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<mods:title id="7629F28F11D0E497821A456953931710">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<mods:namePart id="B8E4C78A3EF29C04B9C6CCA5763B220F">Schwarzhans, Werner</mods:namePart>
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<mods:namePart id="CD2EE66E60B3CBE8ACFE6785D62875AC">Klots, Oleksandr</mods:namePart>
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<mods:namePart id="B97F8E5BE68C883B52A30C773886EC51">Ryabokon, Tamara</mods:namePart>
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<mods:date id="DEF9AFCB56D9533C42A5EED1C81DA395">2022</mods:date>
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<treatment id="039D7D5BFE5DFFB6FBC8FB50FD7CF992" LSID="urn:lsid:plazi:treatment:039D7D5BFE5DFFB6FBC8FB50FD7CF992" httpUri="http://treatment.plazi.org/id/039D7D5BFE5DFFB6FBC8FB50FD7CF992" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection id="C32E9FC6FE5DFFB6FBC8FB50FED1FB52" box="[151,363,1163,1187]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8B8BCC4DFE5DFFB6FBC8FB50FED1FB52" blockId="20.[151,363,1163,1187]" box="[151,363,1163,1187]" pageId="20" pageNumber="21">
<heading id="D0C37B21FE5DFFB6FBC8FB50FED1FB52" bold="true" box="[151,363,1163,1187]" fontSize="10" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B940105FFE5DFFB6FBC8FB50FED1FB52" bold="true" box="[151,363,1163,1187]" pageId="20" pageNumber="21">
<taxonomicName id="4C34B7CEFE5DFFB6FBC8FB50FEADFB52" ID-CoL="G8R" ID-ENA="30871" authority="Swainson, 1839" authorityName="Swainson" authorityYear="1839" box="[151,279,1163,1187]" family="Serranidae" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="21" phylum="Chordata" rank="family">Serranidae</taxonomicName>
? indet.
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32E9FC6FE5DFFB6FBC8FB10FEAFFB12" box="[151,277,1227,1251]" pageId="20" pageNumber="21" type="description">
<paragraph id="8B8BCC4DFE5DFFB6FBC8FB10FEAFFB12" blockId="20.[151,277,1227,1251]" box="[151,277,1227,1251]" pageId="20" pageNumber="21">
<figureCitation id="130FD0C8FE5DFFB6FBC8FB10FEAFFB12" box="[151,277,1227,1251]" captionStart="Fig" captionStartId="21.[168,199,1587,1607]" captionTargetBox="[152,1438,237,1698]" captionTargetId="figure-47@21.[167,1422,253,1574]" captionTargetPageId="21" captionText="Fig. 9 Otoliths of Haemulidae, Serranidae, Sparidae, Leiognathidae and Caproidae:a (reversed)—Brachydeuterus speronatus (Bassoli,1906), Kozatskyi Yar, NMNHU-P PI 2549. b, c Serranidae indet., b (broken and repaired), c (reversed), Kozatskyi Yar,NMNHU-P PI 2579. d, e Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, Mlyntsi, NMNHU-P PI 2578. f (reversed)—Leiognathidae indet., Shydlivshchyna, NMB P1214. g Perciformes indet., Shydlivshchyna,NMB P1221. h (reversed) Capros aper (Linnaeus,1758), Mlyntsi, NMB P1206" figureDoi="http://doi.org/10.5281/zenodo.11999944" httpUri="https://zenodo.org/record/11999944/files/figure.png" pageId="20" pageNumber="21">Figure 9b, c</figureCitation>
</paragraph>
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<subSubSection id="C32E9FC6FE5DFFB6FBC8FAD0FF4AFA92" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8B8BCC4DFE5DFFB6FBC8FAD0FF4AFA92" blockId="20.[151,775,1291,1379]" pageId="20" pageNumber="21">
<emphasis id="B940105FFE5DFFB6FBC8FAD0FF42FAD2" box="[151,248,1291,1315]" italics="true" pageId="20" pageNumber="21">Material</emphasis>
<materialsCitation id="3B5CC610FE5DFFB6FA4FFAD0FEC0FAB2" collectionCode="NMNHU-P" pageId="20" pageNumber="21" specimenCode="PI 2579" specimenCount="2">
5 otoliths:
<specimenCount id="9D3207C4FE5DFFB6FAF9FAD0FDFAFAD2" box="[422,576,1291,1315]" count="2" pageId="20" pageNumber="21" type="generic">2 specimens</specimenCount>
, Kozatskyi Yar,
<collectionCode id="ED255488FE5DFFB6FBC8FAF7FEADFAB2" box="[151,279,1324,1347]" pageId="20" pageNumber="21">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE5DFFB6FA7DFAF7FEC0FAB2" box="[290,378,1323,1347]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="20" pageNumber="21">PI 2579</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B5CC610FE5DFFB6FAD7FAF0FF56FA92" collectionCode="NMNHU-P" pageId="20" pageNumber="21" specimenCode="PI 2580" specimenCount="3">
<specimenCount id="9D3207C4FE5DFFB6FAD7FAF0FDAEFAB2" box="[392,532,1323,1347]" count="3" pageId="20" pageNumber="21" type="generic">3 specimens</specimenCount>
, Mlyntsi,
<collectionCode id="ED255488FE5DFFB6F9D8FAF7FCBDFAB2" box="[647,775,1324,1347]" pageId="20" pageNumber="21">NMNHU-P</collectionCode>
<specimenCode id="DB926436FE5DFFB6FBC8FA97FF56FA92" box="[151,236,1355,1379]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="20" pageNumber="21">PI 2580</specimenCode>
</materialsCitation>
.
</paragraph>
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<subSubSection id="C32E9FC6FE5DFFB6FBC8FA57FD7CF992" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8B8BCC4DFE5DFFB6FBC8FA57FCBDFA52" blockId="20.[151,775,1419,1635]" box="[151,775,1419,1443]" pageId="20" pageNumber="21">
<heading id="D0C37B21FE5DFFB6FBC8FA57FCBDFA52" box="[151,775,1419,1443]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="21" reason="8">
<emphasis id="B940105FFE5DFFB6FBC8FA57FEB0FA52" box="[151,266,1420,1443]" italics="true" pageId="20" pageNumber="21">Discussion</emphasis>
Four small, juvenile, partly eroded specimens
</heading>
</paragraph>
<paragraph id="8B8BCC4DFE5DFFB6FBC8FA70FD7CF992" blockId="20.[151,775,1419,1635]" pageId="20" pageNumber="21">
(
<figureCitation id="130FD0C8FE5DFFB6FBC0FA70FF54FA32" box="[159,238,1451,1475]" captionStart="Fig" captionStartId="21.[168,199,1587,1607]" captionTargetBox="[152,1438,237,1698]" captionTargetId="figure-47@21.[167,1422,253,1574]" captionTargetPageId="21" captionText="Fig. 9 Otoliths of Haemulidae, Serranidae, Sparidae, Leiognathidae and Caproidae:a (reversed)—Brachydeuterus speronatus (Bassoli,1906), Kozatskyi Yar, NMNHU-P PI 2549. b, c Serranidae indet., b (broken and repaired), c (reversed), Kozatskyi Yar,NMNHU-P PI 2579. d, e Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, Mlyntsi, NMNHU-P PI 2578. f (reversed)—Leiognathidae indet., Shydlivshchyna, NMB P1214. g Perciformes indet., Shydlivshchyna,NMB P1221. h (reversed) Capros aper (Linnaeus,1758), Mlyntsi, NMB P1206" figureDoi="http://doi.org/10.5281/zenodo.11999944" httpUri="https://zenodo.org/record/11999944/files/figure.png" pageId="20" pageNumber="21">Fig. 9c</figureCitation>
) and a single large but incomplete and fragmented specimen (
<figureCitation id="130FD0C8FE5DFFB6FA32FA10FE01FA12" box="[365,443,1483,1507]" captionStart="Fig" captionStartId="21.[168,199,1587,1607]" captionTargetBox="[152,1438,237,1698]" captionTargetId="figure-47@21.[167,1422,253,1574]" captionTargetPageId="21" captionText="Fig. 9 Otoliths of Haemulidae, Serranidae, Sparidae, Leiognathidae and Caproidae:a (reversed)—Brachydeuterus speronatus (Bassoli,1906), Kozatskyi Yar, NMNHU-P PI 2549. b, c Serranidae indet., b (broken and repaired), c (reversed), Kozatskyi Yar,NMNHU-P PI 2579. d, e Pshekharus yesinorum Bannikov &amp; Kotlyar, 2015, Mlyntsi, NMNHU-P PI 2578. f (reversed)—Leiognathidae indet., Shydlivshchyna, NMB P1214. g Perciformes indet., Shydlivshchyna,NMB P1221. h (reversed) Capros aper (Linnaeus,1758), Mlyntsi, NMB P1206" figureDoi="http://doi.org/10.5281/zenodo.11999944" httpUri="https://zenodo.org/record/11999944/files/figure.png" pageId="20" pageNumber="21">Fig. 9b</figureCitation>
) are tentatively placed in the family
<taxonomicName id="4C34B7CEFE5DFFB6FBBAFA30FEE0F9F2" authorityName="Swainson" authorityYear="1839" box="[229,346,1515,1539]" family="Serranidae" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="21" phylum="Chordata" rank="family">Serranidae</taxonomicName>
but cannot be further identified. Te otoliths are characterized by a relatively mild bend in the horizontal direction, distinct mid- and postdorsal angles, and a relatively short and only slightly flexed cauda.
</paragraph>
</subSubSection>
</treatment>
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