172 lines
21 KiB
XML
172 lines
21 KiB
XML
<document id="8A1ACBE12C9E3B123B95C5CD02B5EB34" ID-CLB-Dataset="131926" ID-DOI="10.11646/zootaxa.5178.4.4" ID-GBIF-Dataset="69f03f3e-68ec-43c9-a3d3-75927165afae" ID-ISSN="1175-5326" ID-Zenodo-Dep="7031814" ID-ZooBank="60009AB8-4DAC-418E-8378-9155DDDD024B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661769161681" checkinUser="plazi" docAuthor="Nakamine, Hiroshi, Yamamoto, Shûhei & Takahashi, Yui" docDate="2022" docId="D679951B82334C6EFF765D1DE3FCCA06" docLanguage="en" docName="zootaxa.5178.4.4.pdf" docOrigin="Zootaxa 5178 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemerobiidae Latreille 1802" docType="treatment" docVersion="9" lastPageNumber="388" masterDocId="2A40ED6382354C66FFE15A4CE77CC81E" masterDocTitle="Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae)" masterLastPageNumber="390" masterPageNumber="380" pageNumber="386" updateTime="1699374251715" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="093045877F87DD7AF53CA0C03BEA6BC9">
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<mods:title id="D330BE2A86FCE470ECDF04BA4F89FD80">Archaeomegalomus gen. nov.: A remarkable new brown lacewing from mid-Cretaceous Kachin amber from northern Myanmar (Neuroptera: Hemerobiidae)</mods:title>
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<mods:namePart id="0791982A7EA8924E32C7B3FB32BCD63C">Nakamine, Hiroshi</mods:namePart>
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<mods:affiliation id="5BC75C15B16406317C4FC0954BEC8A64">Minoh Park Insect Museum, Minoh Park 1 - 18, Minoh City, Osaka 562 - 0002, Japan.</mods:affiliation>
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<mods:namePart id="61D7B26CFDDEAAF172F9E5CC8AB7546E">Yamamoto, Shûhei</mods:namePart>
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<mods:affiliation id="454E9599DCA70B19978B62C63CA8CB54">Hokkaio University Museum, Hokkaido University, Kita 10 - jo, Nishi 8 - chome, Kita-ku, Sapporo 060 - 0810, Japan.</mods:affiliation>
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<mods:namePart id="8628E27BA24798F3280A7FD69A7515F6">Takahashi, Yui</mods:namePart>
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<mods:affiliation id="62E91B647E474C44D9B38E1D1BF9CD30">Keio Yochisha Elementary School, Ebisu 2 - 35 - 1, Shibuya-ku, Tokyo 150 - 0013, Japan.</mods:affiliation>
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<mods:typeOfResource id="46EEED5B74F3BE155D3DFC2FFC80457B">text</mods:typeOfResource>
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<mods:title id="FBDA37BBDCF9902231F1006700E92E5E">Zootaxa</mods:title>
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<mods:date id="52A72AB5D917D6B645EA0DF243D4C10A">2022</mods:date>
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<mods:number id="E41F1D267D4FB5FC1D1372C4DCFB92E0">2022-08-29</mods:number>
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<treatment id="D679951B82334C6EFF765D1DE3FCCA06" ID-DOI="http://doi.org/10.5281/zenodo.7037082" ID-GBIF-Taxon="199861242" ID-Zenodo-Dep="7037082" LSID="urn:lsid:plazi:treatment:D679951B82334C6EFF765D1DE3FCCA06" httpUri="http://treatment.plazi.org/id/D679951B82334C6EFF765D1DE3FCCA06" lastPageId="8" lastPageNumber="388" pageId="6" pageNumber="386">
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<subSubSection id="16CA778682334C60FF765D1DE565CF75" box="[151,537,1873,1899]" pageId="6" pageNumber="386" type="nomenclature">
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<paragraph id="5E6F240D82334C60FF765D1DE565CF75" blockId="6.[151,537,1873,1935]" box="[151,537,1873,1899]" pageId="6" pageNumber="386">
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<heading id="0527936182334C60FF765D1DE565CF75" bold="true" box="[151,537,1873,1899]" fontSize="11" level="1" pageId="6" pageNumber="386" reason="1">
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<emphasis id="6CA4F81F82334C60FF765D1DE565CF75" bold="true" box="[151,537,1873,1899]" pageId="6" pageNumber="386">
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<taxonomicName id="99D05F8E82334C60FF765D1DE634CF75" ID-CoL="624S6" ID-ENA="50435" authorityName="Latreille" authorityYear="1802" box="[151,328,1873,1899]" class="Insecta" family="Hemerobiidae" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="family">Hemerobiidae</taxonomicName>
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gen. et sp. indet.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="16CA778682334C61FF765D39E352C97B" lastPageId="7" lastPageNumber="387" pageId="6" pageNumber="386" type="description">
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<paragraph id="5E6F240D82334C60FF765D39E797CF91" blockId="6.[151,537,1873,1935]" box="[151,235,1909,1935]" pageId="6" pageNumber="386">
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(
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<figureCitation id="C6EB388882334C60FF7E5D39E79FCF91" box="[159,227,1909,1935]" captionStart="FIGURE 4" captionStartId="6.[151,250,1700,1724]" captionTargetBox="[151,1436,777,1677]" captionTargetId="figure-248@6.[151,1436,777,1677]" captionTargetPageId="6" captionText="FIGURE 4. Habitus and Detail of Hemerobiidae gen. et sp. indet. (AMNH Bu-SY32) in mid-Cretaceous Kachin amber.A, right lateral view; B, left lateral view of terminalia. Abbreviations: ect: ectoproct; gx9: gonocoxite 9; st: gonostylus of gonocoxite 9. Scale bars: 0.5 mm for A; 0.2 mm for B." figureDoi="http://doi.org/10.5281/zenodo.7031823" httpUri="https://zenodo.org/record/7031823/files/figure.png" pageId="6" pageNumber="386">Fig. 4</figureCitation>
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)
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</paragraph>
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<paragraph id="5E6F240D82334C61FF765DF1E352C97B" blockId="6.[151,1436,1981,2043]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="387" pageId="6" pageNumber="386">
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<emphasis id="6CA4F81F82334C60FF765DF1E655CFC9" bold="true" box="[151,297,1981,2007]" pageId="6" pageNumber="386">Description.</emphasis>
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The body and wings are not clearly visible due to many cracks. Length ca.
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<quantity id="992889E882334C60FB4B5DF1E278CFC9" box="[1194,1284,1981,2007]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="386" unit="mm" value="3.6">3.6 mm</quantity>
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as preserved (measured from vertex to apex of the abdomen). Head poorly preserved; eyes large; antennae composed at least 27 flagellomeres with scattered fine setae on each segment. Thorax covered with dense thin setae; pronotum rather short. All legs slender, covered with dense thin setae; all tibia slightly swollen at the middle with paired tibial spurs. Wings poorly preserved; forewing hyaline, oval, ca.
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<quantity id="992889E882324C61FD045A92E440C8E7" box="[741,828,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="387" unit="mm" value="3.5">3.5 mm</quantity>
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long; trichosors present along the entire wing margin, one trichosor between adjacent veins, marginal portions of veins and trichosors with tufts of thin setae, humeral veinlets recurrent, costal space rather broad; hind wing hyaline, oval, ca. 3.0 mm long. Abdomen poorly preserved; terminal segment scattered with thin setae; gonostylus of gonocoxite 9 developed.
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</paragraph>
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</subSubSection>
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<caption id="0AAF748582324C61FF765D1DE50ACFE7" ID-DOI="http://doi.org/10.5281/zenodo.7031825" ID-Zenodo-Dep="7031825" httpUri="https://zenodo.org/record/7031825/files/figure.png" pageId="7" pageNumber="387" startId="7.[151,250,1873,1897]" targetBox="[151,1436,381,1850]" targetPageId="7">
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<paragraph id="5E6F240D82324C61FF765D1DE50ACFE7" blockId="7.[151,1436,1873,2041]" pageId="7" pageNumber="387">
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<emphasis id="6CA4F81F82324C61FF765D1DE66FCF77" bold="true" box="[151,275,1873,1897]" pageId="7" pageNumber="387">FIGURE 5.</emphasis>
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Distribution map and forewing venation of Mesozoic
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<taxonomicName id="99D05F8E82324C61FCDD5D1DE4B3CF77" authorityName="Latreille" authorityYear="1802" box="[828,975,1873,1897]" class="Insecta" family="Hemerobiidae" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="387" phylum="Arthropoda" rank="family">Hemerobiidae</taxonomicName>
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. A, distribution map of Mesozoic
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<taxonomicName id="99D05F8E82324C61FAC65D1DE7C1CF93" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Hemerobiidae" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="387" phylum="Arthropoda" rank="family">Hemerobiidae</taxonomicName>
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; B,
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<taxonomicName id="99D05F8E82324C61FF055D39E699CF93" box="[228,485,1909,1933]" class="Insecta" family="Hemerobiidae" genus="Promegalomus" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="387" phylum="Arthropoda" rank="species" species="anomalus">
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<emphasis id="6CA4F81F82324C61FF055D39E699CF93" box="[228,485,1909,1933]" italics="true" pageId="7" pageNumber="387">Promegalomus anomalus</emphasis>
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</taxonomicName>
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(modified after
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<bibRefCitation id="3A4159FC82324C61FD665D39E46BCF93" author="Panfilov, D. V." box="[647,791,1909,1933]" pageId="7" pageNumber="387" pagination="82 - 111" refId="ref5584" refString="Panfilov, D. V. (1980) New representatives of lacewings (Neuroptera) from the Jurassic of Karatau. In: Dolin, V. G., Panfilov, D. V., Ponomarenko, A. G. & Pritykina, L. N. (Eds.), Fossil insects of the Mesozoic. Naukova Dumka, Kiev, pp. 82 - 111. [in Russian]" type="book chapter" year="1980">Panfilov 1980</bibRefCitation>
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, fig. 91); C,
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<taxonomicName id="99D05F8E82324C61FC755D39E3EACF93" box="[916,1174,1909,1933]" class="Insecta" family="Hemerobiidae" genus="Cretomerobius" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="387" phylum="Arthropoda" rank="species" species="disjunctus">
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<emphasis id="6CA4F81F82324C61FC755D39E3EACF93" box="[916,1174,1909,1933]" italics="true" pageId="7" pageNumber="387">Cretomerobius disjunctus</emphasis>
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</taxonomicName>
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(modified after
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<bibRefCitation id="3A4159FC82324C61FAD85D3AE67FCFAF" author="Ponomarenko, A. G." pageId="7" pageNumber="387" refId="ref5760" refString="Ponomarenko, A. G. (1992) New lacewings (Insecta, Neuroptera) from the Mesozoic of Mongolia. In: Grunt, T. A. (Ed.), New taxa of the fossil invertebrates of Mongolia. Transactions of the Joint Soviet - Mongolian Paleontological Expedition. Vol. 41. Nauka Press, Moscow, pp. 101 - 111. [in Russian]" type="book" year="1992">Ponomarenko 1992</bibRefCitation>
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, fig. 4); D,
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<emphasis id="6CA4F81F82324C61FE975DD6E5C4CFAF" box="[374,696,1945,1969]" italics="true" pageId="7" pageNumber="387">
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<taxonomicName id="99D05F8E82324C61FE975DD6E55CCFAF" authority="Nakamine & Yamamoto & Takahashi, 2022" authorityName="Nakamine & Yamamoto & Takahashi" authorityYear="2022" box="[374,544,1946,1969]" class="Insecta" family="Hemerobiidae" genus="Cretoneuronema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="389" phylum="Arthropoda" rank="genus" status="gen. nov.">Cretoneuronema</taxonomicName>
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jarzembowskii
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</emphasis>
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(modified after
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<bibRefCitation id="3A4159FC82324C61FCBD5DD5E48ECFAF" author="Liu, X. Y. & Chen, Z. L. & Zhuo, D." box="[860,1010,1945,1969]" pageId="7" pageNumber="387" pagination="226 - 232" refId="ref5207" refString="Liu, X. Y., Chen, Z. L. & Zhuo, D. (2022) Cretoneuronema gen. nov. (Neuroptera: Hemerobiidae), a new brown lacewing genus from the mid-Cretaceous Kachin amber. Palaeoentomology, 5 (3), 226 - 232. https: // doi. org / 10.11646 / palaeoentomology. 5.3.4" type="journal article" year="2022">
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Liu
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<emphasis id="6CA4F81F82324C61FC645DD6E4C4CFAF" box="[901,952,1945,1969]" italics="true" pageId="7" pageNumber="387">et al.</emphasis>
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2022
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</bibRefCitation>
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, fig. 3); E,
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<taxonomicName id="99D05F8E82324C61FB835DD5E2E0CFAF" authority="Nakamine & Yamamoto & Takahashi, 2022" authorityName="Nakamine & Yamamoto & Takahashi" authorityYear="2022" box="[1122,1436,1945,1969]" class="Insecta" family="Hemerobiidae" genus="Archaeomegalomus" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="387" phylum="Arthropoda" rank="species" species="burmiticus" status="gen. et sp. nov.">
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<emphasis id="6CA4F81F82324C61FB835DD5E2E0CFAF" box="[1122,1436,1945,1969]" italics="true" pageId="7" pageNumber="387">Archaeomegalomus burmiticus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7797456482324C61FF765DF2E653CFCB" box="[151,303,1982,2006]" pageId="7" pageNumber="387" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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; F,
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<taxonomicName id="99D05F8E82324C61FEAE5DF1E52CCFCB" authorityName="Makarkin & Grohn" authorityYear="2020" box="[335,592,1981,2005]" class="Insecta" family="Hemerobiidae" genus="Hemeroberotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="387" phylum="Arthropoda" rank="species" species="sinefurca">
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<emphasis id="6CA4F81F82324C61FEAE5DF1E52CCFCB" box="[335,592,1981,2005]" italics="true" pageId="7" pageNumber="387">Hemeroberotha sinefurca</emphasis>
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</taxonomicName>
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(modified after
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<bibRefCitation id="3A4159FC82324C61FD135DF1E491CFCB" author="Makarkin, V. N. & Grohn, C." box="[754,1005,1981,2005]" pageId="7" pageNumber="387" pagination="104206" refId="ref5310" refString="Makarkin, V. N. & Grohn, C. (2020) The first unusual Hemerobiidae (Neuroptera) from mid-Cretaceous Burmese amber. Cretaceous Research, 106, 104206. https: // doi. org / 10.1016 / j. cretres. 2019.104206" type="journal article" year="2020">Makarkin & Gröhn 2020</bibRefCitation>
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, fig. 4); G,
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<taxonomicName id="99D05F8E82324C61FBBC5DF1E22FCFCB" box="[1117,1363,1981,2005]" class="Insecta" family="Berothidae" genus="Plesiorobius" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="387" phylum="Arthropoda" rank="species" species="canadensis">
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<emphasis id="6CA4F81F82324C61FBBC5DF1E22FCFCB" box="[1117,1363,1981,2005]" italics="true" pageId="7" pageNumber="387">Plesiorobius canadensis</emphasis>
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</taxonomicName>
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(modified after
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<bibRefCitation id="3A4159FC82324C61FF1B5DADE55BCFE7" author="Klimaszewski, J. & Kevan, D. K. M." box="[250,551,2017,2041]" pageId="7" pageNumber="387" pagination="124 - 132" refId="ref4964" refString="Klimaszewski, J. & Kevan, D. K. M. (1986) A new lacewing-fly (Neuroptera: Planipennia) from Canadian Cretaceous Amber, with an analysis of its fore wing characters. Entomological News, 97, 124 - 132." type="journal article" year="1986">Klimaszewski & Kevan 1986</bibRefCitation>
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, fig. 2).
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</paragraph>
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</caption>
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||
<subSubSection id="16CA7786823D4C6EFF265ADAE675C902" pageId="8" pageNumber="388" type="materials_examined">
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||
<paragraph id="5E6F240D823D4C6EFF265ADAE675C902" blockId="8.[151,1437,150,537]" pageId="8" pageNumber="388">
|
||
<emphasis id="6CA4F81F823D4C6EFF265ADAE64AC8AE" bold="true" box="[199,310,150,176]" pageId="8" pageNumber="388">Material.</emphasis>
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Incomplete female adult (
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<figureCitation id="C6EB3888823D4C6EFDBF5ADAE5DFC8AE" box="[606,675,150,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1700,1724]" captionTargetBox="[151,1436,777,1677]" captionTargetId="figure-248@6.[151,1436,777,1677]" captionTargetPageId="6" captionText="FIGURE 4. Habitus and Detail of Hemerobiidae gen. et sp. indet. (AMNH Bu-SY32) in mid-Cretaceous Kachin amber.A, right lateral view; B, left lateral view of terminalia. Abbreviations: ect: ectoproct; gx9: gonocoxite 9; st: gonostylus of gonocoxite 9. Scale bars: 0.5 mm for A; 0.2 mm for B." figureDoi="http://doi.org/10.5281/zenodo.7031823" httpUri="https://zenodo.org/record/7031823/files/figure.png" pageId="8" pageNumber="388">Fig. 4</figureCitation>
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), indiscernible many morphological details due to numerous cracks and overlapping wings and body, preserved in approximately
|
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<quantity id="992889E8823D4C6EFCA85AF6E4D8C8CB" box="[841,932,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="388" unit="mm" value="15.0">15 mm</quantity>
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||
×
|
||
<quantity id="992889E8823D4C6EFC5D5AF6E376C8CB" box="[956,1034,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="388" unit="mm" value="11.0">11 mm</quantity>
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||
×
|
||
<quantity id="992889E8823D4C6EFBC65AF6E300C8CB" box="[1063,1148,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="388" unit="mm" value="2.5">2.5 mm</quantity>
|
||
flat arcwise, dark yellow, somewhat transparent amber piece with mumerous debris and pigmentation; specimen accession number
|
||
<collectionCode id="38C1BCC8823D4C6EFAA65A92E2E1C8E6" box="[1351,1437,222,248]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="388">AMNH</collectionCode>
|
||
Bu-SY32.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="16CA7786823D4C6EFF265B6AE3FCCA06" pageId="8" pageNumber="388" type="discussion">
|
||
<paragraph id="5E6F240D823D4C6EFF265B6AE3FCCA06" blockId="8.[151,1437,150,537]" pageId="8" pageNumber="388">
|
||
<emphasis id="6CA4F81F823D4C6EFF265B6AE647C95E" bold="true" box="[199,315,294,320]" pageId="8" pageNumber="388">Remarks.</emphasis>
|
||
We assign this undetermined specimen to the
|
||
<taxonomicName id="99D05F8E823D4C6EFCB75B6AE489C95E" authorityName="Latreille" authorityYear="1802" box="[854,1013,294,320]" class="Insecta" family="Hemerobiidae" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="388" phylum="Arthropoda" rank="family">Hemerobiidae</taxonomicName>
|
||
because of the presence of a pair of tibial spurs, slightly swollen tibia at the middle, and recurred humeral veinlets. This undetermined specimen can be distinguished from the three burmite hemerobiids by the number of flagellomeres (this specimen has at least 27 or 28 flagellomeres, whereas there are
|
||
<quantity id="992889E8823D4C6EFDAA5BDEE5F8C9B2" box="[587,644,402,428]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="8" pageNumber="388" unit="in" value="38.0">38 in</quantity>
|
||
<taxonomicName id="99D05F8E823D4C6EFD6A5BDEE414C9B2" authorityName="Nakamine & Yamamoto & Takahashi" authorityYear="2022" box="[651,872,402,428]" class="Insecta" family="Hemerobiidae" genus="Archaeomegalomus" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="388" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6CA4F81F823D4C6EFD6A5BDEE414C9B2" box="[651,872,402,428]" italics="true" pageId="8" pageNumber="388">Archaeomegalomus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="992889E8823D4C6EFC975BDEE4CCC9B2" box="[886,944,402,428]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="8" pageNumber="388" unit="in" value="19.0">19 in</quantity>
|
||
<taxonomicName id="99D05F8E823D4C6EFC565BDEE31AC9B2" authorityName="Makarkin & Gröhn" authorityYear="2020" box="[951,1126,402,428]" class="Insecta" family="Hemerobiidae" genus="Hemeroberotha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="388" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6CA4F81F823D4C6EFC565BDEE31AC9B2" box="[951,1126,402,428]" italics="true" pageId="8" pageNumber="388">Hemeroberotha</emphasis>
|
||
</taxonomicName>
|
||
) and the costal space in the forewing (this specimen is rather broad, whereas it is very broad in
|
||
<taxonomicName id="99D05F8E823D4C6EFC625BFBE343C9CF" authority="Nakamine & Yamamoto & Takahashi, 2022" authorityName="Nakamine & Yamamoto & Takahashi" authorityYear="2022" box="[899,1087,439,465]" class="Insecta" family="Hemerobiidae" genus="Cretoneuronema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="389" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="6CA4F81F823D4C6EFC625BFBE343C9CF" box="[899,1087,439,465]" italics="true" pageId="8" pageNumber="388">Cretoneuronema</emphasis>
|
||
</taxonomicName>
|
||
). Thus, we withheld identification of the specimen as we could not confirm the important traits of the wing venation, although we were able to differentiate it from the three previously recorded fossil species found in Burmese amber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |