193 lines
20 KiB
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193 lines
20 KiB
XML
<document ID-CLB-Dataset="21365" ID-DOI="http://dx.doi.org/10.3897/zookeys.955.51983" ID-GBIF-Dataset="45a98afc-af5a-4734-93d1-205e1ac3694b" ID-PMC="PMC7423785" ID-Pensoft-Pub="1313-2970-955-1" ID-Pensoft-UUID="F7018CF19C6158309D57C83A2F5E9478" ID-PubMed="32855592" ID-ZooBank="C8EF4A4B6F054621B9B5AE47FEE7C217" ModsDocID="1313-2970-955-1" checkinTime="1596691600214" checkinUser="pensoft" docAuthor="Grego, Jozef, Mumladze, Levan, Falniowski, Andrzej, Osikowski, Artur, Rysiewska, Aleksandra, Palatov, Dimitry M. & Hofman, Sebastian" docDate="2020" docId="65E19D736DF65BBC8652527192705777" docLanguage="en" docName="ZooKeys 955: 1-77" docOrigin="ZooKeys 955" docSource="http://dx.doi.org/10.3897/zookeys.955.51983" docTitle="Caucasopsis Grego & Mumladze 2020, gen. nov." docType="treatment" docUuid="7FAE27A6-16B9-4946-A5E8-4CAD2DCF6C33" docUuidSource="ZooBank" docVersion="7" id="F7018CF19C6158309D57C83A2F5E9478" lastPageNumber="1" masterDocId="F7018CF19C6158309D57C83A2F5E9478" masterDocTitle="Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="1" updateTime="1732936627699" updateUser="ExternalLinkService">
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<mods:titleInfo id="F7ED3D287BD1D641628A64B12CA7774F">
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<mods:title id="ADA661132FE4037324B2EBF416E1EAA5">Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species</mods:title>
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<mods:name id="D30E501F6D69C52029283F59D0107F04" type="personal">
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<mods:roleTerm id="7B34217012917BDBD4B9AD2FA5F237FE">Author</mods:roleTerm>
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<mods:namePart id="3693338785318351B76A957286B6C5CC">Grego, Jozef</mods:namePart>
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<mods:affiliation id="B8EF25A359F577717AD537714077FCF3">Horna Micina 219, 97401, Banska Bystrica, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name id="0A1EF325C2E3F31E4B08A99BFE355DFE" type="personal">
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<mods:roleTerm id="D35946B366D9D2BBA9794C0BA762C0CF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="18595E216D575E59095B1FEE3B24AE75">Mumladze, Levan</mods:namePart>
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<mods:affiliation id="70E5D864AA86F209DB4FF771C92028D4">Institute of Zoology, Ilia State University, Kakutsa Cholokashvili Ave 3 / 5, Tbilisi 0162, Georgia</mods:affiliation>
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<mods:nameIdentifier id="3E5D4AB40CA12AE4101245F59CAC23A6" type="orcid">https://orcid.org/0000-0002-2172-6973</mods:nameIdentifier>
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</mods:name>
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<mods:name id="D930AAA24705D1F5473947157E73F7BA" type="personal">
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<mods:roleTerm id="514389C7FED87998A56192006B63F4BB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="45972454F0DEF5CF6866A55E99F490B8">Falniowski, Andrzej</mods:namePart>
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<mods:affiliation id="33C79BF854F71012D245E289A068581A">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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</mods:name>
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<mods:name id="08565D271F2A647A30C0A0261E31835A" type="personal">
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<mods:roleTerm id="97A6392E1F33F0DC7ACBA72866D16663">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9248AD305A214AFD351EDA628570D02E">Osikowski, Artur</mods:namePart>
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<mods:affiliation id="B13631616EBC0953A88EB1E06C28EE49">Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059, Krakow, Poland</mods:affiliation>
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<mods:nameIdentifier id="F59782303FBE3D9D9FA22C55C39D9C27" type="orcid">https://orcid.org/0000-0001-6646-2687</mods:nameIdentifier>
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<mods:nameIdentifier id="4524CC4F4DE404D611BB5305AD00CEAD" type="email">a.osikowski@urk.edu.pl</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="7B424F7757D7CE6A01CDEBBAC63B924F">Rysiewska, Aleksandra</mods:namePart>
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<mods:affiliation id="926A1CFF347630B8F9CD028DAB36E611">Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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</mods:name>
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<mods:namePart id="2F31BF1B8B63F05964317F56CF451A98">Palatov, Dimitry M.</mods:namePart>
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<mods:affiliation id="DCA4A2B70AF7ADDFEBC05ED81B55BF4C">Department of Hydrobiology, Biological Faculty, Moscow State University, 1 - 12 Leninskie Gory, 119991, Moscow, Russian Federation</mods:affiliation>
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</mods:name>
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<mods:name id="E7AC8DF033E65A87F80EE49D05220BF8" type="personal">
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<mods:namePart id="2817872B3D35CACAD2AB574CEA265F34">Hofman, Sebastian</mods:namePart>
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<mods:affiliation id="5817A021D25D1179F1B0E8F7F6C5E89C">Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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<mods:typeOfResource id="61C4C4547300B04194E6DACD4831E8B1">text</mods:typeOfResource>
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<mods:title id="227476BE5F53F6C8AE527685A40EBC36">ZooKeys</mods:title>
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<mods:part id="12CACE76D8E61CAF85EBFE1132327BD1">
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<mods:date id="F0E358DE415D0645DAC1B0C1EB494ACC">2020</mods:date>
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<mods:detail id="9FCB06BCB3EFE50469A7861E63FBED60" type="volume">
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<mods:number id="4343547C7D2519C9E7806E5DA49E67FB">955</mods:number>
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<mods:start id="A115101FEEF34E51EE7A9446C6B936FB">1</mods:start>
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<mods:end id="0B798762F9404534B2756330B4748B18">77</mods:end>
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<mods:url id="8B6E7DBEE58057454232F1CC46229605">http://dx.doi.org/10.3897/zookeys.955.51983</mods:url>
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<mods:classification id="AAF3FE924467CC318890C46C14652F41">journal article</mods:classification>
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<mods:identifier id="BBF3B3AE3C75F9FA6523DE1F7B5F0544" type="DOI">http://dx.doi.org/10.3897/zookeys.955.51983</mods:identifier>
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<mods:identifier id="681A18C16580951AE3BC846851DBB934" type="Pensoft-Pub">1313-2970-955-1</mods:identifier>
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<mods:identifier id="0ACA9EEF9934DE9DD44575793FA87BDA" type="ZooBank">C8EF4A4B6F054621B9B5AE47FEE7C217</mods:identifier>
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<mods:identifier id="28D888EB5D773BC8C99B2A84CB4E98E7" type="Pensoft-UUID">F7018CF19C6158309D57C83A2F5E9478</mods:identifier>
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</mods:mods>
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<treatment id="65E19D736DF65BBC8652527192705777" ID-GBIF-Taxon="166096049" LSID="urn:lsid:zoobank.org:act:7FAE27A6-16B9-4946-A5E8-4CAD2DCF6C33" httpUri="http://treatment.plazi.org/id/65E19D736DF65BBC8652527192705777" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Hydrobiidae" scope_phylum="Mollusca" scope_subFamily="Sadlerianinae" scope_superFamily="Truncatelloidea">
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<subSubSection id="1DAD87CA7FAF27E722B7A4D03CE7B16A" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="ECBE6C607E0B37DED73E7166E0F76D3A" pageId="0" pageNumber="1">
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Genus
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<taxonomicName id="2004B515992903BEBCB6A3F437206F61" ID-CoL="7NZX2" LSID="65E19D73-6DF6-5BBC-8652-527192705777" authority="Grego & Mumladze" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus" status="gen. nov.">Caucasopsis Grego & Mumladze</taxonomicName>
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<taxonomicNameLabel id="6A8ACB49E44BE3900112358F77DFE902" pageId="0" pageNumber="1">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="DC8FE921CCE6910ED95115A09C20705C" pageId="0" pageNumber="1" type="type species">
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<paragraph id="A01ED99558DEAB5F8A0C711907C474F0" pageId="0" pageNumber="1">Type species.</paragraph>
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<paragraph id="89403AA07CE028D9CDCB04E35CE610AF" pageId="0" pageNumber="1">
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<taxonomicName id="0DF55FDB64B8BAF10D47E84529120E9B" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasopsis letsurtsume" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="letsurtsume">
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<emphasis id="6DCAA26950096276FFC57F584EC56772" italics="true" pageId="0" pageNumber="1">Caucasopsis letsurtsume</emphasis>
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</taxonomicName>
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Grego & Mumladze, sp. nov.
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</paragraph>
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</subSubSection>
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<subSubSection id="1317A6E1BF771D2DD8AE760CFE6FC7C8" pageId="0" pageNumber="1" type="species assigned to the genus">
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<paragraph id="7F27E732755A1D5C3025D01578D8A06D" pageId="0" pageNumber="1">Species assigned to the genus.</paragraph>
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<paragraph id="335A827D498D53ABD7F47696AA4FC1D9" pageId="0" pageNumber="1">
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<taxonomicName id="22A354560780BC3C3F41EB4C2917B1A7" authorityName="Starobogatov" authorityYear="1962" class="Gastropoda" family="Moitessieriidae" genus="Paladilhiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paladilhiopsis shadini" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="shadini">
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<emphasis id="799934107CA95B60F809D8FD3C27D417" italics="true" pageId="0" pageNumber="1">Paladilhiopsis shadini</emphasis>
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</taxonomicName>
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Starobogatov, 1962,
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<taxonomicName id="FAC9F921BDA3C94656C2BFDAE06ECFF4" genus="Paldilhiopsis" lsidName="Paldilhiopsis subovata" pageId="0" pageNumber="1" rank="species" species="subovata">
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<emphasis id="AD2777ADE78F8E78664D140D8014EF21" italics="true" pageId="0" pageNumber="1">Paldilhiopsis subovata</emphasis>
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</taxonomicName>
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Starobogatov, 1962;
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<taxonomicName id="2BA21F77BADE754734F47566349B80AF" authorityName="Starobogatov" authorityYear="1962" class="Gastropoda" family="Moitessieriidae" genus="Paladilhiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paladilhiopsis pulcherrima" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pulcherrima">
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<emphasis id="F352E641AE339850967396624059B65E" italics="true" pageId="0" pageNumber="1">Paladilhiopsis pulcherrima</emphasis>
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</taxonomicName>
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Starobogatov, 1962;
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<taxonomicName id="4394D393B78703D1A5CE6D526C4179CA" authorityName="Starobogatov" authorityYear="1962" class="Gastropoda" family="Moitessieriidae" genus="Paladilhiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paladilhiopsis orientalis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="orientalis">
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<emphasis id="E0C1A6CA404D8431D3ADF90AF1AC755A" italics="true" pageId="0" pageNumber="1">Paladilhiopsis orientalis</emphasis>
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</taxonomicName>
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Starobogatov, 1962
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<taxonomicName id="4CE746E280345B40B02C54816580E307" authorityName="Starobogatov" authorityYear="1962" class="Gastropoda" family="Moitessieriidae" genus="Paladilhiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paladilhiopsis schakuranica" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="schakuranica">
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<emphasis id="B70208F65F3F55AA31CFEE671528AD01" italics="true" pageId="0" pageNumber="1">Paladilhiopsis schakuranica</emphasis>
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</taxonomicName>
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Starobogatov, 1962;
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<taxonomicName id="7C565028439C39FB5B803C229FF564D3" genus="Paladilhopsis" lsidName="Paladilhopsis aculeus" pageId="0" pageNumber="1" rank="species" species="aculeus">
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<emphasis id="5AD4120A59BBF88B56BF73BA4C0CC938" italics="true" pageId="0" pageNumber="1">Paladilhopsis aculeus</emphasis>
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</taxonomicName>
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Starobogatov, 1962;
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<taxonomicName id="909D48AF797690A73685DF726FBEAC1A" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasopsis letsurtsume" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="letsurtsume">
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<emphasis id="DB82C6CDC067B5DFE4E78FAD7480AB7C" italics="true" pageId="0" pageNumber="1">Caucasopsis letsurtsume</emphasis>
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</taxonomicName>
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Grego & Mumladze, sp. nov.,
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<taxonomicName id="6314C670E25E81661EF19691D97DE326" authorityName="Mumladze & Grego" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasopsis olsavskyi" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="olsavskyi">
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<emphasis id="0B30B272137AADE406A81DDE4F568A73" italics="true" pageId="0" pageNumber="1">Caucasopsis olsavskyi</emphasis>
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</taxonomicName>
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Grego & Mumladze, sp. nov.,
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<taxonomicName id="2936B33B20B88410CA27764297549F77" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasopsis egrisi" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="egrisi">
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<emphasis id="A82D2A81E963D70A9ACA27672ECBA68F" italics="true" pageId="0" pageNumber="1">Caucasopsis egrisi</emphasis>
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</taxonomicName>
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Grego & Mumladze, sp. nov.
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</paragraph>
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</subSubSection>
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<subSubSection id="9747CFC20E6501E681F048692E4A53A1" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="E73C40E16A8E2F6A8EE71CBDC6F1C4A0" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="8A65FBB3E1230EBBF36BF0F1F893C7A0" pageId="0" pageNumber="1">
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The new genus has a shell shape similar to members of the genus
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<taxonomicName id="AF1BCC261D4162348D0AA2967330D77A" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis id="549D4BCA39C515BA5634C94901276FE3" italics="true" pageId="0" pageNumber="1">Imeretiopsis</emphasis>
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</taxonomicName>
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gen. nov. from more eastern localities of the Imereti region, which have, in contrast, a sinuated labral lateral profile. However, both genera can be clearly distinguished by their penes (Fig.
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<figureCitation id="82CAAA1249DD2EB2CDA224785B14FB50" captionStart="Figure 11" captionStartId="F18" captionText="Figure 11. Morphology of the penis A, B Caucasopsis letsurtsume sp. nov., Samegrelo: A ventrally B dorsally C Imeretiopsis gorgoleti sp. nov. Photograph A. Falniowski and A. Rysiewska." figureDoi="10.3897/zookeys.955.51983.figure11" httpUri="https://binary.pensoft.net/fig/439171" pageId="0" pageNumber="1">11A, B</figureCitation>
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): the penis is long, with the filament (lacking in
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<taxonomicName id="946A4DB310D01CCA8CC5DCBBF6889D7C" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis id="9D9225AB3FF7103AA3A0216A1DD7801F" italics="true" pageId="0" pageNumber="1">Imeretiopsis</emphasis>
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</taxonomicName>
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) and, below the filament, delicately marked outgrowth on the left side (in
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<taxonomicName id="98EBC0019D2F4C9DED1EE7A9EA48609A" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis id="315674BA45DAC9039A117A3149F5ABD3" italics="true" pageId="0" pageNumber="1">Imeretiopsis</emphasis>
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</taxonomicName>
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there are two broad outgrowths).
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</paragraph>
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</subSubSection>
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<subSubSection id="F9BE0CC9A3C36723F61EAD3EC01C87A7" pageId="0" pageNumber="1" type="etymology">
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<paragraph id="754412B7ECDFA054612AB3247CC896A0" pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph id="2B1C4921090A4BC6900530AF4DB950C4" pageId="0" pageNumber="1">
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The name derived from the prefix
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<taxonomicName id="4A0C6A028AF99EA0EB54317A6841E8F0" genus="Caucas" lsidName="Caucas" pageId="0" pageNumber="1" rank="genus">
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<emphasis id="020A393CAACB6095EF367DE438CDE21C" italics="true" pageId="0" pageNumber="1">Caucas</emphasis>
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</taxonomicName>
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- referring to the distribution range in the Caucasus Mountains and suffix -
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<emphasis id="514038FF8CA178F0E30DF906B55CDAD2" italics="true" pageId="0" pageNumber="1">opsis</emphasis>
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reminiscent of the previously applied genus
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<emphasis id="8768BBDFE4292CD24B017DE2A21F69DD" italics="true" pageId="0" pageNumber="1">
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<taxonomicName id="D43A69135AA0E1A179BFEED3EBA9B9AB" authorityName="Pavlovic" authorityYear="1913" class="Gastropoda" family="Moitessieriidae" genus="Paladilhiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paladilhiopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Paladilhiopsis</taxonomicName>
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<normalizedToken id="EBD76A9842219578ED86E90DBCEB62D9" originalValue="Pavlović">Pavlovic</normalizedToken>
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|
</emphasis>
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, 1913, adopted by
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<bibRefCitation id="1135B4DAADF6AE61D57ED95E53D3AA19" author="Starobogatov, YI" journalOrPublisher="Byulleten Moskovskogo Obschchestva Ispytatelei Prirody, Otdel biologicheskiy" pageId="0" pageNumber="1" pagination="42 - 54" refId="B47" refString="Starobogatov, YI, 1962. Contribution to molluscs from subterranean waters of the Caucasus. Byulleten Moskovskogo Obschchestva Ispytatelei Prirody, Otdel biologicheskiy 67: 42 - 54" title="Contribution to molluscs from subterranean waters of the Caucasus." volume="67" year="1962">Starobogatov (1962)</bibRefCitation>
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for the similar shelled species from Abkhazia and from the Sochi region (Russia). Its gender is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="03FB95CD1A60185A9379E9E3F133003E" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="92D94B4CCE3D1A4B83F360BAFB4EE5B0" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="E74DE00DC50B1DA0EA789F53A53D03F8" pageId="0" pageNumber="1">
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The new genus
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<taxonomicName id="5EFC9424BC56FE3B02B2C24FCEEF719B" authorityName="Grego & Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis id="DAA60FE1F44B9F290F63A7DC49B39668" italics="true" pageId="0" pageNumber="1">Caucasopsis</emphasis>
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</taxonomicName>
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is known from the Samegrelo region, and likely from the Abkhazia and Sochi regions in the Russian Federation (Fig.
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<figureCitation id="269EB2AD34213034D96C11A5A9C9B260" captionStart="Figure 10" captionStartId="F15" captionText="Figure 10. Distribution map of Imeretiopsis gen. nov.: (magenta dots) and Caucasopsis gen. nov. (blue dots) 6 I. cameroni sp. nov. and I. iazoni sp. nov. 7 Imeretiopsis prometheus sp. nov. 8 I. nakeralaensis sp. nov. 9 I. gorgoleti sp. nov. 0 Caucasopsis orientalis (Starobogatov, 1962), C. subovata (Starobogatov, 1962) and C. pulcherrima (Starobogatov, 1962) 3 C. shadini (Starobogatov, 1962) 4 C. aculeus (Starobogatov, 1962) and C. schakuranica (Starobogatov, 1962) 22, 23 C. egrisi sp. nov. 24 C. olsavskyi sp. nov. 25, 30, 31 C. letsurtsume sp. nov." figureDoi="10.3897/zookeys.955.51983.figure10" httpUri="https://binary.pensoft.net/fig/439167" pageId="0" pageNumber="1">10</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |