura superior membranous except for two small antero-lateral patches, longitudinally very short; fultura inferior forming a wide, narrow, and strongly sclerotized band. Phallus membranous without cornutus.
</paragraph>
<paragraph id="8BBE6659FF89FFE5C992FE2545B7FE4F" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
Female (
<figureCitation id="133A7ADCFF89FFE5C87FFE2545C5FE6B" box="[298,433,439,464]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="17" pageNumber="18">Figs 7c, 7d</figureCitation>
): Wingspan 
<quantity id="4CF9CBBCFF89FFE5CB14FE2546E1FE74" box="[577,661,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="17" pageNumber="18" unit="mm" value="46.0">46 mm</quantity>
; FW length: 
<quantity id="4CF9CBBCFF89FFE5CA78FE2A47F5FE74" box="[813,897,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="18" unit="mm" value="22.0">22 mm</quantity>
, width: 
<quantity id="4CF9CBBCFF89FFE5CA8BFE2A4044FE74" box="[990,1072,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="18" unit="mm" value="11.0">11 mm</quantity>
, ratio 2: 1; HW length: 
<quantity id="4CF9CBBCFF89FFE5CC11FE2A41ECFE74" box="[1348,1432,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="mm" value="20.0">20 mm</quantity>
, width: 
<quantity id="4CF9CBBCFF89FFE5C9B0FE4E4542FE4F" box="[229,310,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="18" unit="mm" value="14.0">14 mm</quantity>
, ratio 1.4: 1.
</paragraph>
<paragraph id="8BBE6659FF89FFE5C992FE6D403BFD13" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
<emphasis id="B975BA4BFF89FFE5C992FE6D4570FDA3" box="[199,260,511,536]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
. Frons and vertex covered with yellowish brown piliform scales. Intraocular-antennal scales piliform, extending over the width of the eye. Antenna filiform, broken off other than at base; scape rounded. Clypeus unscaled, projecting anteriorly as a transverse ridge with pits lateral to median. Labium covered with dark grayish brown scales. Labium slightly pentagonal, longer than wide. Labial palpus two segmented, proximal palpomere short, oval, distal palpomere long and narrow, about 4.5 times long as basal (
<figureCitation id="133A7ADCFF89FFE5CAA6FD1D4036FD13" box="[1011,1090,655,680]" captionStart="FIGURES 8–9" captionStartId="5.[151,264,1506,1529]" captionTargetBox="[151,1435,193,1486]" captionTargetId="figure@5.[151,1435,193,1486]" captionTargetPageId="5" captionText="FIGURES 8–9. Voucher specimens. 8, Dugdaleiella monticola male. 8a, LT (ZMHB); 8b, Cotopaxi, Ecuador (CMNH); 8c, Volcán Chiles, Ecuador (CMNH); 8d, male, Volcán Chiles, Ecuador (CMNH). 9, Labial palpi (male unless otherwise specified), 9a, Kozloviella viazmenskyi sp. n.; 9b, Pfitzneriella antonkozlovi sp. n.; 9c, P. rawlinsi sp. n.; 9d, P. titarenkoi sp. n.; 9e, P. yuliyakovalevae sp. n.; 9f, P. yuliyakovalevae sp. n. (female); 9g, Dugdaleiella monticola. Photos: Jane Hyland (Fig. 8), John Grehan (Figs 9a–b, d–h), and Carlos Mielke (Fig. 9c). Scale 1 mm (when present)." httpUri="https://zenodo.org/record/1451060/files/figure.png" pageId="17" pageNumber="18">Fig. 9f</figureCitation>
).
</paragraph>
<paragraph id="8BBE6659FF89FFE5C992FD26453FFA8D" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
<emphasis id="B975BA4BFF89FFE5C992FD26456CFD76" box="[199,280,692,717]" italics="true" pageId="17" pageNumber="18">Thorax</emphasis>
. Dorsally covered with long yellowish brown piliform scales, scutum III pale yellowish brown, free of scales other than posterior and medial regions, obscured from direct view by overlapping mesothoracic scales. Ventral thorax pale greyish brown. Legs coloured as the thorax tibial and tarsal scales not obscuring individual tarsal segments; metaleg pale orange-brown; leg length ratio (pro-, meso-, meta-) 1: 1.1: 0.85; epiphysis absent. FW costal margin slightly convex; outer margin convex, merging with anal margin. Venation hepialine (
<bibRefCitation id="EF901BA8FF89FFE5CC49FCD644A7FC3B" pageId="17" pageNumber="18" refId="ref14701" refString="Dumbleton, L. J. (1966) Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera). New Zealand Journal of Science, 9, 920 - 981." type="journal article">Dumbleton 1966</bibRefCitation>
), base of FW anal vein strongly curved anteriorly, distally curved posteriorly (
<figureCitation id="133A7ADCFF89FFE5CD01FCF540C3FC3B" box="[1108,1207,871,896]" captionStart="FIGURE 10" captionStartId="6.[151,250,1649,1671]" captionTargetBox="[152,1437,195,1610]" captionTargetId="figure@6.[152,1437,195,1610]" captionTargetPageId="6" captionText="FIGURE 10. Wing venation (male unless otherwise specified). 10a, Kozloviella viazmenskyi sp. n.; 10b, P. antonkozlovi sp. n.; 10c, P. olafi sp. n.; 10d, P. rawlinsi sp. n.; 10e, P. remota (from photo only, venation incomplete due to damaged condition of HT); 10f, P. yuliyakovalevae sp. n.; 10g, P. yuliyakovalevae sp. n. (female); 10h, Dugdaleiella monticola." httpUri="https://zenodo.org/record/1451062/files/figure.png" pageId="17" pageNumber="18">Fig. 10h</figureCitation>
). FW dorsal ground colour pale greyish yellow with diffuse greyish-brown markings in posterior discal cell; broad postdiscal greyish white band extending between cell of CuA1 and CuA2 and costal margin near apex, curving in parallel with outer margin; marginal region comprising three broken bands of dark brown edged white markings (inner and outer rows) and dark brown (middle row); CuA2 lined with band of white along posterior edge; dark brown spot at base of anal margin, outer and anal margin lined with fringe basally pale yellowish brown and distally as greyish brown, marginal scales longest along basal region of anal margin. FW ventral ground colour greyish brown, diffuse yellowish brown over submarginal region of outer margin and basally; costal pocket with very fine piliform scales; fringe predominantly pale yellowish brown with darker brown tips. HW dorsal greyish brown with pale yellowish brown fringe; costa-subcosta regions covered with fine piliform scales. HW ventral greyish brown with pale yellowish brown fringe. 
<emphasis id="B975BA4BFF89FFE5C8EDFB474656FB55" box="[440,546,1237,1262]" italics="true" pageId="17" pageNumber="18">Abdomen</emphasis>
. Pale yellowish to greyish brown, tergites and sternites weakly sclerotized. Tergum VIII rectangular, wider than long, but longer than preceding tergites; sternite VIII rectangular, longer than wide (
<figureCitation id="133A7ADCFF89FFE5C988FA8F454AFA8D" box="[221,318,1309,1334]" captionStart="FIGURES 15–16" captionStartId="10.[151,264,1890,1913]" captionTargetBox="[197,1386,199,1851]" captionTargetId="figure@10.[197,1386,199,1851]" captionTargetPageId="10" captionText="FIGURES 15–16. Tergosternal connection. 15a, Kozloviella viazmenskyi sp. n.; 15b, Pfitzneriella yuliyakovalevae sp. n.; 15c, P. rawlinsi sp. n.; 15d, Dugdaleiella monticola. 16, Abdomen (males unless otherwise specified). 16a, K. viazmenskyi sp. n.; 16b, P. antonkozlovi sp. n.; 16c, P. olafi sp. n.; 16d, P. rawlinsi sp. n.; 16e, P. titarenkoi sp. n.; 16f, P. yuliyakovalevae sp. n.; 16g, P. yuliyakovalevae sp. n. (female); 16h–16j, Dugdaleiella monticola. 16h, tergum II; 16i, sternum II; 16j, tergum and sternum VIII. Photos: John Grehan (Figs 15d, 16h–16j), Carlos Mielke (Figs 15a–15c, 16a–16g)." httpUri="https://zenodo.org/record/1451068/files/figure.png" pageId="17" pageNumber="18">
Fig. 
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</figureCitation>
).
</paragraph>
<paragraph id="8BBE6659FF89FFE5C992FAD3448EFA52" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
<emphasis id="B975BA4BFF89FFE5C992FAD345FFFAE1" box="[199,395,1345,1370]" italics="true" pageId="17" pageNumber="18">Female genitalia</emphasis>
(
<figureCitation id="133A7ADCFF89FFE5C8CAFAD34632FAE1" box="[415,582,1345,1370]" captionStart="FIGURES 17–25" captionStartId="11.[151,264,1644,1667]" captionTargetBox="[151,1435,193,1629]" captionTargetId="figure@11.[151,1435,193,1629]" captionTargetPageId="11" captionText="FIGURES 17–25. Genitalia (male unless otherwise specified). 17, Kozloviella viazmenskyi sp. n. HT. 17a, ventral; 17b, lateral. 18, Pfitzneriella antonkozlovi sp. n. HT, ventral. 19, P. olafi sp. n. HT, ventral. 20, P. rawlinsi sp. n. HT, ventral. 21, P. remota HT (redrawn from Viette (1951a)). 22, P. titarenkoi sp. n. HT, ventral. 23, P. yuliyakovalevae sp. n. 23a, ventral; 23b, lateral. 24, P. yuliyakovalevae sp. n. PT female. 24a, posterior; 24b, lateral. 25, Dugdaleiella monticola. 25a, LT (ZMHB); 25b, Cotopaxi, Parque Nacional Cotopaxi. Dissection JRG 136 (CMNH). sa = saccus, fs = fultura superior, fi = fultura inferior, as = apodemal suture. Scale 1 mm (when present). Photos: Carlos Mielke (Figs 17–20, 22–25b)." httpUri="https://zenodo.org/record/1451070/files/figure.png" pageId="17" pageNumber="18">Figs 24a, 24b</figureCitation>
). Dorsal plate a narrow band, lightly sclerotized, dorsal edge narrowing slightly at apex, covered in numerous, long setae; lamella antevaginalis as a broad sclerotized plate, central dorsal margin with a broad shallow, medial V-shaped depression bordered each side by rounded concave margin; surface of central region with numerous short setae, remaining surface with fewer scattered setae, ventral margin broadly concave.
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF89FFE5C992FA674001F9ED" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BBE6659FF89FFE5C992FA674001F9ED" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
<emphasis id="B975BA4BFF89FFE5C992FA67452DF9B5" bold="true" box="[199,345,1525,1550]" pageId="17" pageNumber="18">Distribution</emphasis>
. South-central eastern Peruvian Andes (
<figureCitation id="133A7ADCFF89FFE5CA7BFA6747CBF9B5" box="[814,959,1525,1550]" captionStart="FIGURES 27–29" captionStartId="20.[151,264,1710,1733]" captionTargetBox="[156,1435,363,1685]" captionTargetId="figure@20.[151,1436,363,1690]" captionTargetPageId="20" captionText="FIGURES 27–29. Distribution. 27, Dugdaleiella gen. n., Kozloviella gen. n., and Pfitzneriella locality records. 27a, Ecuador and Peru. Tectonic boundaries approximate; 27b, Dugdaleiella gen. n. and P. lucicola in Ecuador. 28, Locality records of Peruvian Pfitzneriella in relation to eastern Andean montane forests (modified from Young &amp; León (2000) with permission by Kenneth Young). 29, Principal groups of Hepialidae in Central and South America. Phassus/Schausiana/Phthius (solid line), cibyrine genera of Hepialidae (dotted line), Dugdaleiella gen. n./Kozloviella gen. n./Pfitzneriella (pale green), Uiridigigas (red), and southern Andean genera (blue)." httpUri="https://zenodo.org/record/1451074/files/figure.png" pageId="17" pageNumber="18">Fig. 27a, 28</figureCitation>
). Male known only from 
<typeStatus id="54BAD8FBFF89FFE5CDB8FA644169F9B5" box="[1261,1309,1526,1550]" pageId="17" pageNumber="18">type</typeStatus>
locality in upper river valley west of Satipo, collected next to disturbed forest near river at 
<quantity id="4CF9CBBCFF89FFE5CD61F98B40E6F98A" box="[1076,1170,1561,1586]" metricMagnitude="3" metricUnit="m" metricValue="2.69" pageId="17" pageNumber="18" unit="m" value="2690.0">2,690 m</quantity>
(
<figureCitation id="133A7ADCFF89FFE5CDF6F98B4134F989" box="[1187,1344,1561,1586]" captionStart="FIGURE 26" captionStartId="14.[151,250,1674,1696]" captionTargetBox="[152,1435,266,1652]" captionTargetId="figure@14.[152,1435,195,1652]" captionTargetPageId="14" captionText="FIGURE 26. Collection localities. 26a–26b, Kozloviella viazmenskyi sp. n. Peru, Cerro Machete, 2,050 m; 26c, Pfitzneriella antonkozlovi sp. n., Peru, Marañon Province, 3,270 m; 26d, P. titarenkoi, sp. n. and P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 3,380 m; 26e–26f, P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 2,960 m. With permission by Anton Kozlov and Yuliya Kovaleva, foreground 26b, 26e, Anton Kozlov and 26f, Yuliya Kovaleva. Photos: Anton Kozlov." httpUri="https://zenodo.org/record/1451072/files/figure.png" pageId="17" pageNumber="18">Figs 26e, 26f</figureCitation>
). Single female from same valley, next to disturbed forest adjacent to river at 
<quantity id="4CF9CBBCFF89FFE5CACEF9AC4783F9EE" box="[923,1015,1598,1622]" metricMagnitude="3" metricUnit="m" metricValue="3.38" pageId="17" pageNumber="18" unit="m" value="3380.0">3,380 m</quantity>
(
<figureCitation id="133A7ADCFF89FFE5CD53F9AF401CF9ED" box="[1030,1128,1597,1622]" captionStart="FIGURE 26" captionStartId="14.[151,250,1674,1696]" captionTargetBox="[152,1435,266,1652]" captionTargetId="figure@14.[152,1435,195,1652]" captionTargetPageId="14" captionText="FIGURE 26. Collection localities. 26a–26b, Kozloviella viazmenskyi sp. n. Peru, Cerro Machete, 2,050 m; 26c, Pfitzneriella antonkozlovi sp. n., Peru, Marañon Province, 3,270 m; 26d, P. titarenkoi, sp. n. and P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 3,380 m; 26e–26f, P. yuliyakovalevae sp. n., Peru, Pampa Hermosa district, 2,960 m. With permission by Anton Kozlov and Yuliya Kovaleva, foreground 26b, 26e, Anton Kozlov and 26f, Yuliya Kovaleva. Photos: Anton Kozlov." httpUri="https://zenodo.org/record/1451072/files/figure.png" pageId="17" pageNumber="18">Fig. 26d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF89FFE5C992F9F3402FF925" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BBE6659FF89FFE5C992F9F3402FF925" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
<emphasis id="B975BA4BFF89FFE5C992F9F34532F9C1" bold="true" box="[199,326,1633,1658]" pageId="17" pageNumber="18">Etymology</emphasis>
. Named for Yuliya Alexandrovna Kovaleva (
<collectingRegion id="49C5A8BBFF89FFE5CA3FF9F047B9F9C1" box="[874,973,1634,1658]" country="Russia" name="Moscow" pageId="17" pageNumber="18">Moscow</collectingRegion>
, 
<collectingCountry id="F31626C9FF89FFE5CA8BF9F34059F9C1" box="[990,1069,1633,1658]" name="Russia" pageId="17" pageNumber="18">Russia</collectingCountry>
), who participated in the 
<collectingCountry id="F31626C9FF89FFE5CC3CF9F041E8F9C2" box="[1385,1436,1634,1657]" name="Peru" pageId="17" pageNumber="18">Peru</collectingCountry>
expedition and assisted with specimen collection, by special request of Anton Kozlov.
</paragraph>
</subSubSection>
<subSubSection id="C31B35D2FF89FFE5C992F93B40F3F8CD" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BBE6659FF89FFE5C992F93B460AF895" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
<materialsCitation id="3B696C04FF89FFE5C992F93B46E4F95D" ID-GBIF-Occurrence="1914299827" collectorName="S. America" country="Peru" location="Satipo Prov" pageId="17" pageNumber="18" specimenCount="2" stateProvince="Junin" typeStatus="holotype">
<emphasis id="B975BA4BFF89FFE5C992F93B4502F979" bold="true" box="[199,374,1705,1730]" pageId="17" pageNumber="18">
<typeStatus id="54BAD8FBFF89FFE5C992F93B4576F979" box="[199,258,1705,1730]" pageId="17" pageNumber="18">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54BAD8FBFF89FFE5C82BF93B4591F979" box="[382,485,1705,1730]" pageId="17" pageNumber="18" type="holotype">
<emphasis id="B975BA4BFF89FFE5C82BF93B4591F979" box="[382,485,1705,1730]" italics="true" pageId="17" pageNumber="18">Holotype</emphasis>
</typeStatus>
male (with the following labels separated by forward slashes): / 
<collectorName id="26F4038FFF89FFE5CD97F9384135F979" box="[1218,1345,1705,1730]" pageId="17" pageNumber="18">S. America</collectorName>
, 
<collectingCountry id="F31626C9FF89FFE5CC18F93841ECF97A" box="[1357,1432,1706,1729]" name="Peru" pageId="17" pageNumber="18">PERU</collectingCountry>
, 
<collectingRegion id="49C5A8BBFF89FFE5C9C2F95F44A0F95D" box="[151,212,1741,1766]" country="Peru" name="Junin" pageId="17" pageNumber="18">Junin</collectingRegion>
reg. 
<location id="8EDE3082FF89FFE5C859F95F45E5F95E" LSID="urn:lsid:plazi:treatment:03A8D74FFF88FFE5C9C2FE4E40F3F8CD:8EDE3082FF89FFE5C859F95F45E5F95E" box="[268,401,1741,1766]" country="Peru" name="Satipo Prov" pageId="17" pageNumber="18" stateProvince="Junin">Satipo Prov</location>
, Pampa Hermosa distr
</materialsCitation>
. 11.5091S, 074.8596W, h= 
<quantity id="4CF9CBBCFF89FFE5CAE8F95F4062F95E" box="[957,1046,1741,1766]" metricMagnitude="3" metricUnit="m" metricValue="2.69" pageId="17" pageNumber="18" unit="m" value="2690.0">2690 m</quantity>
, 
<date id="FFBF4099FF89FFE5CD74F95C40E5F95D" box="[1057,1169,1742,1766]" pageId="17" pageNumber="18" value="2017-01-20">20/I/2017</date>
, Leg. A. Kozlov, Yuliya Kovaleva, Roman Gortovannyi / Holotypus, 
<taxonomicName id="4C011DDAFF89FFE5CBF1F963416EF8B1" authority="♂, Grehan &amp; C. Mielke" authorityName="♂, Grehan &amp; C. Mielke" authorityYear="2018" box="[676,1306,1777,1802]" class="Insecta" family="Hepialidae" genus="Pfitzneriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="yuliyakovalevae">
<emphasis id="B975BA4BFF89FFE5CBF1F963479BF8B1" box="[676,1007,1777,1802]" italics="true" pageId="17" pageNumber="18">Pfitzneriella yuliyakovalevae</emphasis>
♂, Grehan &amp; C. Mielke
</taxonomicName>
det. 2017/ Dissection JRG M261(CMNH). 
<figureCitation id="133A7ADCFF89FFE5C8ABF887460EF895" box="[510,634,1813,1838]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="17" pageNumber="18">Figs 7a, 7b</figureCitation>
.
</paragraph>
<paragraph id="8BBE6659FF89FFE5C992F8A840F3F8CD" blockId="17.[151,1437,151,1910]" pageId="17" pageNumber="18">
<typeStatus id="54BAD8FBFF89FFE5C992F8A84558F8EA" box="[199,300,1850,1873]" pageId="17" pageNumber="18" type="paratype">
<emphasis id="B975BA4BFF89FFE5C992F8A84558F8EA" box="[199,300,1850,1873]" italics="true" pageId="17" pageNumber="18">Paratype</emphasis>
</typeStatus>
. 
<specimenCount id="9D07ADD0FF89FFE5C868F8A8451EF8EA" box="[317,362,1850,1874]" pageId="17" pageNumber="18" type="female">1 ♀</specimenCount>
(
<figureCitation id="133A7ADCFF89FFE5C829F8AB4675F8E9" box="[380,513,1849,1874]" captionStart="FIGURES 1–7" captionStartId="4.[151,264,1707,1730]" captionTargetBox="[151,1436,195,1660]" captionTargetId="figure@4.[151,1436,195,1660]" captionTargetPageId="4" captionText="FIGURES 1–7. Voucher specimens. 1, Kozloviella viazmenskyi gen. n., sp. n. male HT (CMNH). 1a, dorsal; 1b, ventral. 2, Pfitzneriella antonkozlovi sp. n. male HT (CMNH), 2a, dorsal; 2b, ventral. 3, P. olafi sp. n. male HT (MNHN), 3a, dorsal; 3b, ventral. 4, P. rawlinsi sp. n. male, 4a, HT dorsal (CMNH); 4b, PT dorsal; 4c, PT ventral (CGCM). 5, P. remota male HT dorsal (MNHN). 6, P. titarenkoi sp. n. male HT (CMNH), 6a, dorsal; 6b, ventral. 7, P. yuliyakovalevae sp. n. male HT (CMNH), 7a, dorsal; 7b, ventral. Female PT (CMNH); 7c, dorsal; 7d, ventral. Photos: Ernst Brockmann (Fig. 5), Jane Hyland (Fig. 4a), Anton Kozlov (Figs 1, 2, 6, 7), and Carlos Mielke (Figs 3, 4b, 4c)." httpUri="https://zenodo.org/record/1451058/files/figure.png" pageId="17" pageNumber="18">Figs 7c, 7d</figureCitation>
). 
<collectionCode id="ED10FE9CFF89FFE5CB4DF8A84617F8EA" box="[536,611,1850,1873]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14398" name="Universit<69> di Perugia" pageId="17" pageNumber="18">
<collectingCountry id="F31626C9FF89FFE5CB4DF8A84617F8EA" box="[536,611,1850,1873]" name="Peru" pageId="17" pageNumber="18">PERU</collectingCountry>
</collectionCode>
: 
<collectingRegion id="49C5A8BBFF89FFE5CB25F8A846C4F8E9" box="[624,688,1850,1874]" country="Peru" name="Junin" pageId="17" pageNumber="18">Junín</collectingRegion>
, Satipo, Pampa Hermosa distr., 
<geoCoordinate id="EE35009EFF89FFE5CD7DF8AB40E0F8E9" box="[1064,1172,1849,1874]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="5" value="-11.4865">11.4865S</geoCoordinate>
, 
<geoCoordinate id="EE35009EFF89FFE5CDF6F8A84153F8E9" box="[1187,1319,1850,1874]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="5" value="-74.8871">074.8871W</geoCoordinate>
, 
<quantity id="4CF9CBBCFF89FFE5CC63F8A841E3F8EA" box="[1334,1431,1850,1874]" metricMagnitude="3" metricUnit="m" metricValue="3.38" pageId="17" pageNumber="18" unit="m" value="3380.0">3,380 m</quantity>
, 
<date id="FFBF4099FF89FFE5C9C2F8CC4571F8CD" box="[151,261,1886,1910]" pageId="17" pageNumber="18" value="2017-01-21">21.I.2017</date>
, A. Kozlov, Yu. Kovaleva, R. Gortovannyi 
<emphasis id="B975BA4BFF89FFE5CBB9F8CF477AF8CD" box="[748,782,1885,1910]" italics="true" pageId="17" pageNumber="18">leg</emphasis>
. (dissection JRG F262) (
<collectionCode id="ED10FE9CFF89FFE5CD77F8CC400FF8CD" box="[1058,1147,1886,1910]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="17" pageNumber="18">CMNH</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>