133 lines
12 KiB
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133 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. & Tishechkin, Alexey K." docDate="2013" docId="A274B1B4AAF34C1D2042B90FD453CCB3" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus iheringi Bickhardt 1917" docType="treatment" docVersion="3" lastPageNumber="368" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="367" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Caterino, Michael S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>271</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>401</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040889" LSID="urn:lsid:plazi:treatment:A274B1B4AAF34C1D2042B90FD453CCB3" httpUri="http://treatment.plazi.org/id/A274B1B4AAF34C1D2042B90FD453CCB3" lastPageId="367" lastPageNumber="368" pageId="366" pageNumber="367">
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<subSubSection pageId="366" pageNumber="367" type="nomenclature">
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<paragraph pageId="366" pageNumber="367">
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<taxonomicName LSID="http://species-id.net/wiki/Operclipygus_iheringi" authority="Bickhardt, 1917" authorityName="Bickhardt" authorityYear="1917" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus iheringi" order="Coleoptera" pageId="366" pageNumber="367" phylum="Arthropoda" rank="species" species="iheringi">Operclipygus iheringi (Bickhardt, 1917)</taxonomicName>
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Figs 103
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<normalizedToken originalValue="A–B104A–EMap">A-B104A-EMap</normalizedToken>
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34
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</paragraph>
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</subSubSection>
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<subSubSection pageId="366" pageNumber="367" type="reference_group">
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<paragraph pageId="366" pageNumber="367">
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<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister iheringi" order="Coleoptera" pageId="366" pageNumber="367" phylum="Arthropoda" rank="species" species="iheringi">Phelister iheringi</taxonomicName>
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Bickhardt, 1917: 215;
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus iheringi" order="Coleoptera" pageId="366" pageNumber="367" phylum="Arthropoda" rank="species" species="iheringi">Operclipygus iheringi</taxonomicName>
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:
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<bibRefCitation pageId="366" pageNumber="367">Mazur (1984</bibRefCitation>
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: 258).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="366" pageNumber="367" type="type locality">
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<paragraph pageId="366" pageNumber="367">Type locality.</paragraph>
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<paragraph pageId="366" pageNumber="367">
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BRAZIL:
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Paulo: Rair da Serra [exact location unknown].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="366" pageNumber="367" type="type material">
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<paragraph pageId="366" pageNumber="367">Type material.</paragraph>
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<paragraph pageId="366" pageNumber="367">
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Lectotype, here designated: "Rair da Serra,
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Paulo" /
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<normalizedToken originalValue="“Type”">"Type"</normalizedToken>
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/
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<normalizedToken originalValue="“9968”">"9968"</normalizedToken>
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/ "LECTOTYPE
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<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister iheringi" order="Coleoptera" pageId="366" pageNumber="367" phylum="Arthropoda" rank="species" species="iheringi">Phelister iheringi</taxonomicName>
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Bickhardt, 1917 M.S. Caterino & A.K. Tishechkin des. 2010" (ZMHB). Paralectotype: same data as lectotype (ZMHB). This species was described from an unspecified number of specimens, and the lectotype designation fixes primary type status on one of the two known syntypes.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="366" pageNumber="367" type="other material">
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<paragraph pageId="366" pageNumber="367">Other material.</paragraph>
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<paragraph pageId="366" pageNumber="367">
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BRAZIL:
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<normalizedToken originalValue="Paraná">Parana</normalizedToken>
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: 1: Piraquara, Mananciais da Serra, 1000m, i.2007, P. Grossi (UFPR); 1: Piraquara, Sanepar, 18.vii.2001, P. Grossi (UFPR); 2:
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<normalizedToken originalValue="Guartelá">Guartela</normalizedToken>
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, Campo
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Paolo,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-24.533333">24°32'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-50.283333">50°17'W</geoCoordinate>
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, 900m, xi.2007, pitfall, E. Grossi (UFPR).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="367" lastPageNumber="368" pageId="366" pageNumber="367" type="description">
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<paragraph pageId="366" pageNumber="367">Diagnostic description.</paragraph>
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<paragraph lastPageId="367" lastPageNumber="368" pageId="366" pageNumber="367">
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Length: 3.21-3.46 mm, width: 2.56-2.87 mm; body piceous, large, elongate, subdepressed; frons weakly depressed at middle, finely punctate; frontal stria rounded at sides, with a short, detached fragment in front, supraorbital stria complete, connected to sides of frontal; epistoma convex; labrum wide, weakly emarginate; both mandibles with bifid basal teeth; antennal club markedly elongate with basal and middle annuli interrupted; pronotum with lateral margins slightly convergent in basal half, weakly curved inward to apex; pronotal disk with distinct narrow prescutellar impression about as long as scutellum, disk finely punctate throughout, with few barely coarser lateral punctures; marginal pronotal stria complete or narrowly interrupted behind head; submarginal stria continuous from sides across front; central portion of anterior pronotal margin faintly projecting at middle; anterior pronotal margin with a single pair of glands, behind eye on each side; elytron with two complete epipleural punctures, outer subhumeral stria nearly complete, sinuate at middle, frequently abbreviated at base, inner subhumeral stria present in middle, variously abbreviated basally and apically, striae 1-3 complete, 4th stria present in apical third, 5th stria present in apical fourth, sutural stria present in apical half to two-thirds; elytra with broad band of small punctures present along apical margin; prosternal keel broad, flat, weakly emarginate at base, carinal striae convergent to apex, rarely connected by arch; mesoventral margin sinuate, median projection small, acute, marginal stria complete or narrowly interrupted; mesometaventral
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<pageBreakToken pageId="367" pageNumber="368" start="start">stria</pageBreakToken>
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arched to subangulate at middle, reaching anterior half of mesoventral disk; lateral metaventral stria extending posterolaterad to middle of metacoxa; 1st abdominal ventrite with two complete lateral striae very close together, central part of disk with numerous small punctures near these striae; propygidium uniformly covered with small, round punctures separated by slightly less than their diameters; pygidium lacking dense ground punctation, with small punctures diminishing toward apex; marginal pygidial sulcus fine, variably abbreviated at sides, never attaining base. Male genitalia (Figs 104
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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): accessory sclerites present, small; T8 with sides subparallel at middle, slightly divergent basally, desclerotized at apicolateral angles, convergent to apex, basal emargination shallow, reaching less than halfway to basal membrane attachment line, ventrolateral apodemes evenly rounded, distant beneath; S8 with unique median longitudinal sclerite, inner edges of lateral halves strongly divergent, apical guides strongly developed apically, with weakly hooked inner edges; T9 widest near apex, narrowed to basal apodemes, with strongly dentate ventrolateral apodemes, apices narrowed, blunt; T10 completely divided; S9 narrow at middle, weakly widened to narrowly rounded base, apex distinctly emarginate, apical flanges separate; tegmen with sides subparallel in basal half, rather abruptly narrowed to apices, medioventral process narrowly
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<normalizedToken originalValue="‘U’">'U'</normalizedToken>
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-shaped, projecting strongly beneath, about one-fourth from base; basal piece about one-third tegmen length; median lobe about one-half tegmen length, proximal apodemes not obviously differentiated.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="367" pageNumber="368" type="remarks">
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<paragraph pageId="367" pageNumber="368">Remarks.</paragraph>
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<paragraph pageId="367" pageNumber="368">
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This is a very distinctive species of rather obscure relationships. It is much larger than average, with the lateral submarginal pronotal stria close to the margin (Fig. 103A), an unusually elongate antennal club, and a broad band of apical marginal elytral punctures. The median ventral sclerite on the 8th sternite of the male genitalia (Fig. 104B) is unique in the genus. A few characters suggest relationships to the
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="367" pageNumber="368" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
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group, including the desclerotized apicolateral angles of the male 8th tergite. But the many other unique features, particularly the presence of only a single pair of anterior pronotal gland openings, preclude a confident placement at present.
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</paragraph>
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<caption pageId="367" pageNumber="368">
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<paragraph pageId="367" pageNumber="368">
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Figure 103.
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus iheringi" order="Coleoptera" pageId="367" pageNumber="368" phylum="Arthropoda" rank="species" species="iheringi">Operclipygus iheringi</taxonomicName>
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. A Dorsal habitus B Pygidia.
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</paragraph>
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</caption>
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<caption pageId="367" pageNumber="368">
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<paragraph pageId="367" pageNumber="368">
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Figure 104. Male genitalia of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus iheringi" order="Coleoptera" pageId="367" pageNumber="368" phylum="Arthropoda" rank="species" species="iheringi">Operclipygus iheringi</taxonomicName>
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. A T8 B S8 C T9 & T10 D S9 E Aedeagus, dorsal and lateral views.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |