139 lines
16 KiB
XML
139 lines
16 KiB
XML
<document id="F3721E556FAAC06577C382AC230DF411" ID-CLB-Dataset="46828" ID-DOI="10.5281/zenodo.277641" ID-GBIF-Dataset="e945637f-1e4a-433e-b949-8647c3fb20ee" ID-ISSN="1175-5326" ID-Zenodo-Dep="277641" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460359934862" checkinUser="plazi" docAuthor="Ballerio, Alberto, Gill, Bruce D. & Grebennikov, Vasily V." docDate="2011" docId="03A1C115FFD4FFA0FF3FFD63DDDBFA80" docLanguage="en" docName="zt02892p024.pdf" docOrigin="Zootaxa 2892" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ceratocanthinae" docType="treatment" docVersion="9" lastPageNumber="23" masterDocId="FF98B96DFFC2FFB6FFA8FFEDDC1FFFD2" masterDocTitle="Illustrated overview and identification key to Cameroonian Ceratocanthinae beetles (Coleoptera: Scarabaeoidea: Hybosoridae) with description of four new species" masterLastPageNumber="24" masterPageNumber="1" pageNumber="23" updateTime="1698294331078" updateUser="ExternalLinkService">
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<mods:title id="88E854CECB750037416BCA113B255365">Illustrated overview and identification key to Cameroonian Ceratocanthinae beetles (Coleoptera: Scarabaeoidea: Hybosoridae) with description of four new species</mods:title>
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<mods:namePart id="CF9F25264481643363B4C3AC928A81B4">Ballerio, Alberto</mods:namePart>
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<mods:namePart id="16EFC9D47FF686BE62FDB4CE465D3257">Gill, Bruce D.</mods:namePart>
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<mods:name id="7EBEC8375CD9971CBB90F003408EA21F" type="personal">
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<mods:namePart id="34FFFE7A2888FC68BB3F89BD60679D00">Grebennikov, Vasily V.</mods:namePart>
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<mods:title id="AD53655F550CA65235C14AD9936A8FBA">Zootaxa</mods:title>
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<mods:part id="68FE7B215D02AADA8E8EB09E91C2C91C">
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<mods:date id="8F0F107955F4F8F46F0B589074FE670F">2011</mods:date>
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<mods:number id="4643C6F84BEA8F3D4880C03CBFDD603F">2892</mods:number>
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<mods:identifier id="06291588A5F62C60A091EE1E8C854DBB" type="DOI">10.5281/zenodo.277641</mods:identifier>
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<treatment id="03A1C115FFD4FFA0FF3FFD63DDDBFA80" ID-DOI="http://doi.org/10.5281/zenodo.6183927" ID-GBIF-Taxon="119409485" ID-Zenodo-Dep="6183927" LSID="urn:lsid:plazi:treatment:03A1C115FFD4FFA0FF3FFD63DDDBFA80" httpUri="http://treatment.plazi.org/id/03A1C115FFD4FFA0FF3FFD63DDDBFA80" lastPageNumber="23" pageId="22" pageNumber="23">
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<subSubSection id="C3122388FFD4FFA0FF3FFD63DDDBFA80" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8BB77003FFD4FFA0FF3FFD63DF01FD75" blockId="22.[151,1437,654,1362]" box="[151,798,654,679]" pageId="22" pageNumber="23">
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<heading id="D0FFC76FFFD4FFA0FF3FFD63DF01FD75" bold="true" box="[151,798,654,679]" fontSize="10" level="2" pageId="22" pageNumber="23" reason="2">
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<emphasis id="B97CAC11FFD4FFA0FF3FFD63DF01FD75" bold="true" box="[151,798,654,679]" pageId="22" pageNumber="23">
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<taxonomicName id="4C080B80FFD4FFA0FF3FFD63DD41FD75" ID-CoL="6275R" box="[151,350,654,679]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subFamily" subFamily="Ceratocanthinae">Ceratocanthinae</taxonomicName>
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of the Guineo-Congolian forest block
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB77003FFD4FFA0FF6FFD5FD8D9FBA8" blockId="22.[151,1437,654,1362]" pageId="22" pageNumber="23">
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The Guineo-Congolian forest block has the following seven
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<taxonomicName id="4C080B80FFD4FFA0FC24FD5FD85DFD18" box="[908,1090,690,714]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subFamily" subFamily="Ceratocanthinae">Ceratocanthinae</taxonomicName>
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genera strictly endemic to it:
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<taxonomicName id="4C080B80FFD4FFA0FF3FFD3BDDF9FD3C" authority="Paulian, 1968" authorityName="Paulian" authorityYear="1968" box="[151,486,726,750]" class="Insecta" family="Hybosoridae" genus="Baloghianestes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FF3FFD3BDD5DFD3C" box="[151,322,726,750]" italics="true" pageId="22" pageNumber="23">Baloghianestes</emphasis>
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Paulian, 1968
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</taxonomicName>
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,
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<taxonomicName id="4C080B80FFD4FFA0FE5AFD3BDF6BFD3C" authority="Paulian, 1968" authorityName="Paulian" authorityYear="1968" box="[498,884,726,750]" class="Insecta" family="Hybosoridae" genus="Callophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FE5AFD3BDED0FD3C" box="[498,719,726,750]" italics="true" pageId="22" pageNumber="23">Callophilharmostes</emphasis>
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Paulian, 1968
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</taxonomicName>
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,
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<taxonomicName id="4C080B80FFD4FFA0FCD7FD3BD911FD3C" authority="Paulian, 1968" authorityName="Paulian" authorityYear="1968" box="[895,1294,726,750]" class="Insecta" family="Hybosoridae" genus="Carinophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FCD7FD3BD872FD3C" box="[895,1133,726,750]" italics="true" pageId="22" pageNumber="23">Carinophilharmostes</emphasis>
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Paulian, 1968
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</taxonomicName>
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<taxonomicName id="4C080B80FFD4FFA0FABEFD3BDDADFCC0" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" class="Insecta" family="Hybosoridae" genus="Chaetophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FABEFD3BDD15FCC1" italics="true" pageId="22" pageNumber="23">Chaetophilharmostes</emphasis>
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<bibRefCitation id="EF990DF2FFD4FFA0FEBBFD17DDADFCC0" author="Paulian" box="[275,434,762,786]" pageId="22" pageNumber="23" refString="Paulian, R. (1977) Revision des Ceratocanthidae (Coleoptera, Scarabaeidae). I. - Les formes africaines. Revue de Zoologie Africaine, 91, 253 - 316." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C080B80FFD4FFA0FE17FD16DEE2FCC0" authority="Paulian, 1968" authorityName="Paulian" authorityYear="1968" box="[447,765,762,786]" class="Insecta" family="Hybosoridae" genus="Congomostes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FE17FD16DE49FCC0" box="[447,598,763,786]" italics="true" pageId="22" pageNumber="23">Congomostes</emphasis>
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Paulian, 1968
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</taxonomicName>
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,
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<taxonomicName id="4C080B80FFD4FFA0FCA3FD11D850FCC0" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" box="[779,1103,762,787]" class="Insecta" family="Hybosoridae" genus="Petrovitzostes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FCA3FD11DFB8FCC1" box="[779,935,764,787]" italics="true" pageId="22" pageNumber="23">Petrovitzostes</emphasis>
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<bibRefCitation id="EF990DF2FFD4FFA0FC07FD17D850FCC0" author="Paulian" box="[943,1103,762,786]" pageId="22" pageNumber="23" refString="Paulian, R. (1977) Revision des Ceratocanthidae (Coleoptera, Scarabaeidae). I. - Les formes africaines. Revue de Zoologie Africaine, 91, 253 - 316." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
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</taxonomicName>
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, and
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<taxonomicName id="4C080B80FFD4FFA0FB25FD16DD2CFCE4" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" class="Insecta" family="Hybosoridae" genus="Pseudopterorthochaetes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FB25FD16D983FCC1" box="[1165,1436,763,787]" italics="true" pageId="22" pageNumber="23">Pseudopterorthochaetes</emphasis>
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<bibRefCitation id="EF990DF2FFD4FFA0FF3FFCF3DD2CFCE4" author="Paulian" box="[151,307,798,822]" pageId="22" pageNumber="23" refString="Paulian, R. (1977) Revision des Ceratocanthidae (Coleoptera, Scarabaeidae). I. - Les formes africaines. Revue de Zoologie Africaine, 91, 253 - 316." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
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</taxonomicName>
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(
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<bibRefCitation id="EF990DF2FFD4FFA0FEECFCF3DE72FCE4" author="Ocampo" box="[324,621,798,822]" pageId="22" pageNumber="23" refString="Ocampo, F. C. & Ballerio, A. (2006) Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae (Coleoptera: Scarabaeoidea). 4. Catalog of the subfamilies Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, and Pachyplectrinae (Hybosoridae). Bulletin of the University of Nebraska State Museum, 19, 178 - 209." type="journal article" year="2006">Ocampo & Ballerio 2006</bibRefCitation>
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; Ballerio 2008). Additionally, the genus
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<taxonomicName id="4C080B80FFD4FFA0FBEBFCF3DCD0FC88" authority="Paulian, 1968" authorityName="Paulian" authorityYear="1968" class="Insecta" family="Hybosoridae" genus="Melanophilharmostes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FBEBFCF3D928FCE4" box="[1091,1335,798,822]" italics="true" pageId="22" pageNumber="23">Melanophilharmostes</emphasis>
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Paulian, 1968
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</taxonomicName>
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is known from the Guineo-Congolian forest block and from the Kakamega forest,
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<collectingCountry id="F31F3093FFD4FFA0FB2FFCAED8CFFC89" box="[1159,1232,835,859]" name="Kenya" pageId="22" pageNumber="23">Kenya</collectingCountry>
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(Ballerio, unpublished data), which is considered as the easternmost fragment of the Guineo-Congolian block. The genus
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<taxonomicName id="4C080B80FFD4FFA0FA95FC8BDD68FC70" authority="Kolbe, 1895" authorityName="Kolbe" authorityYear="1895" class="Insecta" family="Hybosoridae" genus="Philharmostes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CAC11FFD4FFA0FA95FC8BDCFCFC71" italics="true" pageId="22" pageNumber="23">Philharmostes</emphasis>
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Kolbe, 1895
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</taxonomicName>
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is represented in the area by the endemic subgenus
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<emphasis id="B97CAC11FFD4FFA0FC64FC66D8BCFC71" box="[972,1187,907,931]" italics="true" pageId="22" pageNumber="23">Holophilharmostes</emphasis>
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Paulian, 1968 and by some distinctive species currently placed in the nominotypical subgenus (whose
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<typeStatus id="54B3CEA1FFD4FFA0FBB6FC42D851FC15" box="[1054,1102,943,967]" pageId="22" pageNumber="23">type</typeStatus>
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species is from
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<collectingCountry id="F31F3093FFD4FFA0FAACFC43D98FFC14" box="[1284,1424,942,966]" name="Madagascar" pageId="22" pageNumber="23">Madagascar</collectingCountry>
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), and are strongly differentiated from these representatives occurring elsewhere in Africa. The distribution pattern displayed by species in the area is interesting because of the occurrence of many species, including the flightless ones, across very extensive areas and encompassing both the forests of the Gulf of
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<collectingCountry id="F31F3093FFD4FFA0FB91FBF7D895FBE0" box="[1081,1162,1050,1074]" name="Guinea-Bissau" pageId="22" pageNumber="23">Guinea</collectingCountry>
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and of the
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<collectingCountry id="F31F3093FFD4FFA0FAA0FBF7D94DFBE0" box="[1288,1362,1050,1074]" name="Republic of the Congo" pageId="22" pageNumber="23">Congo</collectingCountry>
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basin. This suggests that the distribution patterns may be influenced by recent biogeographical events, with few localized endemics, making it difficult to find speciation centres and areas of endemism within the block.
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</paragraph>
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<paragraph id="8BB77003FFD4FFA0FF6FFB6ADDDBFA80" blockId="22.[151,1437,654,1362]" pageId="22" pageNumber="23">
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From a systematics point of view, the study of this fauna is complicated by numerous species known only by the
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<typeStatus id="54B3CEA1FFD4FFA0FF17FB46DCF0FB11" box="[191,239,1195,1219]" pageId="22" pageNumber="23">type</typeStatus>
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series or by very few additional specimens, as well as by the existence of a few species whose
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<typeStatus id="54B3CEA1FFD4FFA0FAB8FB46D954FB11" box="[1296,1355,1195,1219]" pageId="22" pageNumber="23">types</typeStatus>
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should be in the Museo do Dundo (
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<collectingCountry id="F31F3093FFD4FFA0FE7FFB23DE2FFB34" box="[471,560,1230,1254]" name="Angola" pageId="22" pageNumber="23">Angola</collectingCountry>
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). The identity of these species, all described by
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<bibRefCitation id="EF990DF2FFD4FFA0FBFBFB23D912FB34" author="Martinez" box="[1107,1293,1230,1254]" pageId="22" pageNumber="23" refString="Martinez, A. (1970) Contribucion al conocimiento de los Ceratocanthini africanos (Coleoptera: Scarabaeidae: Troginae). Publicacoes Culturais de Companhia de Diamantes de Angola, Lisboa, 82, 15 - 35." type="journal article" year="1970">Martínez (1970)</bibRefCitation>
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, will remain problematic until the
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<typeStatus id="54B3CEA1FFD4FFA0FE21FB1EDDA6FAD9" box="[393,441,1267,1291]" pageId="22" pageNumber="23">type</typeStatus>
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material (if still existing) becomes available for study.
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<bibRefCitation id="EF990DF2FFD4FFA0FB83FB1FD8CFFAD8" author="Paulian" box="[1067,1232,1266,1290]" pageId="22" pageNumber="23" refString="Paulian, R. (1977) Revision des Ceratocanthidae (Coleoptera, Scarabaeidae). I. - Les formes africaines. Revue de Zoologie Africaine, 91, 253 - 316." type="journal article" year="1977">Paulian (1977)</bibRefCitation>
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attempted to synonomize some of Martínez’s species, but his actions were tentative, being based on original descriptions only (see
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<bibRefCitation id="EF990DF2FFD4FFA0FF3FFAD7DDA7FA80" author="Ocampo" box="[151,440,1338,1362]" pageId="22" pageNumber="23" refString="Ocampo, F. C. & Ballerio, A. (2006) Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae (Coleoptera: Scarabaeoidea). 4. Catalog of the subfamilies Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, and Pachyplectrinae (Hybosoridae). Bulletin of the University of Nebraska State Museum, 19, 178 - 209." type="journal article" year="2006">Ocampo & Ballerio 2006</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |