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<document id="2D9DD353F982C9D5B1434F37311D41A2" ID-DOI="10.5281/zenodo.157036" ID-GBIF-Dataset="8845a192-a81c-4e81-a280-e19c508ce7dd" ID-ISSN="1175­5326" ID-Zenodo-Dep="157036" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459633983606" checkinUser="plazi" docAuthor="Shear, William A." docDate="2003" docId="F7710943FF813D0898781083FD82F8E4" docLanguage="en" docName="zt00321.pdf" docOrigin="Zootaxa 321" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Nannopetalum pattersonorum Shear, 2003, n. sp." docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="0B48713BFF8C3D0699701225FFF0FFBB" masterDocTitle="The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="14" updateTime="1698204205547" updateUser="plazi">
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<mods:title id="2625F845B35FC623F2B4AD07F4300B76">The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)</mods:title>
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<mods:namePart id="1571A569DFB9ADC479A63332502B73B3">Shear, William A.</mods:namePart>
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<mods:date id="9E914C31DC6184E1866B351B7E0BCDAF">2003</mods:date>
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<treatment id="F7710943FF813D0898781083FD82F8E4" ID-DOI="http://doi.org/10.5281/zenodo.6273578" ID-GBIF-Taxon="119332284" ID-Zenodo-Dep="6273578" LSID="urn:lsid:plazi:treatment:F7710943FF813D0898781083FD82F8E4" httpUri="http://treatment.plazi.org/id/F7710943FF813D0898781083FD82F8E4" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="37C2EBDEFF813D0B98781083FE84FD58" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="7F67B855FF813D0B98781083FD36FD7B" blockId="13.[264,710,678,739]" box="[264,710,678,704]" pageId="13" pageNumber="14">
<heading id="242F0F39FF813D0B98781083FD36FD7B" bold="true" box="[264,710,678,704]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="4DAC6447FF813D0B98781083FD36FD7B" bold="true" box="[264,710,678,704]" pageId="13" pageNumber="14">
<emphasis id="4DAC6447FF813D0B98781083FD8CFD7B" box="[264,636,678,704]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="B8D8C3D6FF813D0B98781083FD87FD7B" ID-CoL="45MLH" box="[264,631,678,704]" class="Diplopoda" family="Trichopetalidae" genus="Nannopetalum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pattersonorum" status="sp. nov.">Nannopetalum pattersonorum</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="569FD93CFF813D0B9BF41083FD36FD7B" box="[644,710,678,704]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7F67B855FF813D0B987810ECFE84FD58" blockId="13.[264,710,678,739]" box="[264,372,713,739]" pageId="13" pageNumber="14">
<figureCitation id="E7E3A4D0FF813D0B987810ECFE84FD58" box="[264,372,713,739]" captionStart="FIGURES 1 4" captionStartId="14.[264,382,1562,1586]" captionTargetBox="[326,1258,313,1561]" captionTargetPageId="14" captionText="FIGURES 1 4. Nannopetalum pattersonorum, n. sp. 1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm)." httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="13" pageNumber="14">Figs. 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF813D0B9878113FFD76FC68" pageId="13" pageNumber="14" type="description">
<paragraph id="7F67B855FF813D0B9878113FFD76FC68" blockId="13.[264,1324,793,1899]" pageId="13" pageNumber="14">
<emphasis id="4DAC6447FF813D0B9878113FFEA2FC88" box="[264,338,794,819]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="A06306F7FF813D0B9878113FFEB8FC88" box="[264,328,794,819]" pageId="13" pageNumber="14">Types</typeStatus>
:
</emphasis>
Male
<typeStatus id="A06306F7FF813D0B98D4113CFDFEFC88" box="[420,526,793,819]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
,
<specimenCount id="69DE73DCFF813D0B9B6B113CFD9DFC88" box="[539,621,793,819]" pageId="13" pageNumber="14" type="male">2 male</specimenCount>
<typeStatus id="A06306F7FF813D0B9B06113CFD16FC88" box="[630,742,793,819]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
and
<specimenCount id="69DE73DCFF813D0B9A54113CFC7CFC88" box="[804,908,793,819]" pageId="13" pageNumber="14" type="female">3 female</specimenCount>
<typeStatus id="A06306F7FF813D0B9AE6113CFBF6FC88" box="[918,1030,793,819]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
(FMNH) from Virginia, Prince Edward County, Hampden­Sydney College campus, Wilson Trail, near stream in maple­sycamore litter, collected
<date id="0B669E95FF813D0B9BEE114CFC8EFC38" box="[670,894,873,899]" pageId="13" pageNumber="14" value="1988-12-07">7 December 1988</date>
by W. A. Shear. Male
<typeStatus id="A06306F7FF813D0B9DB6114CFADBFC38" box="[1222,1323,873,899]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
(FMNH) from same locality, but behind Gilmer Hall, in pine bark­fern litter, collected
<date id="0B669E95FF813D0B9C7D11B4FE36FC68" pageId="13" pageNumber="14" value="1987-11-15">15 November 1987</date>
by W. A. Shear.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF813D0B984811C4FE38FB98" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="7F67B855FF813D0B984811C4FE38FB98" blockId="13.[264,1324,793,1899]" pageId="13" pageNumber="14">
<emphasis id="4DAC6447FF813D0B984811C4FE49FC41" box="[312,441,993,1018]" italics="true" pageId="13" pageNumber="14">Diagnosis:</emphasis>
a larger species of
<taxonomicName id="B8D8C3D6FF813D0B9BDD11C4FCAAFC41" box="[685,858,993,1018]" class="Diplopoda" family="Trichopetalidae" genus="Nannopetalum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF813D0B9BDD11C4FCAAFC41" box="[685,858,993,1018]" italics="true" pageId="13" pageNumber="14">Nannopetalum</emphasis>
</taxonomicName>
with seven (rarely six) ocelli on each side of the head.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF813D0B98481614FCF2FBC8" pageId="13" pageNumber="14" type="etymology">
<paragraph id="7F67B855FF813D0B98481614FCF2FBC8" blockId="13.[264,1324,793,1899]" pageId="13" pageNumber="14">
<emphasis id="4DAC6447FF813D0B98481614FE4EFBF1" box="[312,446,1073,1098]" italics="true" pageId="13" pageNumber="14">Etymology:</emphasis>
Named for Mr. and Mrs. Charles Patterson, whose generous bequest to Hampden­Sydney College aided this work.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF813D0B984816A4FCC8F970" pageId="13" pageNumber="14" type="description">
<paragraph id="7F67B855FF813D0B984816A4FECFFA80" blockId="13.[264,1324,793,1899]" pageId="13" pageNumber="14">
<emphasis id="4DAC6447FF813D0B984816A4FE1CFB21" box="[312,492,1153,1178]" italics="true" pageId="13" pageNumber="14">
Male
<typeStatus id="A06306F7FF813D0B980D16A4FE15FB21" box="[381,485,1153,1178]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
:
</emphasis>
Total length, 6.2 mm; width, 0.6 mm. Seven ocelli in T­shaped rows. Unpigmented. Segmental setae 50% body width. Segments with pronounced lateral striations; segmental shoulders moderately developed. Legpairs 37 enlarged, pairs 3 and 4 the most robust, pairs 5 to 7 more slender but still markedly larger than postgonopodal legs.
</paragraph>
<paragraph id="7F67B855FF813D0B9848176CFC26F9C0" blockId="13.[264,1324,793,1899]" pageId="13" pageNumber="14">
Gonopods (
<figureCitation id="E7E3A4D0FF813D0B98B0176CFDDDFAD8" box="[448,557,1353,1379]" captionStart="FIGURES 1 4" captionStartId="14.[264,382,1562,1586]" captionTargetBox="[326,1258,313,1561]" captionTargetPageId="14" captionText="FIGURES 1 4. Nannopetalum pattersonorum, n. sp. 1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm)." httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="13" pageNumber="14">Figs. 13</figureCitation>
) small, partly concealed. Sternal lobes moderately developed (
<emphasis id="4DAC6447FF813D0B9C68176CFADAFAD9" box="[1304,1322,1353,1378]" italics="true" pageId="13" pageNumber="14">s,</emphasis>
<figureCitation id="E7E3A4D0FF813D0B98781754FEBDFA30" box="[264,333,1393,1419]" captionStart="FIGURES 1 4" captionStartId="14.[264,382,1562,1586]" captionTargetBox="[326,1258,313,1561]" captionTargetPageId="14" captionText="FIGURES 1 4. Nannopetalum pattersonorum, n. sp. 1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm)." httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
). Coxae (
<emphasis id="4DAC6447FF813D0B98BE1754FE00FA31" box="[462,496,1393,1418]" italics="true" pageId="13" pageNumber="14">cc,</emphasis>
<figureCitation id="E7E3A4D0FF813D0B98881754FDCCFA30" box="[504,572,1393,1419]" captionStart="FIGURES 1 4" captionStartId="14.[264,382,1562,1586]" captionTargetBox="[326,1258,313,1561]" captionTargetPageId="14" captionText="FIGURES 1 4. Nannopetalum pattersonorum, n. sp. 1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm)." httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
) with 2 lateral setae each on moderate shoulders; angiocoxites (
<emphasis id="4DAC6447FF813D0B986217BCFECBFA09" box="[274,315,1433,1458]" italics="true" pageId="13" pageNumber="14">acx</emphasis>
,
<figureCitation id="E7E3A4D0FF813D0B983C17BCFE61FA08" box="[332,401,1433,1459]" captionStart="FIGURES 1 4" captionStartId="14.[264,382,1562,1586]" captionTargetBox="[326,1258,313,1561]" captionTargetPageId="14" captionText="FIGURES 1 4. Nannopetalum pattersonorum, n. sp. 1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm)." httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
) strongly projecting basally, basal half set with typical trichomes, becoming longer distolaterally; 3 basal setae; anterior process with 56 coarse teeth, 4 apical setae; posterior process mesal, subtriangular, posteriorly curved. Fimbriate branch reduced. Colpocoxites (
<emphasis id="4DAC6447FF813D0B98C61434FE20F991" box="[438,464,1553,1578]" italics="true" pageId="13" pageNumber="14">cc</emphasis>
,
<figureCitation id="E7E3A4D0FF813D0B98921434FDD9F990" box="[482,553,1553,1579]" captionStart="FIGURES 1 4" captionStartId="14.[264,382,1562,1586]" captionTargetBox="[326,1258,313,1561]" captionTargetPageId="14" captionText="FIGURES 1 4. Nannopetalum pattersonorum, n. sp. 1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm)." httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
;
<figureCitation id="E7E3A4D0FF813D0B9B4B1434FD72F990" box="[571,642,1553,1579]" captionStart="FIGURES 1 4" captionStartId="14.[264,382,1562,1586]" captionTargetBox="[326,1258,313,1561]" captionTargetPageId="14" captionText="FIGURES 1 4. Nannopetalum pattersonorum, n. sp. 1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm)." httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
) lobelike, squarish, small plumose processes on anterior faces. Ninth legs (
<figureCitation id="E7E3A4D0FF813D0B9890141CFDD8F9E8" box="[480,552,1593,1619]" captionStart="FIGURES 1 4" captionStartId="14.[264,382,1562,1586]" captionTargetBox="[326,1258,313,1561]" captionTargetPageId="14" captionText="FIGURES 1 4. Nannopetalum pattersonorum, n. sp. 1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm)." httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
) as usual for the family; coxae slightly excavated on mesal side. Legpairs 10 and 11 with coxal glands, otherwise unmodified.
</paragraph>
<paragraph id="7F67B855FF813D0B984814ACFCC8F970" blockId="13.[264,1324,793,1899]" pageId="13" pageNumber="14">
<emphasis id="4DAC6447FF813D0B984814ACFDF0F919" box="[312,512,1673,1698]" italics="true" pageId="13" pageNumber="14">
Female
<typeStatus id="A06306F7FF813D0B98E814ACFDF0F919" box="[408,512,1673,1698]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
</emphasis>
: Total length 8.0 mm (extended specimens); width, 0.7 mm. Ocelli 7, curved row of 6, plus one out­of­row ocellus.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF813D0B984814FCFD85F8D0" pageId="13" pageNumber="14" type="distribution">
<paragraph id="7F67B855FF813D0B984814FCFD85F8D0" blockId="13.[264,1324,793,1899]" pageId="13" pageNumber="14">
<emphasis id="4DAC6447FF813D0B984814FCFE21F949" box="[312,465,1753,1778]" italics="true" pageId="13" pageNumber="14">Distribution:</emphasis>
Definitely known only from the Hampden­Sydney College campus, but six females taken in Prince Edward Co., Virginia, along the Bush River 3.2 miles northwest of Meherrin, from beech litter, conform to this species. These specimens were collected by me on
<date id="0B669E95FF813D0B98BA1574FD81F8D0" box="[458,625,1873,1899]" pageId="13" pageNumber="14" value="1977-05-15">May 15, 1977</date>
.
</paragraph>
</subSubSection>
<caption id="2BA7E8DDFF823D089878143FFD51F9CC" httpUri="https://zenodo.org/record/168791/files/figure.png" pageId="14" pageNumber="15" targetBox="[326,1258,313,1561]" targetPageId="14">
<paragraph id="7F67B855FF823D089878143FFD51F9CC" blockId="14.[264,1325,1562,1655]" pageId="14" pageNumber="15">
<emphasis id="4DAC6447FF823D089878143FFE43F989" bold="true" box="[264,435,1562,1586]" pageId="14" pageNumber="15">FIGURES 14.</emphasis>
<emphasis id="4DAC6447FF823D0898CA143EFCF1F989" box="[442,769,1563,1586]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="B8D8C3D6FF823D0898CA143EFD0DF989" box="[442,765,1563,1586]" class="Diplopoda" family="Trichopetalidae" genus="Nannopetalum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pattersonorum" status="sp. nov.">Nannopetalum pattersonorum</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="569FD93CFF823D089A76143EFCCFF989" box="[774,831,1563,1586]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
1, gonopod complex, ventral view (SEM). 2, gonopods, anterior view. 3, gonopod colopocoxites, anterior view. 4, male ninth legpair, posterior view. Scale lines = 0.1 mm (100 µm).
</paragraph>
</caption>
<subSubSection id="37C2EBDEFF823D08984814E8FD82F8E4" pageId="14" pageNumber="15" type="discussion">
<paragraph id="7F67B855FF823D08984814E8FD82F8E4" blockId="14.[264,1323,1741,1887]" pageId="14" pageNumber="15">
<emphasis id="4DAC6447FF823D08984814E8FE33F95D" box="[312,451,1741,1766]" italics="true" pageId="14" pageNumber="15">Discussion:</emphasis>
Despite repeated sampling over the years of the exact spot where the
<typeStatus id="A06306F7FF823D089D9D14E8FADBF95C" box="[1261,1323,1741,1767]" pageId="14" pageNumber="15">types</typeStatus>
were collected, no further material came to light; however, none of these later samples were taken in winter. Finally, in January, 2003, a sample from the
<typeStatus id="A06306F7FF823D089D6E1538FBA1F88C" box="[1054,1105,1821,1847]" pageId="14" pageNumber="15">type</typeStatus>
locality contained a single male and two females.
</paragraph>
</subSubSection>
</treatment>
</document>