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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1190.113898" ID-Pensoft-Pub="1313-2970-1190-163" ID-Pensoft-UUID="09087B133ACA53EAA174C3CB94A67735" ID-ZooBank="B41FADCFF7F0419583C5A203D0290B04" ModsDocID="1313-2970-1190-163" checkinTime="1706572078200" checkinUser="pensoft" docAuthor="Chaowvieng, Apisara, Sutcharit, Chirasak, Chanabun, Ratmanee, Srisonchai, Ruttapon, Jeratthitikul, Ekgachai &amp; Siriwut, Warut" docDate="2024" docId="FBC14CBB81435781A2EDCECAF9DDB1B2" docLanguage="en" docName="ZooKeys 1190: 163-193" docOrigin="ZooKeys 1190" docPubDate="2024-01-29" docSource="http://dx.doi.org/10.3897/zookeys.1190.113898" docTitle="Macrobrachium panhai Chaowvieng &amp; Siriwut 2024, sp. nov." docType="treatment" docUuid="42A0A555-C9CF-4DBF-B707-ED49A971F523" docUuidSource="ZooBank" docVersion="1" id="09087B133ACA53EAA174C3CB94A67735" lastPageNumber="163" masterDocId="09087B133ACA53EAA174C3CB94A67735" masterDocTitle="Molecular phylogeny and taxonomic position of Macrobrachium lanchesteri (De Man, 1911), with descriptions of two new species from Thailand (Decapoda, Caridea, Palaemonidae)" masterLastPageNumber="193" masterPageNumber="163" pageNumber="163" updateTime="1706572380357" updateUser="ExternalLinkService">
<mods:mods id="752AD9A018ED20215400167D491D7896" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A985CA99808EC5215430B40838265DFE">
<mods:title id="F0E37AFCDA16A26B3A8F2374DDB8C906">Molecular phylogeny and taxonomic position of Macrobrachium lanchesteri (De Man, 1911), with descriptions of two new species from Thailand (Decapoda, Caridea, Palaemonidae)</mods:title>
</mods:titleInfo>
<mods:name id="669EB93199EB42B435D331066A98E28D" type="personal">
<mods:role id="3A2E2ED90BC782E17E45EDCEF062ABAE">
<mods:roleTerm id="04732A6129FF35CDABE742DF12A07005">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D1A40C1106E5948C53CB3BFFE7D99625">Chaowvieng, Apisara</mods:namePart>
<mods:affiliation id="80D31635EC4868997226E19A3436E0D0">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
</mods:name>
<mods:name id="8D23F2411A4E59A48DECF009008B33BB" type="personal">
<mods:role id="F02AD71A3C1F8CF891FBB85051ECB6F9">
<mods:roleTerm id="901BDEDC3E0FD09E0F22FCF56F1BC458">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97A8C4C42E5DE0B9F98BAB9CB274624A">Sutcharit, Chirasak</mods:namePart>
<mods:nameIdentifier id="28C9F62536A3F1CAFB57EA70602C3020" type="ORCID">https://orcid.org/0000-0001-7670-9540</mods:nameIdentifier>
<mods:affiliation id="16CF9B7C89858ED0256F0EA759FC8FB5">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name id="B7680114555106701EDBB788D28F88D7" type="personal">
<mods:role id="658790D2698B169CF22677D9545F5AF9">
<mods:roleTerm id="04576112429C426F3F932D9188B9F20E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB9C36AF9C294225EA3B66312D2C7B8F">Chanabun, Ratmanee</mods:namePart>
<mods:nameIdentifier id="AA35BAA47FBE49674D51F42F8E94290F" type="ORCID">https://orcid.org/0000-0003-4584-2773</mods:nameIdentifier>
<mods:affiliation id="36AB8DD502CE7466A61A2840B27EC5DD">Program in Animal Science, Faculty of Agricultural Technology, Sakon Nakhon Rajabhat University, Sakon Nakhon 47000, Thailand</mods:affiliation>
</mods:name>
<mods:name id="78E7D7F88C44885FA668F78110A22594" type="personal">
<mods:role id="1390E8B2871B5899129DB5C25FA699A7">
<mods:roleTerm id="F6A3844EDCB68CF3E578F17081718C38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10417FAB6EF83B86B34E6CE60CB016CC">Srisonchai, Ruttapon</mods:namePart>
<mods:nameIdentifier id="63DC61510F52C306ED5F27244A6AEBB5" type="ORCID">https://orcid.org/0000-0002-7142-0999</mods:nameIdentifier>
<mods:affiliation id="2B19D7BB1D3C97D6355CE42FBFB5313D">Biodiversity and Utilization Research Unit, Center of Excellence in Modern Agriculture, Sakon Nakhon Rajabhat University, Sakon Nakhon 47000, Thailand</mods:affiliation>
</mods:name>
<mods:name id="E732F74A0BE141FA286EAB9C84EEFDE2" type="personal">
<mods:role id="B49E5C7B8BE99E6BE83F4204BD05FD42">
<mods:roleTerm id="C64C813EFB9F92652FCBB86C78C66576">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A8BACE970CFAC3BCF118CAD83323CCFE">Jeratthitikul, Ekgachai</mods:namePart>
<mods:affiliation id="D0AC35EA462D95525003ED97DCEFA18C">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
</mods:name>
<mods:name id="046167941ECA9B344DC196FE1F97F835" type="personal">
<mods:role id="616F0EAA00A8F6019FD8899C17B22FF1">
<mods:roleTerm id="1CB7D41061098996FC91A1234991675D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9D0E4D3D7BC2CF2D9E3EEA95FB111731">Siriwut, Warut</mods:namePart>
<mods:nameIdentifier id="3C9E6553F74A1763DDD29E626F643AE7" type="ORCID">https://orcid.org/0000-0001-6760-9724</mods:nameIdentifier>
<mods:affiliation id="D004416FDE016CD68C90E98BBA8E5169">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
<mods:nameIdentifier id="0DC01B4EE61FEC4A23F0510503D127DD" type="email">warut.sir@mahidol.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="13750FADF701538EF9EFE0DF173FD0DF">text</mods:typeOfResource>
<mods:relatedItem id="920A680137D2BC90675B46E8182962B4" type="host">
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<mods:title id="B29BCD37F5AAB18CB315347D5041316B">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="5D02E8A50479C3BB983345921D4C9944">
<mods:date id="A926B5B842C616181F015D79FD3C7DEA">2024</mods:date>
<mods:detail id="D5A93ECDB6587420E7C698671621658B" type="pubDate">
<mods:number id="C8CB2AD1258E5010CCDBBC1CBC2601A9">2024-01-29</mods:number>
</mods:detail>
<mods:detail id="46CB29341FE6C0AD32FFF9646F93CB45" type="volume">
<mods:number id="6B644746CCB76FD15CECD82C3566E396">1190</mods:number>
</mods:detail>
<mods:extent id="9C9CFAE95B4BED64F113B6315F70F621" unit="page">
<mods:start id="084088A75CDCDEBB2F697E84DA02E144">163</mods:start>
<mods:end id="C0C04DA82629CE834CAC0667FE8ABA8D">193</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="B958600B93A6F5BC0792CA4EBBBA5C59">
<mods:url id="349DEAA4306E3265D3BD9BF89D150AB0">http://dx.doi.org/10.3897/zookeys.1190.113898</mods:url>
</mods:location>
<mods:classification id="B3F8A329CD8E84D7E57730E7DCF3E545">journal article</mods:classification>
<mods:identifier id="1998BC380C41D8F6008266C53561CD1D" type="DOI">http://dx.doi.org/10.3897/zookeys.1190.113898</mods:identifier>
<mods:identifier id="79DABEDE3B3494B82B7F815D95751C39" type="Pensoft-Pub">1313-2970-1190-163</mods:identifier>
<mods:identifier id="0C4CC9762BFD61A250C39C4D39E50E4E" type="ZooBank">B41FADCFF7F0419583C5A203D0290B04</mods:identifier>
<mods:identifier id="C7B3CC0D5341EB4364980892E9080164" type="Pensoft-UUID">09087B133ACA53EAA174C3CB94A67735</mods:identifier>
</mods:mods>
<treatment id="FBC14CBB81435781A2EDCECAF9DDB1B2" LSID="urn:lsid:zoobank.org:act:42A0A555-C9CF-4DBF-B707-ED49A971F523" httpUri="http://treatment.plazi.org/id/FBC14CBB81435781A2EDCECAF9DDB1B2" lastPageNumber="163" pageId="0" pageNumber="163">
<subSubSection id="3D2D5D2AF0FF339C4125013044220F01" pageId="0" pageNumber="163" type="nomenclature">
<paragraph id="2D377473C6CF0FED2D8D18C6EFD9D319" pageId="0" pageNumber="163">
<taxonomicName id="F94832B2565C7F396D59C13751B15D2E" LSID="https://zoobank.org/42A0A555-C9CF-4DBF-B707-ED49A971F523" authority="Chaowvieng &amp; Siriwut" authorityName="Chaowvieng &amp; Siriwut" authorityYear="2024" class="Insecta" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrobrachium panhai" order="Decapoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="panhai" status="sp. nov.">Macrobrachium panhai Chaowvieng &amp; Siriwut</taxonomicName>
<taxonomicNameLabel id="C012E7995874EDCD02DF9E4CC6A9AE50" pageId="0" pageNumber="163">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="E2837999E7F7BA43634DB8C282A216F5" pageId="0" pageNumber="163" type="description">
<paragraph id="7204D885013E4675040678374B4BA4D9" pageId="0" pageNumber="163">
<figureCitation id="C2DC90A3284F46D807DC4CC1BC3DC86C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Living habit of specimens of three Macrobrachium species A, B M. lanchesteri from Bang Khiat, Singhanakhon, Songkhla, Thailand C, D M. rostrolevatus sp. nov. from Bueng Khong Long, Bueng Kan, Thailand E M. panhai sp. nov. from Sri Nakarin Dam, Tha Kradan, Si Sawat, Kanchanaburi, Thailand. Scale bars: 1 cm." figureDoi="10.3897/zookeys.1190.113898.figure3" httpUri="https://binary.pensoft.net/fig/974855" pageId="0" pageNumber="163">Figs 3E</figureCitation>
<figureCitation id="28D5266AE718C64F7E55B89E70542CBB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="62709116F142939EEA12AB17A0EBFB31" pageId="0" pageNumber="163" type="materials_examined">
<paragraph id="D89296EED7B9247B1B3BDC9648DB956D" pageId="0" pageNumber="163">Material examined.</paragraph>
<paragraph id="D5B967D01BA9E9CAE979D3B2FB639483" pageId="0" pageNumber="163">
<emphasis id="C575823BB57E61C491D90D5710BDEDC3" bold="true" pageId="0" pageNumber="163">
<emphasis id="979FB8E487CF80DD3FFEC44E8323CBC4" italics="true" pageId="0" pageNumber="163">
<typeStatus id="BDD0F7A3900F1816C0FFA56E8E215D2E" type="holotype">Holotype</typeStatus>
</emphasis>
: Kanchanaburi
</emphasis>
• Ovigerous
<specimenCount id="76E823F279103A7D63FD3B8A0807A79E" count="1" type="female">♀</specimenCount>
from Sri Nakarin Dam, Tha Kradan, Si Sawat;
<geoCoordinate id="EB1F0BB8699103B7CD48EB69FE9CA7C3" degrees="14" direction="north" minutes="24" orientation="latitude" precision="1" seconds="12.1" value="14.40336">14°24'12.1&quot;N</geoCoordinate>
,
<geoCoordinate id="D7615E060D6A8EB3242500B1E791974B" degrees="99" direction="east" minutes="07" orientation="longitude" precision="1" seconds="24.7" value="99.12353">99°07'24.7&quot;E</geoCoordinate>
; CUMZ MP00302.
<emphasis id="332C526D8286CEC7260CBD87521924A6" bold="true" italics="true" pageId="0" pageNumber="163">
<typeStatus id="6084FA671DD7D852C3E81B238FA9EF49" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="D7623FE4BE4C8D568546286638FA7F50" count="7" type="female">7 ♀♀</specimenCount>
, 8 ovigerous,
<materialsCitation id="A9EB4EFCF9CBD72158B09FBC5D511AEA" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" specimenCount="6" specimenCount-male="6" typeStatus="holotype">
<specimenCount id="27708A861BF2E1B751E3B0CCAD0BC177" count="6" type="male">6 ♂♂</specimenCount>
from the same locality of
<typeStatus id="5DA2BCD5D502A21C7E9F7F22DF011B2C" type="holotype">holotype</typeStatus>
; MUMNH MP00303
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="96AA461042572A444A59CE4D0529610A" pageId="0" pageNumber="163" type="materials_examined">
<paragraph id="5B96BDE5255AD8890DD1921C3EC307AF" pageId="0" pageNumber="163">Additional material.</paragraph>
<paragraph id="B6CEAC3A5E61101E15F370554C9BEEF9" pageId="0" pageNumber="163">
<emphasis id="CF00A880D38F47712CB25E0A1B76A9AB" bold="true" pageId="0" pageNumber="163">Chiang Mai</emphasis>
•
<specimenCount id="A5A35FC6C41C59D97089F1AA728EB03E" count="6" type="female">6 ♀♀</specimenCount>
, 6 ovigerous, Nong Han, San Sai;
<geoCoordinate id="A5E0F28DDB1B141042A2ED97EB9D3928" degrees="18" direction="north" minutes="53" orientation="latitude" precision="1" seconds="44.9" value="18.895805">18°53'44.9&quot;N</geoCoordinate>
,
<geoCoordinate id="865D67C8BD930D597DC22C409B2C3D54" degrees="99" direction="east" minutes="01" orientation="longitude" precision="1" seconds="05.7" value="99.01826">99°01'05.7&quot;E</geoCoordinate>
; MUMNH MP00304. •
<specimenCount id="5513D07DC85799E144CA5A7318AF26D5" count="1" type="female">1 ♀</specimenCount>
, Ping, Ki Lek, Mae Taeng;
<geoCoordinate id="E9B8EF64B16040DEED763EB8E2AD7D6D" degrees="19" direction="north" minutes="04" orientation="latitude" precision="1" seconds="37.0" value="19.076946">19°04'37.0&quot;N</geoCoordinate>
,
<geoCoordinate id="6C954CF31828E332D35527D53C0AA4E4" degrees="98" direction="east" minutes="56" orientation="longitude" precision="1" seconds="59.8" value="98.94994">98°56'59.8&quot;E</geoCoordinate>
; MUMNH MP00305.
<emphasis id="AAFC72DA4F12674F2158EE137464B4A5" bold="true" pageId="0" pageNumber="163">Nan</emphasis>
•
<materialsCitation id="DD4A8DB83F2E8B671D1CB19A70C1E46F" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" latitude="18.753555" location="Ban Na Bua" longLatPrecision="1" longitude="101.011024" specimenCount="1" specimenCount-female="1">
<specimenCount id="BD2D3ACE1CCDC9049FBFACD50BA0FEA9" count="1" type="female">1 ♀</specimenCount>
,
<location id="0085D99221AFA70C9A29793C8EB81945" LSID="urn:lsid:plazi:treatment:FBC14CBB81435781A2EDCECAF9DDB1B2:0085D99221AFA70C9A29793C8EB81945" latitude="18.753555" longLatPrecision="1" longitude="101.011024" name="Ban Na Bua">Ban Na Bua</location>
, Nong Daeng, Mae Charim;
<geoCoordinate id="0DF5FEB5D0BA71F41CA419C6D6E92AA7" degrees="18" direction="north" minutes="45" orientation="latitude" precision="1" seconds="12.8" value="18.753555">18°45'12.8&quot;N</geoCoordinate>
,
<geoCoordinate id="547BC261C2A4979EDB709EA814D3CFC6" degrees="101" direction="east" minutes="00" orientation="longitude" precision="1" seconds="39.7" value="101.011024">101°00'39.7&quot;E</geoCoordinate>
; MUMNH MP00306.
<emphasis id="1920AECC9D7AEAF81B55E4371C6D0FEE" bold="true" pageId="0" pageNumber="163">Uttaradit</emphasis>
</materialsCitation>
•
<specimenCount id="1D14C0BF471302644BF3083D430925A6" count="1" type="male">1 ♂</specimenCount>
, Klong Tron, Ban Bueng Pra Kot, Thong Sang Khan;
<geoCoordinate id="E94556D7AAA0C9867AD8E2D478A00BC9" degrees="17" direction="north" minutes="35" orientation="latitude" precision="1" seconds="39.1" value="17.594194">17°35'39.1&quot;N</geoCoordinate>
,
<geoCoordinate id="5B94B3209B529A6B22D053CCF2D76CAE" degrees="100" direction="east" minutes="29" orientation="longitude" precision="1" seconds="10.3" value="100.48619">100°29'10.3&quot;E</geoCoordinate>
; MUMNH MP00307. •
<specimenCount id="CE19A8D659554C0E39EC0DC34C069405" count="3" type="female">3 ♀♀</specimenCount>
, Klong Tron, Nam Khai, Nam Pat;
<geoCoordinate id="34B4EA34D418ED1937F579F346D4F5D4" degrees="17" direction="north" minutes="36" orientation="latitude" precision="1" seconds="15.3" value="17.60425">17°36'15.3&quot;N</geoCoordinate>
,
<geoCoordinate id="3FD29A5A45F44C122444CE741EE0F645" degrees="100" direction="east" minutes="32" orientation="longitude" precision="1" seconds="15.3" value="100.53758">100°32'15.3&quot;E</geoCoordinate>
; MUMNH MP00308.
<emphasis id="6D03889D4413B7E5688BDCDCC65A146F" bold="true" pageId="0" pageNumber="163">Buengkan</emphasis>
•
<specimenCount id="2F1948190A347B89D498C69DCD62FD2F" count="1" type="female">1 ♀</specimenCount>
, Klong Hi, Sang, Seka;
<geoCoordinate id="EDB116295CCDAC469DBE520D479FA415" degrees="17" direction="north" minutes="54" orientation="latitude" precision="1" seconds="17.9" value="17.904972">17°54'17.9&quot;N</geoCoordinate>
,
<geoCoordinate id="087C6C72698ACA40705BD49F756A07DD" degrees="103" direction="east" minutes="57" orientation="longitude" precision="1" seconds="12.0" value="103.95333">103°57'12.0&quot;E</geoCoordinate>
; MUMNH MP00351.
<emphasis id="08FF10E07BA3F6889E98472E62590FD8" bold="true" pageId="0" pageNumber="163">Nong Khai</emphasis>
•
<specimenCount id="94B90D9724FF0A06D7DECD86F47F0E80" count="2" type="female">2 ♀♀</specimenCount>
, Nam Suai, Song Hong, Mueang Nong Khai;
<geoCoordinate id="801413A257CB225568C930B1E883687D" degrees="17" direction="north" minutes="45" orientation="latitude" precision="1" seconds="01.1" value="17.750305">17°45'01.1&quot;N</geoCoordinate>
,
<geoCoordinate id="F39C445B303E3DBE5DDB39493F4274AD" degrees="102" direction="east" minutes="51" orientation="longitude" precision="1" seconds="00.5" value="102.850136">102°51'00.5&quot;E</geoCoordinate>
; MUMNH MP00309.
<emphasis id="BDA3B49075925C2C67BE0CCDC66A7EA3" bold="true" pageId="0" pageNumber="163">Ubon Ratchathani</emphasis>
•
<specimenCount id="1D286D379D6E93F981284B9D84049B6C" count="1" type="female">1 ♀</specimenCount>
, 1 ovigerous,
<materialsCitation id="40737B9B35C175FE24630ED23A1C2005" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" latitude="15.319528" location="Mekong River" longLatPrecision="1" longitude="105.49811" specimenCount="2" specimenCount-male="2">
<specimenCount id="AA5996E42FFCA14B5A003C557FB5CBAC" count="2" type="male">2 ♂♂</specimenCount>
,
<location id="FD974DC2ADE190FA742620F9BF969913" LSID="urn:lsid:plazi:treatment:FBC14CBB81435781A2EDCECAF9DDB1B2:FD974DC2ADE190FA742620F9BF969913" latitude="15.319528" longLatPrecision="1" longitude="105.49811" name="Mekong River">Mekong River</location>
, Khong Chiam;
<geoCoordinate id="C12CFE6446D9AFA97EB2BBD6C796DD0C" degrees="15" direction="north" minutes="19" orientation="latitude" precision="1" seconds="10.3" value="15.319528">15°19'10.3&quot;N</geoCoordinate>
,
<geoCoordinate id="D0E44A1ADA72A6A7FBF3C2FFAB87EEA9" degrees="105" direction="east" minutes="29" orientation="longitude" precision="1" seconds="53.2" value="105.49811">105°29'53.2&quot;E</geoCoordinate>
; MUMNH MP00310
</materialsCitation>
. •
<specimenCount id="F1E7BE5F9E801C6B0CFC7761BB127D79" count="1" type="female">1 ♀</specimenCount>
, 1 ovigerous, Sae Hua Maew Waterfall, Nong Saeng Yai, Khong Chiam;
<geoCoordinate id="04CA880E597E6803DA973A906695034B" degrees="15" direction="north" minutes="20" orientation="latitude" precision="1" seconds="12.2" value="15.336722">15°20'12.2&quot;N</geoCoordinate>
,
<geoCoordinate id="3327878175F863F03F362FEE0C0F49E2" degrees="105" direction="east" minutes="24" orientation="longitude" precision="1" seconds="02.6" value="105.40073">105°24'02.6&quot;E</geoCoordinate>
; MUMNH MP00311.
<emphasis id="BE92EBA7451D5D8D2559F988FAA54194" bold="true" pageId="0" pageNumber="163">Phitsanulok</emphasis>
•
<specimenCount id="861E373446B28FE1ADA732D4E2A99313" count="6" type="female">6 ♀♀</specimenCount>
,
<specimenCount id="F7D1CFB606B2DB419C230AC8F236D001" count="1" type="male">1 ♂</specimenCount>
, Ban Bo, Wang Nok Aen, Wang Thong;
<geoCoordinate id="D55AABCFB0A8D292C8CE2DB608DB7EE3" degrees="16" direction="north" minutes="51" orientation="latitude" precision="1" seconds="13.6" value="16.853779">16°51'13.6&quot;N</geoCoordinate>
,
<geoCoordinate id="214AA01E0A66F34BA25E9C6E352FA98B" degrees="100" direction="east" minutes="36" orientation="longitude" precision="1" seconds="43.2" value="100.612">100°36'43.2&quot;E</geoCoordinate>
; MUMNH MP00312.
<emphasis id="F78F385A90C7F30DAFEAA53EB1443F4F" bold="true" pageId="0" pageNumber="163">Phetchabun</emphasis>
•
<specimenCount id="B4EDA4E921B6918FB59F1F4F6BF33FD4" count="1" type="female">1 ♀</specimenCount>
, Ang Kep Nam Nam Khuen Nam Lang, Wang Hin, Wang Pong;
<geoCoordinate id="17F6F7CA17F418E1CAEDA5843D3ED83D" degrees="16" direction="north" minutes="25" orientation="latitude" precision="1" seconds="19.8" value="16.422167">16°25'19.8&quot;N</geoCoordinate>
,
<geoCoordinate id="2FC2A161CB96A656B3C8D0EFA1E82380" degrees="100" direction="east" minutes="48" orientation="longitude" precision="1" seconds="48.2" value="100.81339">100°48'48.2&quot;E</geoCoordinate>
; MUMNH MP00313.
<emphasis id="B04997F832066B8B481374DCB8DD8C37" bold="true" pageId="0" pageNumber="163">Nakhon Nayok</emphasis>
• 2 ovigerous, Ban Na, Pa Kha, Ban Na;
<geoCoordinate id="A5874138E59A0190BD097929D4CDF442" degrees="14" direction="north" minutes="17" orientation="latitude" precision="1" seconds="11.2" value="14.286445">14°17'11.2&quot;N</geoCoordinate>
,
<geoCoordinate id="2B8C5FE923444A2348C03C6C3D79A773" degrees="101" direction="east" minutes="04" orientation="longitude" precision="1" seconds="13.7" value="101.07047">101°04'13.7&quot;E</geoCoordinate>
; MUMNH MP00314.
<emphasis id="49B3B5F208C2CCAEDA08EA4EFC3D68B3" bold="true" pageId="0" pageNumber="163">Suphan Buri</emphasis>
•
<specimenCount id="140720C61D849C76D2F384800FBDDD53" count="10" type="female">10 ♀♀</specimenCount>
, 8 ovigerous,
<specimenCount id="114DF3485F6010E0AEF713E242F0565F" count="4" type="male">4 ♂♂</specimenCount>
, Ban Chara Mai, Ban Kong, U Thong;
<geoCoordinate id="A1DED833F20453E61543C55047712E22" degrees="14" direction="north" minutes="34" orientation="latitude" precision="1" seconds="51.0" value="14.580833">14°34'51.0&quot;N</geoCoordinate>
,
<geoCoordinate id="BFB31718A2D0FE154F89A047F356369B" degrees="99" direction="east" minutes="52" orientation="longitude" precision="1" seconds="04.7" value="99.86797">99°52'04.7&quot;E</geoCoordinate>
; MUMNH MP00315.
<emphasis id="3E3C3BFBE3A8996745D64DB90210F2E8" bold="true" pageId="0" pageNumber="163">Sa Kaeo</emphasis>
• 1 ovigerous, Huai Yang, Ta Phraya;
<geoCoordinate id="19DB903DD8038137568F5693C777C03D" degrees="14" direction="north" minutes="00" orientation="latitude" precision="1" seconds="46.5" value="14.012917">14°00'46.5&quot;N</geoCoordinate>
,
<geoCoordinate id="BE543FCD2E2BB70B792BB29E667CB907" degrees="102" direction="east" minutes="38" orientation="longitude" precision="1" seconds="37.0" value="102.64361">102°38'37.0&quot;E</geoCoordinate>
; MUMNH MP00316.
<emphasis id="082AEA628F546AEF65EA4D4FC6E2E0BF" bold="true" pageId="0" pageNumber="163">Chachoengsao</emphasis>
•
<specimenCount id="9D919E27A5785AE12E59A98AD41555FC" count="2" type="female">2 ♀♀</specimenCount>
, 3 ovigerous, Nong Prue Mai Kaeo, Wang Yen, Plaeng Yao;
<geoCoordinate id="B1953279EEA4B11D0FCCE4EF84FA6913" degrees="13" direction="north" minutes="33" orientation="latitude" precision="1" seconds="31.5" value="13.55875">13°33'31.5&quot;N</geoCoordinate>
,
<geoCoordinate id="F627D355F0DF2BE634093C117131FD78" degrees="101" direction="east" minutes="17" orientation="longitude" precision="1" seconds="44.0" value="101.295555">101°17'44.0&quot;E</geoCoordinate>
; MUMNH MP00317.
<emphasis id="95FEC0214D2C0298FFD5741754FDEBE8" bold="true" pageId="0" pageNumber="163">Rayong</emphasis>
• 12 ovigerous, Khao Chuk, Kong Din, Klaeng;
<geoCoordinate id="6B872E96ADB68D81000F4522E2327950" degrees="12" direction="north" minutes="51" orientation="latitude" precision="1" seconds="32.0" value="12.85889">12°51'32.0&quot;N</geoCoordinate>
,
<geoCoordinate id="BD8D6F93EE3F285EB03375FB4E29B0D8" degrees="101" direction="east" minutes="46" orientation="longitude" precision="1" seconds="12.0" value="101.770004">101°46'12.0&quot;E</geoCoordinate>
; MUMNH MP00318. •
<specimenCount id="04599A26581207EAAB1AFAA147D2B338" count="7" type="female">7 ♀♀</specimenCount>
, 11 ovigerous,
<specimenCount id="DA32346C13FF7FCB55EF7E3A66AD40FA" count="12" type="male">12 ♂♂</specimenCount>
, Koh Phasuk, Makham, Makham, Chanthaburi; MUMNH MP00319.
<emphasis id="D5D287A4E2CEB170B85E1CA15B339D03" bold="true" pageId="0" pageNumber="163">Trat</emphasis>
•
<specimenCount id="18AB0146BE10EDF0F82554A65593BFD9" count="1" type="female">1 ♀</specimenCount>
, 2 ovigerous, Huai Raeng, Dan Chumphon, Bo Rai;
<geoCoordinate id="D1B99F6F6DE46B813946F50199A5B7EA" degrees="12" direction="north" minutes="23" orientation="latitude" precision="1" seconds="48.4" value="12.396778">12°23'48.4&quot;N</geoCoordinate>
,
<geoCoordinate id="9C22115C3DBF832FE683AE288C015A43" degrees="102" direction="east" minutes="39" orientation="longitude" precision="1" seconds="15.1" value="102.6542">102°39'15.1&quot;E</geoCoordinate>
; MUMNH MP00320. •
<specimenCount id="89B6293DE59175DC60D8B4A31D39A08C" count="1" type="female">1 ♀</specimenCount>
, 2 ovigerous, Ang Kep Nam Dan Chumphon, Dan Chumphon, Bo Rai;
<geoCoordinate id="09FBF861B582AE215A0EA7BF1744C955" degrees="12" direction="north" minutes="27" orientation="latitude" precision="1" seconds="45.4" value="12.462611">12°27'45.4&quot;N</geoCoordinate>
,
<geoCoordinate id="BBD35A0344693495DB7B74F22C34A8A0" degrees="102" direction="east" minutes="38" orientation="longitude" precision="1" seconds="24.6" value="102.64017">102°38'24.6&quot;E</geoCoordinate>
; MUMNH MP00321.
<emphasis id="0A746A35C10CA562069BCA2B2F96F4CC" bold="true" pageId="0" pageNumber="163">Phetburi</emphasis>
•
<specimenCount id="6852E4713A0E8ED89086292226C8DAA2" count="1" type="male">1 ♂</specimenCount>
, Klong Prachan, Yang Nam Klat Tai, Nong Ya Plong;
<geoCoordinate id="9E19F05082F37FD637D2B72F855F06C4" degrees="13" direction="north" minutes="06" orientation="latitude" precision="1" seconds="33.27" value="13.109242">13°06'33.27&quot;N</geoCoordinate>
,
<geoCoordinate id="0A77951A7C774A1F96EBDADA8E5D9ED1" degrees="99" direction="east" minutes="43" orientation="longitude" precision="1" seconds="22.75" value="99.722984">99°43'22.75&quot;E</geoCoordinate>
; MUMNH MP00322.
</paragraph>
</subSubSection>
<subSubSection id="C1B0992F086D1F737781870887AEA574" pageId="0" pageNumber="163" type="diagnosis">
<paragraph id="8020205D44A4176E41D8BF8F59F3AE16" pageId="0" pageNumber="163">Diagnosis.</paragraph>
<paragraph id="2784CF9B49719A0A88F861F934E5B7AD" pageId="0" pageNumber="163">
Rostrum straight proximally and slightly upward distally. Rostrum length reaching beyond end of antennular peduncle and exceeding the scaphocerite. Rostral formula: 8-12/3-6 teeth including two or three distal teeth with small gap separate from rest. Carapace smooth. Epistome bilobed. First pereiopods reaching end of scaphocerite. Second pereiopods thin and long, similar in form and equals in length, exceeding scaphocerite. Fingers covered with scattered setae, slightly shorter than palm. Translucent razor edge present anteriorly between fingers and no teeth on inner side of cutting edges. Carpus cylindrical shape and articulation margin expanded. Carpus 1.5
<normalizedToken id="FB48A7971B50993975385F0B4E05BCB7" originalValue="×">x</normalizedToken>
longer than chela. Merus subcylindrical. Carpus 1.5
<normalizedToken id="EFAF04A59510920B954E1086B7203709" originalValue="×">x</normalizedToken>
longer than merus. Third pereiopods thin and long, reaching end of scaphocerite. Dactylus curved distally with short setae. Propodus 2
<normalizedToken id="61A6687357F3710C146F21449963FC5D" originalValue="×">x</normalizedToken>
longer than dactylus. Propodus with three or four pairs of spines and fine setae present scarcely on articulation margin. Propodus 2
<normalizedToken id="C30CD7D81854BBFCCBA904CE59E7D465" originalValue="×">x</normalizedToken>
longer than carpus. Sixth to eighth thoracic sternites smooth. First and second pleonal sternites with small median process or smooth. Third and fourth pleonal sternites smooth. Fifth pleonal sternite with triangular ridge. Uropodal diaeresis with inner movable spine slightly longer than outer angle.
</paragraph>
</subSubSection>
<subSubSection id="D8B0A627DB30FE429D27A073A288CDBD" pageId="0" pageNumber="163" type="description">
<paragraph id="B58B8B75D9E4AE14346C2A965419B45B" pageId="0" pageNumber="163">Composite description</paragraph>
<paragraph id="CB9D1B785D09E722733FBB303EDCC972" pageId="0" pageNumber="163">
<emphasis id="FEB54262707EE3E095D24B96924F4280" bold="true" pageId="0" pageNumber="163">
(holotype in parentheses).
<emphasis id="2AB578D86FBC8E71AE038431D7259F8C" italics="true" pageId="0" pageNumber="163">Rostrum</emphasis>
</emphasis>
(Fig.
<figureCitation id="DFA6C8E9A2DC278C9A4E8D2050ABC93E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">5B</figureCitation>
). Straight or proximal convex and slightly distal upward. Rostrum length exceeding end of antennular peduncle and slightly exceeding scaphocerite (rl 7.32 mm). Dorsal margin with 8-12 (10) teeth including two or three (3) teeth distally with small gap from rest. Postorbital margin with one or two (1) teeth, reaching one-third of carapace length. First dorsal tooth positioned slightly behind hepatic spine. Ventral margin with 3-6 (4) teeth, starting from middle to distal margin. Short setae present between rostral teeth.
</paragraph>
<caption id="C19933DB23A864B9E59B4082E6BEDA6F" doi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163" start="Figure 5" startId="F5">
<paragraph id="8F89C352EF3B7D00D24D15308C82556A" pageId="0" pageNumber="163">
<emphasis id="E2F2B2D1A67532529222338DAAE30F39" bold="true" pageId="0" pageNumber="163">Figure 5.</emphasis>
Morphological characteristics of
<taxonomicName id="0C99FA3C998DA9BA831FDF9AE2D3DBC6" authorityName="Chaowvieng &amp; Siriwut" authorityYear="2024" class="Insecta" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrobrachium panhai" order="Decapoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="98C528F011DEC6A0273D1A6B6445B2D2" italics="true" pageId="0" pageNumber="163">Macrobrachium panhai</emphasis>
</taxonomicName>
sp. nov. (
<emphasis id="4D83F319EBCE0A0F9BE94FC43B8042B1" bold="true" pageId="0" pageNumber="163">A, F</emphasis>
ovigerous female paratype MUMNH MP00303
<emphasis id="38A047DDE2C7F96B0D778B4FB296D908" bold="true" pageId="0" pageNumber="163">B-E, G-H</emphasis>
ovigerous female holotype CUMZ MP00302)
<emphasis id="F136B69500A64110B83C5F8F68733DC3" bold="true" pageId="0" pageNumber="163">A</emphasis>
lateral view
<emphasis id="D45D5F7E863CDCCB5DB1DD70ED24041D" bold="true" pageId="0" pageNumber="163">B</emphasis>
carapace
<emphasis id="43C85CAD7BAF9628662D482AB2896E95" bold="true" pageId="0" pageNumber="163">C</emphasis>
epistome
<emphasis id="003FBDFE20266E6DC78E0F79418A0AE9" bold="true" pageId="0" pageNumber="163">D</emphasis>
second pereiopod
<emphasis id="3159CB093CAE58D5EBACAFF98CD16C2A" bold="true" pageId="0" pageNumber="163">E</emphasis>
teeth between fingers
<emphasis id="D57A90377B86BA0ECE888562146E6FA4" bold="true" pageId="0" pageNumber="163">F</emphasis>
third pereiopod
<emphasis id="013793340B62EB67BC9DA578D80C9691" bold="true" pageId="0" pageNumber="163">G</emphasis>
uropod and
<emphasis id="49970A538462B13511E80DED678883D1" bold="true" pageId="0" pageNumber="163">H</emphasis>
movable spine at uropodal diaeresis. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="F0E719A53AC22D5C1A60CA96BC269A24" pageId="0" pageNumber="163">
<emphasis id="F21067D3E97C580EBD143C9AE6AA0C9D" bold="true" italics="true" pageId="0" pageNumber="163">Cephalon</emphasis>
(Fig.
<figureCitation id="37FE3546D7AB352CE420749B84A70353" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">5B</figureCitation>
). Eye well developed. Ocular beak without laterally expanded tip. Cornea longer and broader than stalk. Postantennular carapace margin rounded. Cornea osculum longer than stalk. Antennular peduncle longer than wide, with fine setae. Basal segment short, second segment shorter than third segment. Stylocerite projection sharp, reaching beyond basal segment. Antennal spine sharp, situated below orbital margin. Hepatic spine slightly larger than antennal spine, positioned posteriorly and lower than antennal spine. Scaphocerite with straight margin, distolateral tooth sharp and not reaching end of lamella. Epistome bilobed (Fig.
<figureCitation id="9864D387542DC0B641E97329BFEC6AE7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">5C</figureCitation>
). Branchiostegal suture beginning at carapace margin to behind hepatic spine. Carapace surface smooth (cl 5.76 mm).
</paragraph>
<paragraph id="04C0B8B47FDB611065A0C9AD69289EEC" pageId="0" pageNumber="163">
<emphasis id="F47910BC5FB45AC1507BBA7EA5D01739" bold="true" italics="true" pageId="0" pageNumber="163">First pereiopods.</emphasis>
Thin and long, reaching end of scaphocerite. Fingers as long as palm, tips with fine setae. Series of setae present on anterior inner part of palm. Carpus slightly longer than merus. Distal articulation of carpus with series of fine setae. Ischium shorter than merus. Scattered setae present on all segments.
</paragraph>
<paragraph id="788D2D44DE63C57DBBE5DA1D00AA0573" pageId="0" pageNumber="163">
<emphasis id="3CC8CA7C4CC1C4F6DEAC13C4BD89B5B8" bold="true" italics="true" pageId="0" pageNumber="163">Second pereiopods</emphasis>
(Fig.
<figureCitation id="207541F3469755347618E25921986C1C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">5D</figureCitation>
). Thin and long, similar in form and exceeding scaphocerite. Fingers subcylindrical covered with scattered setae. Palm 1.1-1.5
<normalizedToken id="ABDE9E1B0C2E307CF7302F2651F9A318" originalValue="×">x</normalizedToken>
longer than fingers (Fin 1.06: Pal 1.39 mm). Fingers with translucent razor edges present anteriorly and cutting edge between fingers smooth. Tip of fingers crossed and covered by fine setae (Fig.
<figureCitation id="544986F7333D6EF5350BFB2A90598C4E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">5E</figureCitation>
). Carpus cylindrical shape and articulation margin expanded. Carpus 1.2-2
<normalizedToken id="9CAB7261E1996314960AC23A64DBF08D" originalValue="×">x</normalizedToken>
longer than chela (Che 2.45: Car 4.22 mm). Merus subcylindrical. Carpus 1.2-1.7
<normalizedToken id="8316392B0358093620D795A1B19684E5" originalValue="×">x</normalizedToken>
longer than merus (Mer 2.85: Car 4.22 mm). Ischium as long as merus. Scattered short setae present on all segments.
</paragraph>
<paragraph id="3437B961CB28B296203EEC8E5660A062" pageId="0" pageNumber="163">
<emphasis id="2303808545201549619BA62120309BD7" bold="true" italics="true" pageId="0" pageNumber="163">Third pereiopods</emphasis>
(Fig.
<figureCitation id="CB9D6149023E5622954099010DC5FA38" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">5F</figureCitation>
). Thin and slender, reaching end of scaphocerite. Dactylus short and curved distally. Propodus 2
<normalizedToken id="CAA865340DEB2BA4931C4696A9649130" originalValue="×">x</normalizedToken>
longer than dactylus. Propodus with three or four pairs of spines along inferior-lateral margin and fine setae at distal articulation, 2
<normalizedToken id="18445A2F65F60FD11A83114F85AA439C" originalValue="×">x</normalizedToken>
longer than carpus. Ischium shorter than carpus. Scattered short setae present on all segments.
</paragraph>
<paragraph id="071C426C5A6F46181C39172224D3687E" pageId="0" pageNumber="163">
<emphasis id="90BB9B6573D9BDAFC1716F61887D457D" bold="true" italics="true" pageId="0" pageNumber="163">Fourth and fifth pereiopods.</emphasis>
Long and slender, exceeding scaphocerite. Propodus of fourth pereiopods with 3-6 (4) pairs of spines distributed along its length, 2.5
<normalizedToken id="6B1B3B3376EB200DE0077022D2917252" originalValue="×">x</normalizedToken>
longer than dactylus. Propodus as long as merus. Ischium shorter than merus. Propodus with fine setae at distal articulation. Scattered short setae present on all segments. Propodus of fifth pereiopods with 4-8 pairs of spines distributed along its length and fine setae at distal articulation. Propodus 2.5
<normalizedToken id="41E63DE53F507BAA46FBE02285AC3DCC" originalValue="×">x</normalizedToken>
longer than carpus. Propodus as long as merus. Scattered short setae present on all segments.
</paragraph>
<paragraph id="279E9F7A6AD3F3E358AA55F7E0F0E3EF" pageId="0" pageNumber="163">
<emphasis id="EBBD7CCC9ED46C4A67AA0BEF3F3D50D0" bold="true" italics="true" pageId="0" pageNumber="163">Thoracic sternum.</emphasis>
Fourth and fifth thoracic sternites with moderately transverse plate without median process, and seventh thoracic sternite smooth. Eighth thoracic sternite usually smooth.
</paragraph>
<paragraph id="BFF0DCC0EAFDFE6DE292E40CDB76945F" pageId="0" pageNumber="163">
<emphasis id="0761966D5F96300CD9BF69AE50709EDF" bold="true" italics="true" pageId="0" pageNumber="163">Pleon.</emphasis>
Smooth. All pleonal sternites with transverse ridges. First and second pleonal sternites with or without small median processes. Third and fourth pleonal sternites smooth. Fifth pleonal sternite with triangular ridge. Preanal carina present, obtuse ridge developed without spine or setae. Ventral margin of pleural tergum with small setae.
</paragraph>
<paragraph id="1788028AB04C5B16A0216E389B844D4F" pageId="0" pageNumber="163">
<emphasis id="235AFB2FE45835FA4D33644BB3254BAF" bold="true" italics="true" pageId="0" pageNumber="163">Telson</emphasis>
(Fig.
<figureCitation id="5E6DF7ADD27BFE6E5FED8315A23F47F2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">5G</figureCitation>
). Tapered posteriorly, protruding point on middle margin with lateral spines and few fine setae. Inner spines longer than outer spines. Dorsal surface with two pairs of small spines, similar in size.
</paragraph>
<paragraph id="E06A342E9D6A9A243FB434ECFD3AB15F" pageId="0" pageNumber="163">
<emphasis id="5D18A340A2211AE332DDA1CB8E57D934" bold="true" italics="true" pageId="0" pageNumber="163">Uropods</emphasis>
(Fig.
<figureCitation id="43C9A2CF85403B794899BA304B4CB6FE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphological characteristics of Macrobrachium panhai sp. nov. (A, F ovigerous female paratype MUMNH MP 00303 B-E, G-H ovigerous female holotype CUMZ MP 00302) A lateral view B carapace C epistome D second pereiopod E teeth between fingers F third pereiopod G uropod and H movable spine at uropodal diaeresis. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.113898.figure5" httpUri="https://binary.pensoft.net/fig/974857" pageId="0" pageNumber="163">5G</figureCitation>
). Uropodal diaeresis with inner movable spine, as long as or slightly longer than outer angle. Exopods longer than endopods.
</paragraph>
</subSubSection>
<subSubSection id="2DC45DEDACB2E6B723FC6101A3817248" pageId="0" pageNumber="163" type="etymology">
<paragraph id="771F9F71B0A57EDE098E362963D22EF8" pageId="0" pageNumber="163">Etymology.</paragraph>
<paragraph id="C9FC70F6810632C3CDE2AC2A92CE9100" pageId="0" pageNumber="163">
The specific name
<emphasis id="A0DDC21E9430E71D4213D7DF9D80A7AE" italics="true" pageId="0" pageNumber="163">panhai</emphasis>
is dedicated to Prof. Dr. Somsak Panha, a taxonomist from Faculty of Science, Chulalongkorn University, Thailand well known for his remarkable contributions and endorsement to the study of invertebrate fauna in Thailand.
</paragraph>
</subSubSection>
<subSubSection id="E1114F804A752218E3495CF430567A11" pageId="0" pageNumber="163" type="distribution">
<paragraph id="06D5B30EDEBADB3F8EC02CD5DEC6A976" pageId="0" pageNumber="163">Distribution.</paragraph>
<paragraph id="84CF62CC2A11CA8B05191F82FDCE5878" pageId="0" pageNumber="163">This species is distributed in the Chao Phraya and Mekong River Basins, Thailand.</paragraph>
</subSubSection>
<subSubSection id="BCDE71CC7E9A92BFD01130F1B7C01A57" pageId="0" pageNumber="163" type="remarks">
<paragraph id="04709ECE6438CC5DADF018E909E7AB98" pageId="0" pageNumber="163">Remarks.</paragraph>
<paragraph id="7ACA9AFF0FC987D4FE3D067B0EF69A19" pageId="0" pageNumber="163">
<taxonomicName id="1F61F0E668BC4D8CF663E80AB2F42B58" authorityName="Chaowvieng &amp; Siriwut" authorityYear="2024" class="Insecta" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrobrachium panhai" order="Decapoda" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="13F97AAB03F356CCCB88838AB35FD02D" italics="true" pageId="0" pageNumber="163">Macrobrachium panhai</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName id="AF8BD5A4DB46FACFD944DC8797A83515" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="8271CB1310C1AE397D09CEA56AAD16FC" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
s. str. due to having the rostral formula with 8-12/3-6 teeth (vs 6-10/1-6 teeth in
<taxonomicName id="54482EAF6BDDDD4BEEAA566BCCAB76F2" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="87CDAD6CD8854C32B645B02A0B2B278D" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
). Movable spine at uropodal diaeresis is slightly longer than outer angle (vs movable spine is shorter in
<taxonomicName id="9576116ECE99D7994097EF4C7EB7C149" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="3D363910148887DC26C62E0C89F32706" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
). Third propodus has 3-6 pairs of spines (vs 4-8 pairs of spines in
<taxonomicName id="BE1D154CD08BE17F40703835810DA669" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="335707552395603BD40C5F87202598CD" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
). The teeth between fingers of second pereiopods are absent (vs 1 or 2 teeth on fixed and movable finger in
<taxonomicName id="FF9F3C08BB3B324AAB4B9C99271BB08E" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="694FC497C7335D4D4D99C4C65A51F73B" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
). The ratio between rostrum and carapace length is 0.89-1.53 (vs 0.78-1.14 in
<taxonomicName id="15E9C6F8A12FA02AFF420235971D1CDA" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="A4A60E42A9EE06215208BB9180D1315C" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
) and the ratio between chela and carpus is 1.2-2.08 (vs 1.15-1.72 in
<taxonomicName id="ECA67958136AB48801C4A60115BE9B51" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="AA3C1DDB2E976BF4651F90F4D92F8DA6" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
).
<taxonomicName id="C56FC792BFEDF565D6E9B613CAB993CF" lsidName="M. panhai" pageId="0" pageNumber="163" rank="species" species="panhai">
<emphasis id="D83FC10D08874A4BD6DDAFEC408671B2" italics="true" pageId="0" pageNumber="163">M. panhai</emphasis>
</taxonomicName>
sp. nov. occasionally co-exists with
<taxonomicName id="5F03738A793BE3B129C328917E817007" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="A1161EB01CDE3C0CBB916D0C9EE48EF8" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
in the Chao Phraya and Mekong River Basins.
</paragraph>
<paragraph id="675CC8D22CC7859554E7EF2EB3ACF488" pageId="0" pageNumber="163">
This new species also differs from
<taxonomicName id="4A57D4A8726914E5ECA629DDAC18844F" lsidName="M. peguense" pageId="0" pageNumber="163" rank="species" species="peguense">
<emphasis id="9D7115E526DC2EA4E5B14F93D0DA6A7C" italics="true" pageId="0" pageNumber="163">M. peguense</emphasis>
</taxonomicName>
sensu
<bibRefCitation id="334E14541EDB9C1C00AEAA1368816278" DOI="https://doi.org/10.1080/00222935208654260" author="Tiwari, KK" journalOrPublisher="Annals &amp; Magazine of Natural History" pageId="0" pageNumber="163" pagination="27 - 32" refId="B76" refString="Tiwari, KK, 1952. Diagnosis of new species and subspecies of the genus Palaemon Fabricius (Crustacea: Decapoda). Annals &amp; Magazine of Natural History 12 (49): 27 - 32, DOI: https://doi.org/10.1080/00222935208654260" title="Diagnosis of new species and subspecies of the genus Palaemon Fabricius (Crustacea: Decapoda)." url="https://doi.org/10.1080/00222935208654260" volume="12" year="1952">Tiwari (1952)</bibRefCitation>
by processes of rostral formula 8-12/3-6 teeth (vs 6-9/2-4 teeth in
<taxonomicName id="887B90AF8FE13F634FC8ACD1054A1679" lsidName="M. peguense" pageId="0" pageNumber="163" rank="species" species="peguense">
<emphasis id="DF40FCA7C4B2A5A47D2DD73CF2AB82B4" italics="true" pageId="0" pageNumber="163">M. peguense</emphasis>
</taxonomicName>
). Second pereiopods had palms shorter than half of carpus (vs palm slightly more than half of carpus in
<taxonomicName id="5D945E80586AAECC0D18EAEBBE70A1F3" lsidName="M. peguense" pageId="0" pageNumber="163" rank="species" species="peguense">
<emphasis id="370E621222B2C0E125B6C8A8940EE2BA" italics="true" pageId="0" pageNumber="163">M. peguense</emphasis>
</taxonomicName>
). Propodus of third pereiopods are 2
<normalizedToken id="8FB7F2A0760F2161AE57B617E24A2893" originalValue="×">x</normalizedToken>
longer than dactylus (vs 3 in
<taxonomicName id="74FBCB0B925233FABF81BF317F26F3D3" lsidName="M. peguense" pageId="0" pageNumber="163" rank="species" species="peguense">
<emphasis id="2C805AFB5D7169E79499E6D0DAC3CFCF" italics="true" pageId="0" pageNumber="163">M. peguense</emphasis>
</taxonomicName>
). Dorsal surface of telson is without depression (vs longitudinal depression in
<taxonomicName id="09F519E6AE48ADFCE3DD2A90339BE8A3" lsidName="M. peguense" pageId="0" pageNumber="163" rank="species" species="peguense">
<emphasis id="00A59AB9DF8E16D3BCC1D28155B8BC9E" italics="true" pageId="0" pageNumber="163">M. peguense</emphasis>
</taxonomicName>
). Movable spine at uropodal diaeresis is slightly longer than outer angle (vs movable spine is shorter in
<taxonomicName id="940B7CDC19A86FF7EA7641CF76A9A17A" lsidName="M. peguense" pageId="0" pageNumber="163" rank="species" species="peguense">
<emphasis id="BC501B7218FADE2B9C399CD4FE2D904B" italics="true" pageId="0" pageNumber="163">M. peguense</emphasis>
</taxonomicName>
).
<bibRefCitation id="328894A6F3C27979B3C4DC55F694F8A9" DOI="https://doi.org/10.1023/A:1022991224381" author="Cai, Y" journalOrPublisher="Hydrobiologia" pageId="0" pageNumber="163" pagination="59 - 83" refId="B1" refString="Cai, Y, Ng, PKL, 2002. The freshwater palaemonid prawns (Crustacea: Decapoda: Caridea) of Myanmar. Hydrobiologia 487 (1): 59 - 83, DOI: https://doi.org/10.1023/A:1022991224381" title="The freshwater palaemonid prawns (Crustacea: Decapoda: Caridea) of Myanmar." url="https://doi.org/10.1023/A:1022991224381" volume="487" year="2002">Cai and Ng (2002)</bibRefCitation>
also mentioned that the egg size can be used to distinguish
<taxonomicName id="F6D128D6C2821FEB87B4D312BE17E5A7" lsidName="M. peguense" pageId="0" pageNumber="163" rank="species" species="peguense">
<emphasis id="E5C111C0C9803D064ED6FDFB8B012378" italics="true" pageId="0" pageNumber="163">M. peguense</emphasis>
</taxonomicName>
and
<taxonomicName id="BA147C4B4F4EB486507426462E68DD06" lsidName="M. lanchesteri" pageId="0" pageNumber="163" rank="species" species="lanchesteri">
<emphasis id="8C458D2F6E90390D009C89D48D473C3F" italics="true" pageId="0" pageNumber="163">M. lanchesteri</emphasis>
</taxonomicName>
group (1.15-1.5
<normalizedToken id="BFC5F12BC2A0D79838D4A2083F5627A7" originalValue="×">x</normalizedToken>
1.6-2.1 mm and 0.6-0.7
<normalizedToken id="C3E800C838060F6760A55AAD9CE39C03" originalValue="×">x</normalizedToken>
0.8-1 mm, respectively). Currently, the distribution range of
<taxonomicName id="FC6A1B1433A9B0B3CE521A773BC2CFEC" lsidName="M. peguense" pageId="0" pageNumber="163" rank="species" species="peguense">
<emphasis id="F0221260D24C4E9A9D4B8C345F352B6C" italics="true" pageId="0" pageNumber="163">M. peguense</emphasis>
</taxonomicName>
was found only from Myanmar.
</paragraph>
</subSubSection>
</treatment>
</document>
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