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<document id="C19E6B601121BD6B8B3FB3F6E4A3FCF1" ID-DOI="10.5281/zenodo.173668" ID-GBIF-Dataset="8ce269f9-2380-43d9-b3ca-b8d799e9dd65" ID-ISSN="1175­5326" ID-Zenodo-Dep="173668" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459782665572" checkinUser="plazi" docAuthor="Nieto, Carolina &amp; Salles, Frederico Falcaõ" docDate="2006" docId="D155154796637B02FE90A885FBB98843" docLanguage="en" docName="zt01303p033.pdf" docOrigin="Zootaxa 1303" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Paracloeodes pacawara Nieto &amp; Salles, 2006, sp. n." docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="2D6C6D3F96697B09FF98AD21FFA08D5D" masterDocTitle="Revision of the Genus Paracloeodes (Ephemeroptera: Baetidae) in South America" masterLastPageNumber="33" masterPageNumber="1" pageNumber="11" updateTime="1698217998394" updateUser="plazi">
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<mods:title id="D85E0D40609D6DAD668437E44D21ACC2">Revision of the Genus Paracloeodes (Ephemeroptera: Baetidae) in South America</mods:title>
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<mods:namePart id="98781393E14CAE3E22F9A4F85E075BA6">Nieto, Carolina</mods:namePart>
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<mods:namePart id="D1549F6B7E0EC5ACD51B50313777689B">Salles, Frederico Falcaõ</mods:namePart>
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<mods:date id="2A503EF0ABF2E0DC16013FDAE1DF7DE2">2006</mods:date>
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<mods:identifier id="0BD91885B00D9498F85B69490FEAA8FB" type="ISSN">1175­5326</mods:identifier>
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<treatment id="D155154796637B02FE90A885FBB98843" ID-DOI="http://doi.org/10.5281/zenodo.5659796" ID-GBIF-Taxon="119348395" ID-Zenodo-Dep="5659796" LSID="urn:lsid:plazi:treatment:D155154796637B02FE90A885FBB98843" httpUri="http://treatment.plazi.org/id/D155154796637B02FE90A885FBB98843" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="11E6F7DA96637B03FE90A885FD278A2C" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="5943A45196637B03FE90A885FCB688E3" blockId="10.[264,790,1444,1470]" box="[264,790,1444,1470]" pageId="10" pageNumber="11">
<heading id="020B133D96637B03FE90A885FCB688E3" bold="true" box="[264,790,1444,1470]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="6B88784396637B03FE90A885FCB688E3" bold="true" box="[264,790,1444,1470]" pageId="10" pageNumber="11">
<taxonomicName id="9EFCDFD296637B03FE90A885FD8588E3" ID-CoL="4CQL4" box="[264,549,1444,1470]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pacawara" status="sp. nov.">
<emphasis id="6B88784396637B03FE90A885FD8588E3" bold="true" box="[264,549,1444,1470]" italics="true" pageId="10" pageNumber="11">Paracloeodes pacawara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BBC53896637B03FDB5A885FDCF88E3" box="[557,623,1444,1470]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="C1C7B8D496637B03FDE7A885FCAE88E3" box="[639,782,1444,1470]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="10" pageNumber="11">Figs. 50–62</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="5943A45196637B03FE90A8CFFD278A2C" blockId="10.[264,1323,1518,1905]" pageId="10" pageNumber="11">
<emphasis id="6B88784396637B03FE90A8CFFEC58B55" bold="true" box="[264,357,1518,1544]" pageId="10" pageNumber="11">Nymph</emphasis>
(
<figureCitation id="C1C7B8D496637B03FEE4A8CEFE7B8B54" box="[380,475,1519,1545]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="10" pageNumber="11">Fig. 50</figureCitation>
). Length: body, 3.0–
<quantity id="9E0409B496637B03FD7AA8CEFCE58B54" box="[738,837,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="11" unit="mm" value="3.4">3.4 mm</quantity>
; cerci,
<quantity id="9E0409B496637B03FC3FA8CEFB9E8B54" box="[935,1086,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="10" pageNumber="11" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
; terminal filament,
<quantity id="9E0409B496637B03FE90AB36FE388B6C" box="[264,408,1559,1585]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
. Antennae, 1.0–
<quantity id="9E0409B496637B03FDC1AB36FD148B6C" box="[601,692,1559,1585]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="11" unit="mm" value="1.1">1.1 mm</quantity>
; 1.5–1.8 times the head capsule. Head yellowish with small dark spots, ocelli dark brown. Turbinate portion of compound eyes in male nymphs yellowish­brown. Frontal keel present. Antennae pale yellow. Mouthparts: labrum (
<figureCitation id="C1C7B8D496637B03FB63AB46FE878BF4" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="10" pageNumber="11">Fig. 51</figureCitation>
a), dorsally without a pair of subapical setae near middle. Mandibles (
<figureCitation id="C1C7B8D496637B03FB1FABAEFABC8BF4" box="[1159,1308,1679,1705]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="10" pageNumber="11">Figs. 52–53</figureCitation>
): external margins straight. Left mandible (
<figureCitation id="C1C7B8D496637B03FC95AB96FCC98B8C" box="[781,873,1719,1745]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="10" pageNumber="11">Fig. 52</figureCitation>
) with prostheca with 2–3 denticles. Hypopharynx with lingua (
<figureCitation id="C1C7B8D496637B03FDFBABFEFD638BA4" box="[611,707,1759,1785]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="10" pageNumber="11">Fig. 54</figureCitation>
) subequal to superlingua and rounded apically. Maxillae (
<figureCitation id="C1C7B8D496637B03FE1AAA26FE788A7C" box="[386,472,1799,1825]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="10" pageNumber="11">Fig. 55</figureCitation>
) with palpi 1.5 times the length of galea­lacinia. Labium (
<figureCitation id="C1C7B8D496637B03FB0DAA26FB578A7C" box="[1173,1271,1799,1825]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="10" pageNumber="11">Figs. 56</figureCitation>
a–b) with segment II of palpi with a strong rounded distomedial projection, 3.1 times width of segment III, segment III conical.
</paragraph>
</subSubSection>
<subSubSection id="11E6F7DA96627B02FEA0AC3DFB248FC3" pageId="11" pageNumber="12" type="description">
<paragraph id="5943A45196627B02FEA0AC3DFE7A8C8B" blockId="11.[264,1323,284,1310]" pageId="11" pageNumber="12">
Thorax (
<figureCitation id="C1C7B8D496627B02FE39AC3DFE5F8C6B" box="[417,511,284,310]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 50</figureCitation>
): pro­, meso­ and metanota yellowish­brown with small brownish spots distributed throughout, and with a narrow pale yellow median band. Pleura yellowish­brown, sterna pale yellow. Legs (
<figureCitation id="C1C7B8D496627B02FC86AC4DFCD88CDB" box="[798,888,364,390]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 57</figureCitation>
) pale yellow, with tarsal claws 0.5 times the length of tarsi and with two rows of minute denticles basally (
<figureCitation id="C1C7B8D496627B02FBE4ACB5FB778CF3" box="[1148,1239,404,430]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 58</figureCitation>
).
<collectingCountry id="21EBE4C196627B02FB77ACB5FA8A8CF3" box="[1263,1322,404,430]" name="India" pageId="11" pageNumber="12">Hind</collectingCountry>
wing pads absent.
</paragraph>
<paragraph id="5943A45196627B02FEA0ACC5FB248FC3" blockId="11.[264,1323,284,1310]" pageId="11" pageNumber="12">
Abdomen: segments II–III and IX brownish, remaining segments yellowish­brown, all segments with small brownish spots as in
<figureCitation id="C1C7B8D496627B02FD6DAF2DFCEC8F7B" box="[757,844,524,550]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 50</figureCitation>
. Posterior margin of terga with spines as long as wide (
<figureCitation id="C1C7B8D496627B02FE2FAF15FDB38F13" box="[439,531,564,590]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 59</figureCitation>
). Sterna pale yellow, anterior margin of each segment and entire sterna IX–X brownish. Gills whitish (
<figureCitation id="C1C7B8D496627B02FD42AF7DFC978F2B" box="[730,823,604,630]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 60</figureCitation>
). Paraprocts with 9–10 spines (
<figureCitation id="C1C7B8D496627B02FB26AF7DFABA8F2B" box="[1214,1306,604,630]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 61</figureCitation>
). Caudal filaments yellowish with crown of spines on each segment (
<figureCitation id="C1C7B8D496627B02FB87AFA5FBD78FC3" box="[1055,1143,644,670]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11E6F7DA96627B02FEA0AF8DFD9C8FB3" pageId="11" pageNumber="12" type="etymology">
<paragraph id="5943A45196627B02FEA0AF8DFD9C8FB3" blockId="11.[264,1323,284,1310]" pageId="11" pageNumber="12">
<emphasis id="6B88784396627B02FEA0AF8DFE648F9B" bold="true" box="[312,452,684,710]" pageId="11" pageNumber="12">Etymology.</emphasis>
<taxonomicName id="9EFCDFD296627B02FE56AF8DFD9E8F9B" box="[462,574,684,710]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="29" phylum="Arthropoda" rank="species" species="pacawara" status="sp. nov.">Pacawara</taxonomicName>
is the name of a tribe that inhabit the north of
<collectingCountry id="21EBE4C196627B02FBE4AF8DFB758F9B" box="[1148,1237,684,710]" name="Bolivia" pageId="11" pageNumber="12">Bolivia</collectingCountry>
, where this species was collected.
</paragraph>
</subSubSection>
<subSubSection id="11E6F7DA96627B02FEA0AFDDFC7C8923" pageId="11" pageNumber="12" type="discussion">
<paragraph id="5943A45196627B02FEA0AFDDFC7C8923" blockId="11.[264,1323,284,1310]" pageId="11" pageNumber="12">
<emphasis id="6B88784396627B02FEA0AFDDFE638E4B" bold="true" box="[312,451,764,790]" pageId="11" pageNumber="12">Discussion.</emphasis>
This species can be distinguished from the other species of the genus by the following combination of characters, 1) frontal keel present; 2) antennae 1.5–1.8 times the head capsule; 3) labrum without a pair of subapical setae near middle (
<figureCitation id="C1C7B8D496627B02FB08AE6DFB498E3B" box="[1168,1257,844,870]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 51</figureCitation>
a); 4) lingua (
<figureCitation id="C1C7B8D496627B02FEF9AE55FE168ED3" box="[353,438,884,910]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 54</figureCitation>
) subequal to superlingua and rounded apically; 5) maxillary palpi 1.5 times the length of galea­lacinia (
<figureCitation id="C1C7B8D496627B02FDC1AEBDFD158EEB" box="[601,693,924,950]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 55</figureCitation>
); 6) segment II of labial palpi with strong rounded distomedial projection 3.1 times width of segment III (
<figureCitation id="C1C7B8D496627B02FC1BAEE5FC798E83" box="[899,985,964,990]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 56</figureCitation>
b); 7) hind wing pads absent; 8) tarsal claws long, 0.5 times the length of tarsi (
<figureCitation id="C1C7B8D496627B02FCFAAECDFC19895B" box="[866,953,1004,1030]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 57</figureCitation>
) and with two rows of minute denticles basally (
<figureCitation id="C1C7B8D496627B02FE45A935FD968973" box="[477,566,1044,1070]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 58</figureCitation>
); 9) posterior margin of abdominal terga with spines as long as wide (
<figureCitation id="C1C7B8D496627B02FECAA91DFE0B890B" box="[338,427,1084,1110]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 59</figureCitation>
); 10) caudal filaments with crown of spines on each segment (
<figureCitation id="C1C7B8D496627B02FB0BA91DFB4C890B" box="[1171,1260,1084,1110]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 62</figureCitation>
); 11) body color pattern with small dark spots throughout (
<figureCitation id="C1C7B8D496627B02FCEFA945FC6E8923" box="[887,974,1124,1150]" captionStart="FIGURE 50 – 62" captionStartId="28.[264,368,1721,1745]" captionTargetBox="[314,1219,325,1685]" captionTargetId="figure@28.[107,1283,310,1716]" captionTargetPageId="28" captionText="FIGURE 50 – 62. Paracloeodes pacawara sp. n. Nymph. 50, General view. Mouthparts (Figs. 51 – 56): 51 a, labrum, left d. v., right v. v.; 51 b, labrum: apical setae; 52, left mandible v. v.; 53, right mandible v. v.; 54, hypopharynx v. v.; 55, maxilla v. v.; 56 a, labium, left d. v., right v. v.; 56 b, palp: segments II – III. 57, leg I; 58, tarsal claw I. 59, posterior margin of tergum IV. 60, gill IV. 61, paraproct. 62, detail of cercus." httpUri="https://zenodo.org/record/173674/files/figure.png" pageId="11" pageNumber="12">Fig. 50</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="11E6F7DA96627B02FEA0A9ADFBB98843" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="5943A45196627B02FEA0A9ADFBB98843" blockId="11.[264,1323,284,1310]" pageId="11" pageNumber="12">
<emphasis id="6B88784396627B02FEA0A9ADFE0C89FB" bold="true" box="[312,428,1164,1190]" pageId="11" pageNumber="12">Material.</emphasis>
<typeStatus id="86471AF396627B02FE2FA9ADFD8489FB" box="[439,548,1164,1190]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
male nymph:
<collectingCountry id="21EBE4C196627B02FD4FA9ADFCF389FB" box="[727,851,1164,1190]" name="Bolivia" pageId="11" pageNumber="12">BOLIVIA</collectingCountry>
, Depto. Beni, Prov. Ballivian, Río San Bernardo,
<geoCoordinate id="3CC8C29696627B02FE1BA995FDBD8993" box="[387,541,1204,1230]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="-14.754444">S 14°45´16´´</geoCoordinate>
,
<geoCoordinate id="3CC8C29696627B02FDB1A995FD6F8993" box="[553,719,1204,1230]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-67.16861">W 67°10´07´´</geoCoordinate>
,
<quantity id="9E0409B496627B02FD42A995FC878993" box="[730,807,1204,1230]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="12" unit="m" value="220.0">220 m</quantity>
,
<date id="2D42829196627B02FCAAA995FC1F8993" box="[818,959,1204,1230]" pageId="11" pageNumber="12" value="2002-05-22">22/ V/ 2002</date>
, Domínguez col.
<typeStatus id="86471AF396627B02FB14A995FAA28993" box="[1164,1282,1204,1230]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
: 44 nymphs same data and collector. The
<typeStatus id="86471AF396627B02FD48A9FDFC9689AB" box="[720,822,1244,1270]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
and 5
<typeStatus id="86471AF396627B02FC14A9FDFC5C89AB" box="[908,1020,1244,1270]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
are deposited at
<collectionCode id="3FED3C9496627B02FB54A9FDFA8589AB" box="[1228,1317,1244,1270]" httpUri="http://grbio.org/cool/jxh3-1jar" name="Instituto de Ecologia" pageId="11" pageNumber="12">UMSA</collectionCode>
. Five
<typeStatus id="86471AF396627B02FEDBA825FE138843" box="[323,435,1284,1310]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
deposited at
<collectionCode id="3FED3C9496627B02FDD7A825FD2E8843" box="[591,654,1284,1310]" pageId="11" pageNumber="12">IBRJ</collectionCode>
, other
<typeStatus id="86471AF396627B02FD45A825FCED8843" box="[733,845,1284,1310]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
housed at
<collectionCode id="3FED3C9496627B02FC54A825FBB68843" box="[972,1046,1284,1310]" pageId="11" pageNumber="12">IFML</collectionCode>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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